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Sea snail is a
common name In biology, a common name of a taxon or organism (also known as a vernacular name, English name, colloquial name, country name, popular name, or farmer's name) is a name that is based on the normal language of everyday life; and is often contrast ...
for slow-moving
marine Marine is an adjective meaning of or pertaining to the sea or ocean. Marine or marines may refer to: Ocean * Maritime (disambiguation) * Marine art * Marine biology * Marine debris * Marine habitats * Marine life * Marine pollution Military * ...
gastropod The gastropods (), commonly known as snails and slugs, belong to a large taxonomic class of invertebrates within the phylum Mollusca called Gastropoda (). This class comprises snails and slugs from saltwater, from freshwater, and from land. T ...
molluscs, usually with visible external shells, such as whelk or
abalone Abalone ( or ; via Spanish , from Rumsen ''aulón'') is a common name for any of a group of small to very large marine gastropod molluscs in the family (biology), family Haliotidae. Other common name In biology, a common name of a taxon o ...
. They share the taxonomic class
Gastropoda The gastropods (), commonly known as snails and slugs, belong to a large taxonomic class of invertebrates within the phylum Mollusca called Gastropoda (). This class comprises snails and slugs from saltwater, from freshwater, and from land. ...
with slugs, which are distinguished from snails primarily by the absence of a visible shell.


Definition

Determining whether some gastropods should be called sea snails is not always easy. Some species that live in brackish water (such as certain neritids) can be listed as either
freshwater snails Freshwater snails are gastropod mollusks which live in fresh water. There are many different families. They are found throughout the world in various habitats, ranging from ephemeral pools to the largest lakes, and from small seeps and springs ...
or marine snails, and some species that live at or just above the high tide level (for example species in the genus '' Truncatella'') are sometimes considered to be sea snails and sometimes listed as
land snails A land snail is any of the numerous species of snail that live on land, as opposed to the sea snails and freshwater snails. ''Land snail'' is the common name for terrestrial gastropod mollusks that have shells (those without shells are known as ...
.


Anatomy

Sea snails are a very large group of animals and a very diverse one. Most snails that live in salt water respire using a gill or gills; a few species, though, have a lung, are intertidal, and are active only at low tide when they can move around in the air. These air-breathing species include false limpets in the family Siphonariidae and another group of false limpets in the family Trimusculidae. Many, but not all, sea snails have an operculum.


Shell

The shells of most species of sea snails are spirally coiled. Some, though, have conical shells, and these are often referred to by the common name of
limpet Limpets are a group of aquatic snails that exhibit a conical shell shape (patelliform) and a strong, muscular foot. Limpets are members of the class Gastropoda, but are polyphyletic, meaning the various groups called "limpets" descended indep ...
s. In one unusual family ( Juliidae), the shell of the snail has become two hinged plates closely resembling those of a
bivalve Bivalvia (), in previous centuries referred to as the Lamellibranchiata and Pelecypoda, is a class of marine and freshwater molluscs that have laterally compressed bodies enclosed by a shell consisting of two hinged parts. As a group, bival ...
; this family is sometimes called the "bivalved gastropods". Their shells are found in a variety of shapes and sizes, but are normally very small. Those of living species of sea snails range in size from ''
Syrinx aruanus ''Syrinx aruanus'', common name the Australian trumpet or false trumpet, is a species of extremely large sea snail measuring up to 91 cm long and weighing up to 18 kg. It is a marine gastropod mollusk in the family Turbinellidae, and ...
'', the largest living shelled gastropod species at , to minute species whose shells are less than 1 mm at adult size. Because the shells of sea snails are strong and durable in many cases, as a group they are well represented in the fossil record. The shells of snails are complex and grow at different speeds. The speed of growth is affected by a few variables such as the temperature of the water, depth of the water, food present for the snail, as well as isotopic oxygen levels. By looking at the composition of aragonite in the growth layers of
mollusks Mollusca is the second-largest phylum of invertebrate animals after the Arthropoda, the members of which are known as molluscs or mollusks (). Around 85,000  extant species of molluscs are recognized. The number of fossil species is esti ...
you can predict the size the mollusk shell can reach.


