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The sand goanna (''Varanus gouldii)'', also known commonly as Gould's monitor, the racehorse goanna, and the sand monitor, is a
species A species () is often defined as the largest group of organisms in which any two individuals of the appropriate sexes or mating types can produce fertile offspring, typically by sexual reproduction. It is the basic unit of Taxonomy (biology), ...
of large
Australia Australia, officially the Commonwealth of Australia, is a country comprising mainland Australia, the mainland of the Australia (continent), Australian continent, the island of Tasmania and list of islands of Australia, numerous smaller isl ...
n monitor
lizard Lizard is the common name used for all Squamata, squamate reptiles other than snakes (and to a lesser extent amphisbaenians), encompassing over 7,000 species, ranging across all continents except Antarctica, as well as most Island#Oceanic isla ...
in the
family Family (from ) is a Social group, group of people related either by consanguinity (by recognized birth) or Affinity (law), affinity (by marriage or other relationship). It forms the basis for social order. Ideally, families offer predictabili ...
Varanidae The Varanidae are a family of lizards in the superfamily Varanoidea and order Anguimorpha. The family, a group of carnivorous and frugivorous lizards, includes the living genus '' Varanus'' and a number of extinct genera more closely related ...
.


Taxonomy

John Edward Gray John Edward Gray (12 February 1800 – 7 March 1875) was a British zoologist. He was the elder brother of zoologist George Robert Gray and son of the pharmacologist and botanist Samuel Frederick Gray (1766–1828). The same is used for a z ...
described the species in 1838 as ''Hydrosaurus gouldii'', noting the source of the type specimen as " New Holland" and distinguishing the new varanid by "two yellow streaks on the sides of the neck" and small flat scales at the orbits. An earlier description, ''Tupinambis endrachtensis'' Péron, F. 1807, was determined as likely to refer to this animal, but the epithet ''gouldii'' was conserved and a new specimen designated as the type. This neotype was obtained in 1997 at the near coastal Western Australian suburb of Karrakatta, and placed with the
British Museum of Natural History The Natural History Museum in London is a museum that exhibits a vast range of specimens from various segments of natural history. It is one of three major museums on Exhibition Road in South Kensington, the others being the Science Museum (Lo ...
. The decision of a nomenclatural commission (
ICZN The International Code of Zoological Nomenclature (ICZN) is a widely accepted convention in zoology that rules the formal scientific naming of organisms treated as animals. It is also informally known as the ICZN Code, for its formal author, t ...
) was to issue an opinion suppressing the earlier name ''Tupinambis endrachtensis'' and the name ''Hydrosaurus ocellarius'' Blyth, 1868, that were unsatisfactory to some who had commented on the case, but provided taxonomic certainty for future revisions of the associated taxa.


Etymology

The
specific name Specific name may refer to: * in Database management systems, a system-assigned name that is unique within a particular database In taxonomy, either of these two meanings, each with its own set of rules: * Specific name (botany), the two-part (bino ...
, ''gouldii'', is assumed to be a Latinised form of the surname of an associate of the describing author, the English
ornithologist Ornithology, from Ancient Greek ὄρνις (''órnis''), meaning "bird", and -logy from λόγος (''lógos''), meaning "study", is a branch of zoology dedicated to the study of birds. Several aspects of ornithology differ from related discip ...
John Gould John Gould (; 14 September 1804 – 3 February 1881) was an English ornithologist who published monographs on birds, illustrated by plates produced by his wife, Elizabeth Gould (illustrator), Elizabeth Gould, and several other artists, includ ...
, who was actively assembling specimens of fauna from Australia but is not thought to have any direct connection to this species. In some Aboriginal languages, the sand goanna is called ''bungarra'', a term also commonly used by non-Aboriginal people in
Western Australia Western Australia (WA) is the westernmost state of Australia. It is bounded by the Indian Ocean to the north and west, the Southern Ocean to the south, the Northern Territory to the north-east, and South Australia to the south-east. Western Aust ...
. In Pitjantjatjara and other central Australian languages it is called "Tingka".


Subspecies

Two subspecies are recognised, *'' Varanus gouldii gouldii'' – Gould's goanna *'' Varanus gouldii flavirufus'' – desert sand monitor ''
Nota bene ( ; plural: ) is the Latin language, Latin phrase meaning ''note well''. In manuscripts, ''nota bene'' is abbreviated in upper-case as NB and N.B., and in lower-case as n.b. and nb; the editorial usages of ''nota bene'' and ''notate bene'' fi ...
'': A trinomial authority in parentheses indicates that the subspecies was originally described in a
genus Genus (; : genera ) is a taxonomic rank above species and below family (taxonomy), family as used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In bino ...
other than ''Varanus''.


