''Myrica rubra'', also called yangmei (; Cantonese: yeung4 mui4;

Shanghainese The Shanghainese language, also known as the Shanghai dialect, or Hu language, is a variety of Wu Chinese spoken in the central districts of the city of Shanghai and its surrounding areas. It is classified as part of the Sino-Tibetan langua ...

: ), , Chinese bayberry, red bayberry, yumberry, waxberry, or Chinese strawberry (and often mistranslated from Chinese as

arbutus ''Arbutus'' is a genus of 12 accepted speciesAct. Bot. Mex no.99 Pátzcuaro abr. 2012.''Arbutus bicolor''/ref> of flowering plants in the family Ericaceae, native to temperate regions of the Mediterranean, western Europe, the Canary Islands a ...

) is a subtropical

tree In botany, a tree is a perennial plant with an elongated stem, or trunk, usually supporting branches and leaves. In some usages, the definition of a tree may be narrower, e.g., including only woody plants with secondary growth, only ...

grown for its

fruit In botany, a fruit is the seed-bearing structure in flowering plants (angiosperms) that is formed from the ovary after flowering. Fruits are the means by which angiosperms disseminate their seeds. Edible fruits in particular have long propaga ...

Description

''Myrica rubra'' is an

evergreen In botany, an evergreen is a plant which has Leaf, foliage that remains green and functional throughout the year. This contrasts with deciduous plants, which lose their foliage completely during the winter or dry season. Consisting of many diffe ...

tree that grows to a height of up to high, with smooth gray bark and a uniform spherical to hemispherical crown. Leaves are leathery, bare, elliptic-obovate to oval lanceolate in shape, wedge-shaped at the base and rounded to pointed or tapered at the apex, margin is serrated or serrated in the upper half, with a length of and a width of . Leaves are alternately arranged on the branches are divided into petiole and leaf blade. The petiole is long. The leaf underside is pale green and sparsely to moderately golden glandular, the top surface is dark green. The species is

dioecious Dioecy ( ; ; adj. dioecious, ) is a characteristic of certain species that have distinct unisexual individuals, each producing either male or female gametes, either directly (in animals) or indirectly (in seed plants). Dioecious reproduction is ...

. Male flowers with simple or unobtrusively branched bracts, are held in inflorescences individually or occasionally in groups of a few inflorescences in the leaf axils. Female flowers are long, in inflorescences with bare stems, the bracts almost circular with a diameter of about 1 millimeter, and have golden glands on the underside. The male flowers are accompanied by two to four egg-shaped, sparse lanceolate leaves. Each male flower contains four to six stamens with dark red, elliptical anthers. Female inflorescences are single with multi-flowered spikes of in length standing in the leaf axils. The rhachis is hairy and glandular. The cover sheets overlap, are hairless and only unobtrusively glandular. Female flowers are accompanied by four leaves. The upper ovary is velvety hairy, with a stylus with a two-lobed scar. There are two slender scar lobes that are colored bright red. The flowering period extends from March to April in China, with fruits developing from May to June. The fruit is spherical, typically in diameter, with diameters up to 3 centimeters, a knobby surface. The surface is a thick-skinned, typically a crimson red, but may vary from white to purple, with similar or somewhat lighter flesh color. At the center is a single seed, with a diameter about half that of the whole fruit. The flesh is sweet and very tart. Myrica rubra4.jpg, Flowers Morella rubra - Leping, Jiangxi, China 2014.06.15 11-43-07.jpg, Fruit Myrica rubra1.jpg, Leaves File:Berkeley Yangmei Tree.jpg, Tree File:Ripe yangmei.jpg, Ripe Fruit

Taxonomy

The plant was first described by

João de Loureiro João de Loureiro (1717, Lisbon – 18 October 1791) was a Portuguese people, Portuguese Jesuit missionary and botanist. Biography After receiving admission to the Jesuit Order, João de Loureiro served as a missionary in Goa, capital of Port ...

in Flora Cochinchinensis, 2, page 548 in 1790 under the name (

basionym In the scientific name of organisms, basionym or basyonym means the original name on which a new name is based; the author citation of the new name should include the authors of the basionym in parentheses. The term "basionym" is used in both botan ...

