Ruderal
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A ruderal species is a
plant Plants are the eukaryotes that form the Kingdom (biology), kingdom Plantae; they are predominantly Photosynthesis, photosynthetic. This means that they obtain their energy from sunlight, using chloroplasts derived from endosymbiosis with c ...
species A species () is often defined as the largest group of organisms in which any two individuals of the appropriate sexes or mating types can produce fertile offspring, typically by sexual reproduction. It is the basic unit of Taxonomy (biology), ...
that is first to colonize disturbed lands. The disturbance may be natural for example,
wildfire A wildfire, forest fire, or a bushfire is an unplanned and uncontrolled fire in an area of Combustibility and flammability, combustible vegetation. Depending on the type of vegetation present, a wildfire may be more specifically identified as a ...
s or
avalanche An avalanche is a rapid flow of snow down a Grade (slope), slope, such as a hill or mountain. Avalanches can be triggered spontaneously, by factors such as increased precipitation or snowpack weakening, or by external means such as humans, othe ...
sor the consequences of human activities, such as construction ( of roads, of buildings,
mining Mining is the Resource extraction, extraction of valuable geological materials and minerals from the surface of the Earth. Mining is required to obtain most materials that cannot be grown through agriculture, agricultural processes, or feasib ...
, etc.) or agriculture (abandoned fields,
irrigation Irrigation (also referred to as watering of plants) is the practice of applying controlled amounts of water to land to help grow crops, landscape plants, and lawns. Irrigation has been a key aspect of agriculture for over 5,000 years and has bee ...
, etc.). The term '' ruderal'' originates from the Latin word '' rudus'', meaning " rubble". Ruderal species typically dominate the disturbed area for a few years, gradually losing the
competition Competition is a rivalry where two or more parties strive for a common goal which cannot be shared: where one's gain is the other's loss (an example of which is a zero-sum game). Competition can arise between entities such as organisms, indi ...
to other native species. However, in extreme disturbance circumstances, such as when the natural
topsoil Topsoil is the upper layer of soil. It has the highest concentration of organic matter and microorganisms and is where most of the Earth's biological soil activity occurs. Description Topsoil is composed of mineral particles and organic mat ...
is covered with a foreign substance, a single-species ruderal community may become permanently established. In addition, some ruderal
invasive species An invasive species is an introduced species that harms its new environment. Invasive species adversely affect habitats and bioregions, causing ecological, environmental, and/or economic damage. The term can also be used for native spec ...
may have such a competitive advantage over the native species that they, too, may permanently prevent a disturbed area from returning to its original state despite natural topsoil.


Features

Features contributing to a species' success as ruderal are: * Massive seed production *
Seedling A seedling is a young sporophyte developing out of a plant embryo from a seed. Seedling development starts with germination of the seed. A typical young seedling consists of three main parts: the radicle (embryonic root), the hypocotyl (embry ...
s whose nutritional requirements are modest * Fast-growing roots * Independence of mycorrhizae * Polyploidy


Quantification

Ecologists have proposed various scales for quantifying ruderality, which can be defined as the "ability to thrive where there is disturbance through partial or total destruction of plant biomass" (Grime, Hodgson & Hunt, 1988). The ruderality scale of Grime presents values that are readily available, and it takes into account disturbance factors as well as other indicators such as the annual or perennial character of the plants.


See also

* Edge effect * Hemeroby * Hemerochory * Adventive plant * Pioneer species * Restoration ecology * Supertramp (ecology) * Examples of ruderal species: ** '' Cannabis ruderalis'' (family
Cannabaceae Cannabaceae is a small family of flowering plants, known as the hemp family. As now circumscribed, the family includes about 170 species grouped in about 11 genera, including '' Cannabis'' (hemp), '' Humulus'' (hops) and '' Celtis'' (hackberri ...
) ** '' Conyza bonariensis'' (family
Asteraceae Asteraceae () is a large family (biology), family of flowering plants that consists of over 32,000 known species in over 1,900 genera within the Order (biology), order Asterales. The number of species in Asteraceae is rivaled only by the Orchi ...
) ** '' Dittrichia viscosa'' (
Asteraceae Asteraceae () is a large family (biology), family of flowering plants that consists of over 32,000 known species in over 1,900 genera within the Order (biology), order Asterales. The number of species in Asteraceae is rivaled only by the Orchi ...
) ** '' Nicotiana glauca'' ( Solanaceae)


References


External links

*
St. John TV. 1987. SOIL DISTURBANCE AND THE MINERAL NUTRITION OF NATIVE PLANTS in Proceedings of the 2nd Native Plant Revegetation Symposium

Chapin. FS. III. 1980. The mineral nutrition of wild plants. Annu. Rev. Ecol. Syst., 11:233–260.
*{{Cite EB1911 , wstitle=Ruderal , volume=23 , page=814 , short=1 Ecology terminology Habitat Botany