Taxonomy


2005 taxonomy

The following cladogram is an overview of the main clades of living gastropods based on the taxonomy of Bouchet & Rocroi (2005), with taxa that contain saltwater or brackish water species marked in boldface (some of the highlighted taxa consist entirely of marine species, but some of them also contain freshwater or land species.) * Clade Patellogastropoda * Clade Vetigastropoda * Clade
Cocculiniformia The Cocculinoidea is a superfamily of deepwater limpets (marine gastropods), the only superfamily in the order Cocculinida , one of the main orders of gastropods according to the taxonomy as set up by ( Bouchet & Rocroi, 2005). The clade Coccil ...
* Clade Neritimorpha ** Clade
Cycloneritimorpha Cycloneritida (nerites and false-limpets) is an order of land snails, freshwater snails, and sea snails.MolluscaBase eds. (2020). MolluscaBase. Cycloneritida. Accessed through: World Register of Marine Species at: http://www.marinespecies.org/ap ...
* Clade
Caenogastropoda Caenogastropoda is a taxonomic clade, a large diverse group which are mostly sea snails and other marine gastropod mollusks, but also includes some freshwater snails and some land snails. The clade is the most diverse and ecologically successfu ...
** Informal group
Architaenioglossa Architaenioglossa is a taxonomic group of snails which have gills and often an operculum. They are primarily land and freshwater gastropod mollusks within the clade Caenogastropoda. This " informal group" has been shown to be polyphyletic in a s ...
** Clade
Sorbeoconcha Sorbeoconcha is a taxonomic clade of snails, i.e. gastropods, mainly marine species with gills and opercula, within the clade Caenogastropoda. The taxon Sorbeoconcha was named by Winston Ponder and David R. Lindberg in 1997. Taxonomy 1997 taxo ...
** Clade
Hypsogastropoda Hypsogastropoda is a clade containing marine gastropods within the clade Caenogastropoda. MolluscaBase eds. (2021). MolluscaBase. Hypsogastropoda. Accessed through: World Register of Marine Species at: http://www.marinespecies.org/aphia.php?p=t ...
*** Clade Littorinimorpha *** Informal group
Ptenoglossa The Ptenoglossa is an informal taxonomic group of sea snails. This group was considered paraphyletic or polyphyletic by Ponder and Lindberg (1997) in their classification of gastropod molluscs. The database WoRMS considers this suborder as bel ...
*** Clade
Neogastropoda Neogastropoda is an order of sea snails, both freshwater and marine gastropod molluscs. Description The available fossil record of Neogastropoda is relatively complete, and supports a widely accepted evolutionary scenario of an Early Creta ...
* Clade Heterobranchia ** Informal group Lower Heterobranchia ** Informal group
Opisthobranchia Opisthobranchs () is now an informal name for a large and diverse group of specialized complex gastropods which used to be united in the subclass Opisthobranchia. That taxon is no longer considered to represent a monophyletic grouping. Euopisth ...
*** Clade Cephalaspidea *** Clade Thecosomata *** Clade Gymnosomata *** Clade Aplysiomorpha *** Group Acochlidiacea *** Clade Sacoglossa *** Group
Cylindrobullida ''Cylindrobulla'' is a genus of sea snails or bubble snails, marine gastropod mollusks in the clade Sacoglossa. ''Cylindrobulla'' is the type genus of the family Cylindrobullidae and it is the only genus in the family. Taxonomy The subord ...
*** Clade Umbraculida *** Clade Nudipleura **** Clade Pleurobranchomorpha **** Clade Nudibranchia ***** Clade Euctenidiacea ***** Clade Dexiarchia ****** Clade Pseudoeuctenidiacea ****** Clade Cladobranchia ******* Clade Euarminida ******* Clade Dendronotida ******* Clade Aeolidida ** Informal group
Pulmonata Pulmonata or pulmonates, is an informal group (previously an order, and before that a subclass) of snails and slugs characterized by the ability to breathe air, by virtue of having a pallial lung instead of a gill, or gills. The group includ ...
*** Informal group
Basommatophora Basommatophora was a term that was previously used as a taxonomic informal group, a group of snails within the informal group Pulmonata, the air-breathing slugs and snails. According to the taxonomy of the Gastropoda (Bouchet & Rocroi, 2005), w ...
*** Clade
Eupulmonata Eupulmonata is a taxonomic clade of air-breathing snails. The great majority of this group are land snails and slugs, but some are marine and some are saltmarsh snails that can tolerate salty conditions. Linnean taxonomy *Suborder Eupulmonata H ...
**** Clade
Systellommatophora The Systellommatophora (synonym Gymnomorpha) is a clade of primitive, air-breathing slugs, according to the taxonomy of the Gastropoda (Bouchet & Rocroi, 2005). They are marine and terrestrial pulmonate gastropods within the Heterobranchi ...
**** Clade
Stylommatophora Stylommatophora is an orderPhilippe Bouchet, Jean-Pierre Rocroi, Bernhard Hausdorf, Andrzej Kaim, Yasunori Kano, Alexander Nützel, Pavel Parkhaev, Michael Schrödl and Ellen E. Strong. 2017. Revised Classification, Nomenclator and Typification ...
***** Clade Elasmognatha ***** Clade Orthurethra ***** Informal group
Sigmurethra Sigmurethra is a taxonomic category of air-breathing land snails and slugs, terrestrial pulmonate gastropod molluscs. This is an informal group which includes most land snails and slugs. The two strong synapomorphies of Sigmurethra are a long p ...


