''Retrophyllum vitiense'' is a species of
conifer
Conifers are a group of cone-bearing seed plants, a subset of gymnosperms. Scientifically, they make up the division Pinophyta (), also known as Coniferophyta () or Coniferae. The division contains a single extant class, Pinopsida. All ex ...
in the family
Podocarpaceae
Podocarpaceae is a large family of mainly Southern Hemisphere conifers, known in English as podocarps, comprising about 156 species of evergreen trees and shrubs.James E. Eckenwalder. 2009. ''Conifers of the World''. Portland, Oregon: Timber Pre ...
. It is a large
evergreen
In botany, an evergreen is a plant which has foliage that remains green and functional through more than one growing season. This also pertains to plants that retain their foliage only in warm climates, and contrasts with deciduous plants, which ...
rainforest emergent
tree
In botany, a tree is a perennial plant with an elongated stem, or trunk, usually supporting branches and leaves. In some usages, the definition of a tree may be narrower, including only woody plants with secondary growth, plants that are ...
native to
Fiji,
Indonesia
Indonesia, officially the Republic of Indonesia, is a country in Southeast Asia and Oceania between the Indian and Pacific oceans. It consists of over 17,000 islands, including Sumatra, Java, Sulawesi, and parts of Borneo and New Guine ...
,
Papua New Guinea
Papua New Guinea (abbreviated PNG; , ; tpi, Papua Niugini; ho, Papua Niu Gini), officially the Independent State of Papua New Guinea ( tpi, Independen Stet bilong Papua Niugini; ho, Independen Stet bilong Papua Niu Gini), is a country i ...
, and
Solomon Islands
Solomon Islands is an island country consisting of six major islands and over 900 smaller islands in Oceania, to the east of Papua New Guinea and north-west of Vanuatu. It has a land area of , and a population of approx. 700,000. Its ca ...
.
Description
''Retrophyllum vitiense'' is a large evergreen tree. The
trunk
Trunk may refer to:
Biology
* Trunk (anatomy), synonym for torso
* Trunk (botany), a tree's central superstructure
* Trunk of corpus callosum, in neuroanatomy
* Elephant trunk, the proboscis of an elephant
Computing
* Trunk (software), in rev ...
is usually erect, straight and terete. Large trees frequently feature a bole clear of branches for the first 20 meters and a buttressed trunk base.
[ The ]crown
A crown is a traditional form of head adornment, or hat, worn by monarchs as a symbol of their power and dignity. A crown is often, by extension, a symbol of the monarch's government or items endorsed by it. The word itself is used, partic ...
of a young plant is often pyramidal while older trees generally have rounded or spreading crowns. Branch
A branch, sometimes called a ramus in botany, is a woody structural member connected to the central trunk of a tree (or sometimes a shrub). Large branches are known as boughs and small branches are known as twigs. The term '' twig'' usually ...
es are ascending or spreading in the upper parts of the crown, but pendulous in the shaded parts.[ The initially brown and smooth ]bark
Bark may refer to:
* Bark (botany), an outer layer of a woody plant such as a tree or stick
* Bark (sound), a vocalization of some animals (which is commonly the dog)
Places
* Bark, Germany
* Bark, Warmian-Masurian Voivodeship, Poland
Arts, en ...
weathers gray and develops vertical fissures with age, flaking in strips.
The leaves are mostly flat with a decurrent base and a spreading blade connected by a short twisting petiole, but leading and cone-bearing shoots also have smaller scale-like leaves.[ The ]phyllotaxis
In botany, phyllotaxis () or phyllotaxy is the arrangement of leaves on a plant stem. Phyllotactic spirals form a distinctive class of patterns in nature.
Leaf arrangement
The basic arrangements of leaves on a stem are opposite and alternat ...
is spiral though the leaves of the lateral shoots are twisted to lie pectinately in two ranks and appear nearly opposite.[ The pectinate leaves are twisted at their petioles in opposite directions on each side of the shoot causing the adaxial sides of the leaves face up on one side of the shoot and down on the other side. The leaf blades are usually 15-25 millimeters long, 3-5 millimeters wide and ovate-lanceolate or ovate-elliptic in shape.][ Juvenile leaves are often larger.][ The narrow midrib of the leaf is conspicuous on the abaxial side. ]Stoma
In botany, a stoma (from Greek ''στόμα'', "mouth", plural "stomata"), also called a stomate (plural "stomates"), is a pore found in the epidermis of leaves, stems, and other organs, that controls the rate of gas exchange. The pore is bo ...
ta are present on both sides of the leaf.[
''Retrophyllum vitiense'' is ]dioecious
Dioecy (; ; adj. dioecious , ) is a characteristic of a species, meaning that it has distinct individual organisms (unisexual) that produce male or female gametes, either directly (in animals) or indirectly (in seed plants). Dioecious reproducti ...
