Rødvig Formation
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The Rødvig Formation is a geological formation deposited during the earliest part of the
Danian The Danian is the oldest age or lowest stage of the Paleocene Epoch or Series, of the Paleogene Period or System, and of the Cenozoic Era or Erathem. The beginning of the Danian (and the end of the preceding Maastrichtian) is at the Cretac ...
(early
Paleocene The Paleocene ( ), or Palaeocene, is a geological epoch (geology), epoch that lasted from about 66 to 56 mya (unit), million years ago (mya). It is the first epoch of the Paleogene Period (geology), Period in the modern Cenozoic Era (geology), ...
; ) and it was first identified by Richard Taylor and Richard Phillips in 1827. It is known from exposures at
Stevns Klint Stevns Klint, known as the Cliffs of Stevns in English, is a white chalk cliff located some southeast of Store Heddinge on the Danish island of Zealand. Stretching along the coast, it is of geological importance as one of the best exposed C ...
in Denmark. The unit lies directly above the K–Pg boundary and contains fossils that provide a record of the recovery of various groups following the Cretaceous–Paleogene extinction. The upper boundary of the formation is an unconformity in the form of a
hardground Carbonate hardgrounds are surfaces of synsedimentarily cemented carbonate layers that have been exposed on the seafloor (Wilson and Palmer, 1992). A hardground is essentially, then, a lithified seafloor. Ancient hardgrounds are found in limeston ...
, beneath which the formation is sometimes missing. The base of the unit is irregular due to the presence of mounding associated with
bryozoa Bryozoa (also known as the Polyzoa, Ectoprocta or commonly as moss animals) are a phylum of simple, aquatic animal, aquatic invertebrate animals, nearly all living in sedentary Colony (biology), colonies. Typically about long, they have a spe ...
, causing variations in thickness. The unit is subdivided into the lower ''Fiskeler Member'' mainly formed of
marl Marl is an earthy material rich in carbonate minerals, Clay minerals, clays, and silt. When Lithification, hardened into rock, this becomes marlstone. It is formed in marine or freshwater environments, often through the activities of algae. M ...
and the overlying ''Cerithium Limestone Member''.


Geology

The dark layer of , mainly five to ten centimeters thick, clearly marks the
Cretaceous–Paleogene boundary The Cretaceous–Paleogene (K–Pg) boundary, formerly known as the Cretaceous–Tertiary (K–T) boundary, is a geological signature, usually a thin band of rock containing much more iridium than other bands. The K–Pg boundary marks the end o ...
and overlies the
Maastrichtian The Maastrichtian ( ) is, in the International Commission on Stratigraphy (ICS) geologic timescale, the latest age (geology), age (uppermost stage (stratigraphy), stage) of the Late Cretaceous epoch (geology), Epoch or Upper Cretaceous series (s ...
age
Tor Formation The Tor Formation is a geological formation of late Campanian to Maastrichtian (uppermost Cretaceous) age. It forms the part of the Chalk Group in the North Sea. It is an important reservoir for oil and gas in fields such as Valhall. It overlies ...
. The is enriched in
iridium Iridium is a chemical element; it has the symbol Ir and atomic number 77. This very hard, brittle, silvery-white transition metal of the platinum group, is considered the second-densest naturally occurring metal (after osmium) with a density ...
, a fact used as an argument for the
Alvarez hypothesis The Alvarez hypothesis posits that the Extinction event, mass extinction of the non-avian dinosaurs and many other living things during the Cretaceous–Paleogene extinction event was caused by the Impact event, impact of a large asteroid on the Ea ...
that the worldwide Cretaceous–Paleogene mass extinction was caused by the impact of an asteroid. Following the boundary is a layer of darker clay and chalk between 10 and 30 cm thick, corresponding to a period of low biological diversity on the sea floor immediately after the K-Pg Boundary.


Paleontology

The Rødvig Formation contains a remarkably detailed and complete fossil record of the biota in Northern Europe during the early Paleogene. The layers are rich in microfossils, containing many species of millimeter-long suspension feeders. A wide variety of
benthic The benthic zone is the ecological region at the lowest level of a body of water such as an ocean, lake, or stream, including the sediment surface and some sub-surface layers. The name comes from the Ancient Greek word (), meaning "the depths". ...
foraminifera Foraminifera ( ; Latin for "hole bearers"; informally called "forams") are unicellular organism, single-celled organisms, members of a phylum or class (biology), class of Rhizarian protists characterized by streaming granular Ectoplasm (cell bio ...
species have been identified from the Rødvig Formation, with significant differences in abundance across the K–Pg boundary demonstrating the biotic turnover that occurred during the mass extinction event.


Ammonites

The Rødvig Formation was the first known site to document the short-term survival of ammonites into the
Paleogene The Paleogene Period ( ; also spelled Palaeogene or Palæogene) is a geologic period and system that spans 43 million years from the end of the Cretaceous Period Ma (million years ago) to the beginning of the Neogene Period Ma. It is the fir ...
, when they were originally thought to have gone extinct at the K-Pg boundary. Ammonites are thus only known from the first 200,000 years of the Cerithium Limestone, before disappearing from the formation around 64.8 million years ago. Two ammonite species are known from the Rødvig Formation: '' Baculites vertebralis'', notable for having a nearly straight shell, and '' Hoploscaphites constrictus'', which may have been a subspecies of ''
Scaphites ''Scaphites'' is a genus of heteromorph ammonites belonging to the Scaphitidae family. They were a widespread genus that thrived during the Late Cretaceous period. Morphology ''Scaphites'' generally have a chambered, boat-shaped shell. The ini ...
'',''Hoploscaphites''
at
Fossilworks Fossilworks was a portal which provides query, download, and analysis tools to facilitate access to the Paleobiology Database, a large relational database assembled by hundreds of paleontologists from around the world. History Fossilworks was cr ...
.org
and is also the most common ammonite known from the
Paleocene The Paleocene ( ), or Palaeocene, is a geological epoch (geology), epoch that lasted from about 66 to 56 mya (unit), million years ago (mya). It is the first epoch of the Paleogene Period (geology), Period in the modern Cenozoic Era (geology), ...
.