Description

A species of '' Varanus'', lizards known as monitors and goannas. Due to the taxonomic uncertainty during the twentieth century the species form and behaviour has included taxa later recognised as distinct species, this includes '' V. rosenbergi'', formerly treated as a subspecies and later elevated, and '' V. panoptes'', described as a new species in 1980 and resolved as a legitimate publication in 2000.


Biology

Gould's monitor is a terrestrial reptile that excavates large
burrow file:Chipmunk-burrow (exits).jpg, An eastern chipmunk at the entrance of its burrow A burrow is a hole or tunnel excavated into the ground by an animal to construct a space suitable for habitation or temporary refuge, or as a byproduct of Animal lo ...
s for shelter. Rock escarpments and tree hollows are also suitable dwellings. It inhabits a vast range throughout Australia, reaching an average length of and weighing as much as . It can be found in northern and eastern Australia, where they inhabit open
woodland A woodland () is, in the broad sense, land covered with woody plants (trees and shrubs), or in a narrow sense, synonymous with wood (or in the U.S., the '' plurale tantum'' woods), a low-density forest forming open habitats with plenty of sunli ...
s and
grassland A grassland is an area where the vegetation is dominance (ecology), dominated by grasses (Poaceae). However, sedge (Cyperaceae) and rush (Juncaceae) can also be found along with variable proportions of legumes such as clover, and other Herbaceo ...
s. ''V. g. flavirufus'', a slightly smaller
subspecies In Taxonomy (biology), biological classification, subspecies (: subspecies) is a rank below species, used for populations that live in different areas and vary in size, shape, or other physical characteristics (Morphology (biology), morpholog ...
, resides in Australia's interior. In some places, however, the ranges of Gould's monitor, ''V. g. flavirufus'' and the Argus monitor overlap. The similarities between the species and their close proximity frequently cause confusion. The sand monitor is a relentless
forager A forager is a person who collects edible plants or fungi for consumption. Urban foragers may collect in city parks, private lands, and sidewalks. Urban foraging has gained in popularity in the 21st century, as people share their knowledge, exper ...
. It is diurnal, meaning most of its activities take place during the day. Anything smaller than itself will be eagerly devoured. The diets of hatchlings and juveniles often consist mostly of
insect Insects (from Latin ') are Hexapoda, hexapod invertebrates of the class (biology), class Insecta. They are the largest group within the arthropod phylum. Insects have a chitinous exoskeleton, a three-part body (Insect morphology#Head, head, ...
s and small
lizard Lizard is the common name used for all Squamata, squamate reptiles other than snakes (and to a lesser extent amphisbaenians), encompassing over 7,000 species, ranging across all continents except Antarctica, as well as most Island#Oceanic isla ...
s, but generally varies more with age. Adult monitors will
prey Predation is a biological interaction in which one organism, the predator, kills and eats another organism, its prey. It is one of a family of common feeding behaviours that includes parasitism and micropredation (which usually do not ki ...
on
mice A mouse (: mice) is a small rodent. Characteristically, mice are known to have a pointed snout, small rounded ears, a body-length scaly tail, and a high breeding rate. The best known mouse species is the common house mouse (''Mus musculus' ...
, other small
mammal A mammal () is a vertebrate animal of the Class (biology), class Mammalia (). Mammals are characterised by the presence of milk-producing mammary glands for feeding their young, a broad neocortex region of the brain, fur or hair, and three ...
s, small
bird Birds are a group of warm-blooded vertebrates constituting the class (biology), class Aves (), characterised by feathers, toothless beaked jaws, the Oviparity, laying of Eggshell, hard-shelled eggs, a high Metabolism, metabolic rate, a fou ...
s,
crustacean Crustaceans (from Latin meaning: "those with shells" or "crusted ones") are invertebrate animals that constitute one group of arthropods that are traditionally a part of the subphylum Crustacea (), a large, diverse group of mainly aquatic arthrop ...
s, scorpions, large insects, small lizard species,
