''Samotherium'' ("beast of

Samos Samos (, also ; , ) is a Greek island in the eastern Aegean Sea, south of Chios, north of Patmos and the Dodecanese archipelago, and off the coast of western Turkey, from which it is separated by the Mycale Strait. It is also a separate reg ...

") is an extinct genus of

Giraffidae The Giraffidae are a family (biology), family of ruminant artiodactyl mammals that share a recent common ancestor with deer and bovids. This family, once a diverse group spread throughout Eurasia and Africa, presently comprises only two extant ge ...

from the

Miocene The Miocene ( ) is the first epoch (geology), geological epoch of the Neogene Period and extends from about (Ma). The Miocene was named by Scottish geologist Charles Lyell; the name comes from the Greek words (', "less") and (', "new") and mea ...

and

Pliocene The Pliocene ( ; also Pleiocene) is the epoch (geology), epoch in the geologic time scale that extends from 5.33 to 2.58Eurasia Eurasia ( , ) is a continental area on Earth, comprising all of Europe and Asia. According to some geographers, Physical geography, physiographically, Eurasia is a single supercontinent. The concept of Europe and Asia as distinct continents d ...

and

Africa Africa is the world's second-largest and second-most populous continent after Asia. At about 30.3 million km2 (11.7 million square miles) including adjacent islands, it covers 20% of Earth's land area and 6% of its total surfac ...

. ''Samotherium'' had two

ossicone Ossicones are columnar or conical skin-covered bone structures on the heads of giraffes, male okapi, and some of their extinct relatives. Ossicones are distinguished from the superficially similar structures of Horn (anatomy), horns and antlers ...

s on its head, and long

legs A leg is a weight-bearing and locomotive anatomical structure, usually having a columnar shape. During locomotion, legs function as "extensible struts". The combination of movements at all joints can be modeled as a single, linear element cap ...

. The ossicones usually pointed upward, and were curved backwards, with males having larger, more curved ossicones, though in the Chinese species, ''S. sinense'', the straight ossicones point laterally, not upwards. The genus is closely related to '' Shansitherium''. Fossil evidence suggests that ''Samotherium'' had a rounded muzzle, which would suggest a grazing lifestyle and a habitat composed of grassland. One common predator of this animal was the ''

Amphimachairodus ''Amphimachairodus'' is an extinct genus of large machairodonts. It is also a member of the tribe Homotherini within Machairodontinae and is most closely related to genera as '' Xenosmilus'', ''Homotherium'' itself, and ''Machairodus''. It inhab ...

''. Biologist Richard Ellis has proposed that the skull of ''Samotherium'' is portrayed on an ancient Greek vase as a monster that

Heracles Heracles ( ; ), born Alcaeus (, ''Alkaios'') or Alcides (, ''Alkeidēs''), was a Divinity, divine hero in Greek mythology, the son of ZeusApollodorus1.9.16/ref> and Alcmene, and the foster son of Amphitryon.By his adoptive descent through ...

is fighting. However, other authors have argued that it is more likely to be the skull of a

monitor lizard Monitor lizards are lizards in the genus ''Varanus,'' the only extant genus in the family Varanidae. They are native to Africa, Asia, and Oceania, and West African Nile monitor, one species is also found in south America as an invasive species. A ...

instead.

Description

A 2015 study found that ''Samotherium'' had a neck intermediate in length between the

giraffe The giraffe is a large Fauna of Africa, African even-toed ungulate, hoofed mammal belonging to the genus ''Giraffa.'' It is the Largest mammals#Even-toed Ungulates (Artiodactyla), tallest living terrestrial animal and the largest ruminant on ...

and the

okapi The okapi (; ''Okapia johnstoni''), also known as the forest giraffe, Congolese giraffe and zebra giraffe, is an artiodactyl mammal that is endemic to the northeast Democratic Republic of the Congo in central Africa. However, non-invasive gen ...

, judging from examination of specimens of ''S. major'' from Greece.

References

Prehistoric giraffes Prehistoric Artiodactyla genera Miocene Artiodactyla Pliocene Artiodactyla Neogene mammals of Asia Transitional fossils Fossil taxa described in 1888 Taxa named by Charles Immanuel Forsyth Major {{paleo-eventoedungulate-stub