Serendipaceratops
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''Serendipaceratops'' (meaning " serendipitous horned face") is a
genus Genus (; : genera ) is a taxonomic rank above species and below family (taxonomy), family as used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In bino ...
of herbivorous
ornithischia Ornithischia () is an extinct clade of mainly herbivorous dinosaurs characterized by a pelvic structure superficially similar to that of birds. The name ''Ornithischia'', or "bird-hipped", reflects this similarity and is derived from the Greek ...
n
dinosaur Dinosaurs are a diverse group of reptiles of the clade Dinosauria. They first appeared during the Triassic Geological period, period, between 243 and 233.23 million years ago (mya), although the exact origin and timing of the #Evolutio ...
, possibly an
ankylosaur Ankylosauria is a group of herbivorous dinosaurs of the clade Ornithischia. It includes the great majority of dinosaurs with armor in the form of bony osteoderms, similar to turtles. Ankylosaurs were bulky quadrupeds, with short, powerful l ...
, from the early
Cretaceous The Cretaceous ( ) is a geological period that lasted from about 143.1 to 66 mya (unit), million years ago (Mya). It is the third and final period of the Mesozoic Era (geology), Era, as well as the longest. At around 77.1 million years, it is the ...
Period of Australia. The type species, ''S. arthurcclarkei'', was named in 2003.


Discovery and species

The first known bone from ''Serendipaceratops'', an
ulna The ulna or ulnar bone (: ulnae or ulnas) is a long bone in the forearm stretching from the elbow to the wrist. It is on the same side of the forearm as the little finger, running parallel to the Radius (bone), radius, the forearm's other long ...
(lower arm bone), was in 1993 discovered in Australia near Kilcunda, on the south-east coast of Victoria. The find took place during the " Dinosaur Cove" project, uncovering remains at the basis of a cliff face at the shoreline; the fossil was excavated in the "Arch", a small area of sea-floor protected from the waves by a dam. In 2003, the
type species In International_Code_of_Zoological_Nomenclature, zoological nomenclature, a type species (''species typica'') is the species name with which the name of a genus or subgenus is considered to be permanently taxonomically associated, i.e., the spe ...
''Serendipaceratops arthurcclarkei'' was named and described by Tom Rich and
Patricia Vickers-Rich Patricia Arlene Vickers-Rich (born 11 July 1944), also known as Patricia Rich, is an Australian Professor of Palaeontology and Palaeobiology, who researches the environmental changes that have impacted Australia (including the ancient super ...
.T. Rich and P. Vickers-Rich. 2003. Protoceratopsian? ulnae from Australia. ''Records of the Queen Victoria Museum Launceston'' 113:1-12 Initially, the discoverers had not considered the possibility that the fossil might have been ceratopsian in nature as at this would have been the last group of
dinosaur Dinosaurs are a diverse group of reptiles of the clade Dinosauria. They first appeared during the Triassic Geological period, period, between 243 and 233.23 million years ago (mya), although the exact origin and timing of the #Evolutio ...
s one would have expected to find evidence of in Australia. Instead they had tried to convince themselves it was a
theropod Theropoda (; from ancient Greek , (''therion'') "wild beast"; , (''pous, podos'') "foot"">wiktionary:ποδός"> (''pous, podos'') "foot" is one of the three major groups (clades) of dinosaurs, alongside Ornithischia and Sauropodom ...
bone. Some months later, however, on a visit to the Royal Tyrrell Museum of Palaeontology in
Alberta Alberta is a Provinces and territories of Canada, province in Canada. It is a part of Western Canada and is one of the three Canadian Prairies, prairie provinces. Alberta is bordered by British Columbia to its west, Saskatchewan to its east, t ...
, Canada, colleague
Dale Russell Dale Alan Russell (27 December 1937 – 21 December 2019) was an American-Canadian geologist and palaeontologist. Throughout his career Russell worked as the Curator of Fossil Vertebrates at the Canadian Museum of Nature, Research Professor at ...
pointed out its striking similarity to the ulna of ''
Leptoceratops ''Leptoceratops'' (meaning 'small horn face') is a genus of ceratopsian dinosaur from the Late Cretaceous of North America. First found in Alberta in 1910, the type species ''Leptoceratops gracilis'' was named in 1914 by Barnum Brown for a parti ...
''. Hence the genus name, referring to
serendipity Serendipity is an unplanned fortunate discovery. The term was coined by Horace Walpole in 1754. The concept is often associated with scientific and technological breakthroughs, where accidental discoveries led to new insights or inventions. Ma ...
and combining this reference with ~''ceratops'', a common suffix in ceratopsian generic names. The specific name honours
science fiction Science fiction (often shortened to sci-fi or abbreviated SF) is a genre of speculative fiction that deals with imaginative and futuristic concepts. These concepts may include information technology and robotics, biological manipulations, space ...
writer Arthur C. Clarke. A personal friend of the couple and author of books such as '' 2001: A Space Odyssey'' and '' Rendezvous with Rama'', Clarke first became interested in science as a child because he became fascinated by dinosaurs. Perhaps coincidentally, ''"Serendip"'' is a former name for
Sri Lanka Sri Lanka, officially the Democratic Socialist Republic of Sri Lanka, also known historically as Ceylon, is an island country in South Asia. It lies in the Indian Ocean, southwest of the Bay of Bengal, separated from the Indian subcontinent, ...
, Clarke's adoptive country. ''Serendipaceratops'' is known only from its
holotype A holotype (Latin: ''holotypus'') is a single physical example (or illustration) of an organism used when the species (or lower-ranked taxon) was formally described. It is either the single such physical example (or illustration) or one of s ...
NMV P186385, a single left
ulna The ulna or ulnar bone (: ulnae or ulnas) is a long bone in the forearm stretching from the elbow to the wrist. It is on the same side of the forearm as the little finger, running parallel to the Radius (bone), radius, the forearm's other long ...
found in the Wonthaggi Formation, dating from the early
Aptian The Aptian is an age (geology), age in the geologic timescale or a stage (stratigraphy), stage in the stratigraphic column. It is a subdivision of the Early Cretaceous, Early or Lower Cretaceous epoch (geology), Epoch or series (stratigraphy), S ...
, in 2003 considered to have been about 115 million years old.