Known fossil fauna

The known fossil fauna found in the Rødvig Formation includes: * ''
Acar Acar is a type of vegetable pickle of Maritime Southeast Asia, most prevalent in Indonesia, Malaysia, Singapore and Brunei. It is a localised version of Indian '' achar''. It is known as atjar in Dutch cuisine, derived from Indonesian ''acar' ...
'' sp. *
Arcida The Arcida is an Extant taxon, extant order (biology), order of bivalve molluscs. This order dates back to the lower Ordovician period. They are distinguished from related groups, such as the mussels, by having a straight hinge to the shells, a ...
indet. * '' Arcopsis christinae'' * '' Arcopsis'' sp. * '' Baculites vertebralis'' * '' Barbatia'' sp. * '' Bathyarca'' sp. *
Bivalvia Bivalvia () or bivalves, in previous centuries referred to as the Lamellibranchiata and Pelecypoda, is a class (biology), class of aquatic animal, aquatic molluscs (marine and freshwater) that have laterally compressed soft bodies enclosed b ...
indet. * ''
Brachidontes ''Brachidontes'' is a genus of mussels in the family Mytilidae The Mytilidae are a family (biology), family of small to large Marine life, marine and Brackish water, brackish-water bivalve molluscs in the order (biology), order Mytilida. One o ...
'' sp. * ''Carcharias'' aff. ''gracilis'' * '' Centroscymnus praecursor'' * '' Chlamydoselachus'' sp. * '' Cuspidaria ( Halonympha) kanae'' * '' Corbulamella'' sp. * '' Crassatella'' sp. * '' Crassescyliorhinus germanicus'' * ''
Cretalamna ''Cretalamna'' is a genus of extinction, extinct Otodontidae, otodontid shark that lived from the latest Early Cretaceous to Eocene epoch (about 103 to 46 million years ago). It is considered by many to be the ancestor of the largest sharks to ha ...
'' sp. * '' Cuspidaria'' sp. * '' Cyclaster danicus'' * '' Cyrtodaria'' sp. * ''
Dacrydium ''Dacrydium'' is a genus of conifers belonging to the podocarp family Podocarpaceae. Sixteen species of evergreen Plant sexuality, dioecious trees and shrubs are presently recognized. The genus was first described by Solander in 1786, and former ...
'' sp. * ''
Echinorhinus ''Echinorhinus'' is the only extant genus in the family Echinorhinidae. Taxonomy Echinorhinidae are traditionally classified in the order Squaliformes, together with kitefin and gulper sharks.Compagno, 2005. "Sharks of the World". However, a ...
'' sp. * '' Eriphylopsis'' sp. * '' Euciroa'' sp. * '' Hemiscyllium hermani'' * '' Heterodontus rugosus'' * '' Hoploscaphites constrictus'' * '' Gregariella'' sp. * '' Leptosolen'' sp. * '' Limopsis ravni'' * '' Limopsis'' sp. * '' Loripes'' sp. *
Lucinidae Lucinidae, common name hatchet shells, is a family of saltwater clams, marine bivalve molluscs. These bivalves are remarkable for their endosymbiosis with sulphide-oxidizing bacteria. Characteristics The members of this family have a worldwid ...
indet. * '' Martesia'' sp. * '' Miocardiopsis'' sp. * '' Myrtea'' sp. * ''
Nebrius ''Nebrius'' is a genus of carpet sharks in the family Ginglymostomatidae. It contains a single Extant taxon, extant species, the tawny nurse shark (''Nebrius ferrugineus''), as well as a number of extinct species dating back to the Early Paleoce ...
'' sp. * '' Nielonella'' sp. * '' Palaeocypraea spirata'' * ''Palaeogaleus'' cf. ''faujasi'' * ''Palaeohypotodus'' aff. ''bronni'' * '' Parasquatina cappettai'' * '' Paratriakis curtirostris'' * ''
Portlandia arctica ''Portlandia'' is an American sketch comedy television series starring Fred Armisen and Carrie Brownstein, set in and around Portland, Oregon, and spoofing the city's reputation as a haven for eccentric hipsters. The show was produced by Broa ...
'' * '' Protocardia'' sp. * '' Pycnodonte vesicularis'' (='' Phygraea vesiculare'') * '' “Scyliorhinus” biddlei'' * '' “Scyliorhinus” elongatus'' * ''Spondylus fimbriatus'' * '' Squalus gabrielsoni'' * '' Syncylonema nilsoni'' * '' Synechodus faxensis'' (='' Paleospinax'') * ''
Thalassinoides ''Thalassinoides'' is an ichnogenus of trace fossil (fossil records of lifeforms' movement, rather than of the lifeforms themselves) used to refer to "dichotomously or T-branched boxworks, mazes and shafts, unlined and unornamented". Facies of ' ...
'' sp. * ''
Thyasira ''Thyasira'' is a genus of saltwater clams, marine bivalve mollusks in the family Thyasiridae. Species There are more than 50 species in the genus, including both extant (living) and extinct fossil species: * ''Thyasira gouldi'' * ''Thyasira ...
'' sp. * '' Tylocidaris oedumi'' * '' Uddenia'' sp. * '' Unicordium'' sp, * '' Vetericardiella'' sp. * '' Yoldiella'' sp.


References

{{reflist, group=lower-alpha Danian Danian Stage Geology of Denmark