Uses


By humans

A number of species of sea snails are harvested in
aquaculture Aquaculture (less commonly spelled aquiculture), also known as aquafarming, is the controlled cultivation ("farming") of aquatic organisms such as fish, crustaceans, mollusks, algae and other organisms of value such as aquatic plants (e.g. lot ...
and used by humans for food, including
abalone Abalone ( or ; via Spanish , from Rumsen ''aulón'') is a common name for any of a group of small to very large marine gastropod molluscs in the family (biology), family Haliotidae. Other common name In biology, a common name of a taxon o ...
,
conch Conch () is a common name of a number of different medium-to-large-sized sea snails. Conch shells typically have a high spire and a noticeable siphonal canal (in other words, the shell comes to a noticeable point at both ends). In North Am ...
,
limpet Limpets are a group of aquatic snails that exhibit a conical shell shape (patelliform) and a strong, muscular foot. Limpets are members of the class Gastropoda, but are polyphyletic, meaning the various groups called "limpets" descended indep ...
s, whelks (such as the North American ''
Busycon ''Busycon'' is a genus of very large edible sea snails in the subfamily Busyconinae. These snails are commonly known in the United States as ''whelks'' or ''Busycon whelks''. Less commonly they are loosely, and somewhat misleadingly, called "conc ...
'' species and the North Atlantic ''
Buccinum undatum ''Buccinum undatum'', the common whelk or the waved buccinum, is a large, edible marine gastropod in the family Buccinidae, the "true whelks".Fraussen, K.; Gofas, S. (2014). Buccinum undatum Linnaeus, 1758. Accessed through: World Register of M ...
'') and periwinkles including '' Littorina littorea''. The shells of sea snails are often found washed up on beaches. Because many are attractive and durable, they have been used to make necklaces and other jewelry since prehistoric times. The shells of a few species of large sea snails within the Vetigastropoda have a thick layer of
nacre Nacre ( , ), also known as mother of pearl, is an organicinorganic composite material produced by some molluscs as an inner shell layer; it is also the material of which pearls are composed. It is strong, resilient, and iridescent. Nacre is f ...
and have been used as a source of mother of pearl. Historically, the button industry relied on these species for a number of years.


By non-human animals

The shells of sea snails are used for protection by many kinds of hermit crabs. A hermit crab carries the shell by grasping the central columella of the shell using claspers on the tip of its abdomen.


See also

*
Freshwater snail Freshwater snails are gastropod mollusks which live in fresh water. There are many different families. They are found throughout the world in various habitats, ranging from ephemeral pools to the largest lakes, and from small seeps and springs ...
* Terrestrial molluscs *
Land snail A land snail is any of the numerous species of snail that live on land, as opposed to the sea snails and freshwater snails. ''Land snail'' is the common name for terrestrial gastropod mollusks that have shells (those without shells are known as ...
* Sea slug *
Slug Slug, or land slug, is a common name for any apparently shell-less terrestrial gastropod mollusc. The word ''slug'' is also often used as part of the common name of any gastropod mollusc that has no shell, a very reduced shell, or only a smal ...


References

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