snake Snakes are elongated limbless reptiles of the suborder Serpentes (). Cladistically squamates, snakes are ectothermic, amniote vertebrates covered in overlapping scales much like other members of the group. Many species of snakes have s ...
s, amphibians, eggs, and carrion. They have also been seen to congregate around
fish A fish (: fish or fishes) is an aquatic animal, aquatic, Anamniotes, anamniotic, gill-bearing vertebrate animal with swimming fish fin, fins and craniate, a hard skull, but lacking limb (anatomy), limbs with digit (anatomy), digits. Fish can ...
kills. They are seemingly immune to snake venom and will kill and eat even the most venomous species such as the
inland taipan The inland taipan (''Oxyuranus microlepidotus''), also Common name, commonly known as the western taipan, small-scaled snake, or fierce snake,White, Julian (November 1991)''Oxyuranus microlepidotus'' "Neurotoxic paralysis usually takes 2-4 hours ...
. The sand goanna does consume smaller species of monitors: Ackies, rock monitors and other dwarf species are often found and eaten. It is common to see a Gould's monitor disturbing rock piles in an attempt to flush out any '' Odatria''. It lays its
egg An egg is an organic vessel grown by an animal to carry a possibly fertilized egg cell (a zygote) and to incubate from it an embryo within the egg until the embryo has become an animal fetus that can survive on its own, at which point the ...
s in termite mounds to protect them from the harsh
desert A desert is a landscape where little precipitation occurs and, consequently, living conditions create unique biomes and ecosystems. The lack of vegetation exposes the unprotected surface of the ground to denudation. About one-third of the la ...
climate. Like all goannas and
snake Snakes are elongated limbless reptiles of the suborder Serpentes (). Cladistically squamates, snakes are ectothermic, amniote vertebrates covered in overlapping scales much like other members of the group. Many species of snakes have s ...
s the sand monitor has a forked tongue, with which it "tastes" the air. Scents are picked up on each side of the fork which, when retracted, brushes over an area in the mouth called the
Jacobson's organ The vomeronasal organ (VNO), or Jacobson's organ, is the paired auxiliary olfactory (smell) sense organ located in the soft tissue of the nasal septum, in the nasal cavity just above the roof of the mouth (the hard palate) in various tetrapods. T ...
, which identifies on which side the scent is stronger. Upon identifying this difference in scent strength the sand monitor can alter its direction accordingly. A 2021 study had shown that during the
Miocene The Miocene ( ) is the first epoch (geology), geological epoch of the Neogene Period and extends from about (Ma). The Miocene was named by Scottish geologist Charles Lyell; the name comes from the Greek words (', "less") and (', "new") and mea ...
,
Komodo dragon The Komodo dragon (''Varanus komodoensis''), also known as the Komodo monitor, is a large reptile of the monitor lizard family Varanidae that is endemic to the Indonesian islands of Komodo (island), Komodo, Rinca, Flores, Gili Dasami, and Gili ...
s had hybridized with the
common ancestor Common descent is a concept in evolutionary biology applicable when one species is the ancestor of two or more species later in time. According to modern evolutionary biology, all living beings could be descendants of a unique ancestor commonl ...
of ''V. gouldii'', '' V. rosenbergi'', '' V. panoptes'', and '' V. spenceri'', thus providing evidence that the Komodo dragon had once inhabited Australia.


Gallery

File:Sand monitor walking tracks in sand.jpg, Sand monitor walking tracks in sand. File:Varanus gouldii at Sunrise Dam Gold Mine.jpg, At the Sunrise Dam Gold Mine (WA). File:Varanus gouldii - Chace ranges.JPG, At
Wilpena Pound Wilpena Pound ("Ikara" in the Adnyamathanha language) is a major natural amphitheatre of mountains located north of Adelaide, South Australia, Australia in the heart of the Ikara-Flinders Ranges National Park. Its fringe is accessible by a sea ...
(SA) File:Sand Goanna on road (15734829198).jpg, Mungo National Park (NSW) File:Varanus gouldii camera lure.JPG, At a camera lure, Angas Downs IPA (NT)


Citations


General sources

* 226 pp. * * *


External links


Photo of ''Varanus gouldii'' at AROD.com.au

Photo of ''Varanus gouldii'' at Pbase.com
{{Taxonbar, from=Q671373 Varanus Reptiles described in 1838 Taxa named by John Edward Gray Goanna, sand Reptiles of Queensland Reptiles of New South Wales Monitor lizards of Australia