. The cylindrical male pollen cones are borne apically on short lateral or subterminal branchlets. They grow in groups of two or three. A pollen cone consists of numerous spirally arranged microsporophylls around a 10-25 millimeter long rachis. The microsporophylls are triangular and keeled, bearing two pollen
Pollen is a powdery substance produced by seed plants. It consists of pollen grains (highly reduced microgametophytes), which produce male gametes (sperm cells). Pollen grains have a hard coat made of sporopollenin that protects the gametop ...
sacs each.[
The female ]seed cones
A conifer cone (in formal botanical usage: strobilus, plural strobili) is a seed-bearing organ on gymnosperm plants. It is usually woody, ovoid to globular, including scales and bracts arranged around a central axis, especially in conifers ...
are borne on short lateral branchlets. A seed cone has several sterile cone scales and usually just one fertile scale. The fertile cone scale has a single inverted ovule
In seed plants, the ovule is the structure that gives rise to and contains the female reproductive cells. It consists of three parts: the '' integument'', forming its outer layer, the ''nucellus'' (or remnant of the megasporangium), and the f ...
developing into a seed
A seed is an embryonic plant enclosed in a protective outer covering, along with a food reserve. The formation of the seed is a part of the process of reproduction in seed plants, the spermatophytes, including the gymnosperm and angiosper ...
. The seed is entirely enclosed by a modified ovuliferous scale known as the epimatium. The epimatium is green or glaucous at first and becomes fleshy and red in color at maturity.[ The mature epimatium is generally 14-20 millimeters long, 10-13 millimeters wide and pyriform in shape. The subglobose seed inside is 12-16 millimeters long.][
]
Distribution
''Retrophyllum vitiense'' is native to Southeast Asia and the southwestern Pacific. It occurs naturally on the Indonesian Maluku Islands
The Maluku Islands (; Indonesian: ''Kepulauan Maluku'') or the Moluccas () are an archipelago in the east of Indonesia. Tectonically they are located on the Halmahera Plate within the Molucca Sea Collision Zone. Geographically they are located ...
, Papua New Guinean New Guinea
New Guinea (; Hiri Motu: ''Niu Gini''; id, Papua, or , historically ) is the world's second-largest island with an area of . Located in Oceania in the southwestern Pacific Ocean, the island is separated from Australia by the wide Torr ...
and New Britain
New Britain ( tpi, Niu Briten) is the largest island in the Bismarck Archipelago, part of the Islands Region of Papua New Guinea. It is separated from New Guinea by a northwest corner of the Solomon Sea (or with an island hop of Umboi the D ...
, the Fiji islands and the Santa Cruz Islands
The Santa Cruz Islands are a group of islands in the Pacific Ocean, part of Temotu Province of the nation of Solomon Islands discovered by the Spaniards. They lie approximately 250 miles (400 km) to the southeast of the Solomon Island ...
of Solomon Islands.[
]
Habitat and ecology
''Retrophyllum vitiense'' is a large rainforest
Rainforests are characterized by a closed and continuous tree canopy, moisture-dependent vegetation, the presence of epiphytes and lianas and the absence of wildfire. Rainforest can be classified as tropical rainforest or temperate rainfo ...
tree. It grows in tropical lowland and montane rainforests ranging in altitude from near the sea level to 1800 meters.[ The mean annual precipitation of its natural habitat is 3290 millimeters.][ The species usually occurs as an emergent tree growing above the ]canopy
Canopy may refer to:
Plants
* Canopy (biology), aboveground portion of plant community or crop (including forests)
* Canopy (grape), aboveground portion of grapes
Religion and ceremonies
* Baldachin or canopy of state, typically placed over an ...
in forests dominated by other trees. It commonly occurs together with other tropical conifers such as trees of the genera ''Podocarpus
''Podocarpus'' () is a genus of conifers, the most numerous and widely distributed of the podocarp family, the Podocarpaceae. The name comes from Greek πούς (poús, “foot”) + καρπός (karpós, “fruit”). ''Podocarpus'' speci ...
'', ''Dacrycarpus
''Dacrycarpus'' is a genus of conifers belonging to the family Podocarpaceae.Christopher N. Page. 1990. "Podocarpaceae" pages 332-346. In: Klaus Kubitzki (general editor); Karl U. Kramer and Peter S. Green (volume editors) ''The Families and Gene ...
'', ''Dacrydium
''Dacrydium'' is a genus of conifers belonging to the podocarp family Podocarpaceae. Sixteen species of evergreen dioecious trees and shrubs are presently recognized. The genus was first described by Solander in 1786, and formerly included many ...
'' and ''Agathis
''Agathis'', commonly known as kauri or dammara, is a genus of 22 species of evergreen tree. The genus is part of the ancient conifer family Araucariaceae, a group once widespread during the Jurassic and Cretaceous periods, but now largely res ...
''.[
]
References
{{Taxonbar, from=Q5413473
Podocarpaceae
Least concern plants
Taxonomy articles created by Polbot