Description

The holotype ulna has a preserved length of about sixteen centimetres. At the rear top, its
olecranon The olecranon (, ), is a large, thick, curved bony process on the proximal, posterior end of the ulna. It forms the protruding part of the elbow and is opposite to the cubital fossa or elbow pit (trochlear notch). The olecranon serves as a lever ...
process has broken off. The shaft is rather flattened.


Classification

''Serendipaceratops'' was originally described as a member of the
Neoceratopsia Ceratopsia or Ceratopia ( or ; Greek: "horned faces") is a group of herbivorous, beaked dinosaurs that thrived in what are now North America, Asia and Europe, during the Cretaceous Period, although ancestral forms lived earlier, in the Late Ju ...
and one of the earliest known ceratopsian dinosaurs, and the only one known from the southern hemisphere, with the possible exception of the dubious
South America South America is a continent entirely in the Western Hemisphere and mostly in the Southern Hemisphere, with a considerably smaller portion in the Northern Hemisphere. It can also be described as the southern Subregion#Americas, subregion o ...
n genus '' Notoceratops'', which also may be another kind of
ornithischia Ornithischia () is an extinct clade of mainly herbivorous dinosaurs characterized by a pelvic structure superficially similar to that of birds. The name ''Ornithischia'', or "bird-hipped", reflects this similarity and is derived from the Greek ...
n dinosaur. In addition to the holotype ulna, another supposed ceratopsian ulna was found at Dinosaur Cove, in south-west Victoria. It is a little younger at 106 million years old, having been found in the Eumeralla Formation. The scientists who first studied the ulnae said that they most closely resembled those of ''Leptoceratops'', but subsequent studies have shown that this interpretation is likely incorrect. In a comprehensive survey of the dinosaurs from Australia and New Zealand, Federico Angolin and colleagues in 2010 found that the ulna did not especially resemble that of a basal ceratopsian any more than of any other quadruped
genasauria Ornithischia () is an extinct clade of mainly herbivorous dinosaurs characterized by a pelvic structure superficially similar to that of birds. The name ''Ornithischia'', or "bird-hipped", reflects this similarity and is derived from the Greek st ...
n ornithischian, lacking ceratopsian
synapomorphies In phylogenetics, an apomorphy (or derived trait) is a novel character or character state that has evolved from its ancestral form (or plesiomorphy). A synapomorphy is an apomorphy shared by two or more taxa and is therefore hypothesized to ...
, and that in fact it was more generally similar to the ulna of the Australian
ankylosaur Ankylosauria is a group of herbivorous dinosaurs of the clade Ornithischia. It includes the great majority of dinosaurs with armor in the form of bony osteoderms, similar to turtles. Ankylosaurs were bulky quadrupeds, with short, powerful l ...
'' Minmi''. It could thus not be confidently referred to any ornithischian group, and should be considered a ''
nomen dubium In binomial nomenclature, a ''nomen dubium'' (Latin for "doubtful name", plural ''nomina dubia'') is a scientific name that is of unknown or doubtful application. Zoology In case of a ''nomen dubium,'' it may be impossible to determine whether a ...
''.Agnolin, F.L., Ezcurra, M.D., Pais, D.F. and Salisbury, S.W. (2010). "A reappraisal of the Cretaceous non-avian dinosaur faunas from Australia and New Zealand: Evidence for their Gondwanan affinities." ''Journal of Systematic Palaeontology'', 8(2): 257-300 However, in 2014 Rich ''
et al References Notes References Further reading * * External links * {{Latin phrases E ...
.'' published a statistical study showing that the proportions of the holotype ulna fell within ceratopsian morphospace and found the taxon to be valid based on a unique combination of characters. Most recently, Rozadilla ''et al.'' in 2021 found that while the taxon was valid, the ulna of ''Serendipaceratops'' was deformed, making Rich et al's conclusions about its classification invalid. Instead, Rozadilla et al. classified ''Serendipaceratops'' as an ankylosaur, because of the ulna's robust proportions.


References


External links


Geoscience Australia
(Department of Australian Government)
BBC article
on 19 July 2005

in "Dinosaurs of Darkness" Exhibition
National Geographic ''National Geographic'' (formerly ''The National Geographic Magazine'', sometimes branded as ''Nat Geo'') is an American monthly magazine published by National Geographic Partners. The magazine was founded in 1888 as a scholarly journal, nine ...
Article
Planet DinosaurDinobase
fro
PALAEONTOLOGY RESEARCH GROUP University Of Bristol
{{Taxonbar, from=Q135287 Ornithischia Dinosaur genera Aptian dinosaurs Taxa named by Tom Rich Taxa named by Patricia Vickers-Rich Fossil taxa described in 2003 Dinosaurs of Australia