) ''Morella rubra'' The species was moved to the genus ''

Myrica ''Myrica'' is a genus of about 35–50 species of small trees and shrubs in the family Myricaceae, order Fagales. The genus has a wide distribution, including Africa, Asia, Europe, North America, and South America, and missing only from Antar ...

'' as ''Myrica rubra'' . by Philipp Franz von Siebold and Joseph Gerhard von Zuccarini in treatises of the Bavarian Academy of Sciences. Mathematical and natural science class, volume 4, number 3, page 230 published. In studies of

germplasm Germplasm refers to genetic resources such as seeds, tissues, and DNA sequences that are maintained for the purpose of animal and plant breeding, conservation efforts, agriculture, and other research uses. These resources may take the form of s ...

, it was clearly distinguished from wax myrtle, and could be subdivided into two groups unrelated to the sex of the plant, but more so by the geographic region in

China China, officially the People's Republic of China (PRC), is a country in East Asia. With population of China, a population exceeding 1.4 billion, it is the list of countries by population (United Nations), second-most populous country after ...

where the accession originated. Among regions in China, accessions varied within regions, indicating extensive

gene In biology, the word gene has two meanings. The Mendelian gene is a basic unit of heredity. The molecular gene is a sequence of nucleotides in DNA that is transcribed to produce a functional RNA. There are two types of molecular genes: protei ...

mixing. Nearly 100

cultivar A cultivar is a kind of Horticulture, cultivated plant that people have selected for desired phenotypic trait, traits and which retains those traits when Plant propagation, propagated. Methods used to propagate cultivars include division, root a ...

s of ''M. rubra'' exist in China alone.

Zhejiang Province ) , translit_lang1_type2 = , translit_lang1_info2 = ( Hangzhounese) ( Ningbonese) (Wenzhounese) , image_skyline = 玉甑峰全貌 - panoramio.jpg , image_caption = View of the Yandang Mountains , image_map = Zhejiang i ...

is a possible center of diversity for the plant in China. Chromosome count is 2n = 16

Distribution and habitat

It is native to eastern

Asia Asia ( , ) is the largest continent in the world by both land area and population. It covers an area of more than 44 million square kilometres, about 30% of Earth's total land area and 8% of Earth's total surface area. The continent, which ...

, mainly in south-central

in province of

Fujian Fujian is a provinces of China, province in East China, southeastern China. Fujian is bordered by Zhejiang to the north, Jiangxi to the west, Guangdong to the south, and the Taiwan Strait to the east. Its capital is Fuzhou and its largest prefe ...

Guangdong ) means "wide" or "vast", and has been associated with the region since the creation of Guang Prefecture in AD 226. The name "''Guang''" ultimately came from Guangxin ( zh, labels=no, first=t, t= , s=广信), an outpost established in Han dynasty ...

Guangxi Guangxi,; officially the Guangxi Zhuang Autonomous Region, is an Autonomous regions of China, autonomous region of the China, People's Republic of China, located in South China and bordering Vietnam (Hà Giang Province, Hà Giang, Cao Bằn ...

Guizhou ) , image_skyline = , image_caption = , image_map = Guizhou in China (+all claims hatched).svg , mapsize = 275px , map_alt = Map showing the location of Guizhou Province , map_caption = Map s ...

Hainan Hainan is an island provinces of China, province and the southernmost province of China. It consists of the eponymous Hainan Island and various smaller islands in the South China Sea under the province's administration. The name literally mean ...

Hunan Hunan is an inland Provinces of China, province in Central China. Located in the middle reaches of the Yangtze watershed, it borders the Administrative divisions of China, province-level divisions of Hubei to the north, Jiangxi to the east, Gu ...

Jiangsu Jiangsu is a coastal Provinces of the People's Republic of China, province in East China. It is one of the leading provinces in finance, education, technology, and tourism, with its capital in Nanjing. Jiangsu is the List of Chinese administra ...

Jiangxi ; Gan: ) , translit_lang1_type2 = , translit_lang1_info2 = , translit_lang1_type3 = , translit_lang1_info3 = , image_map = Jiangxi in China (+all claims hatched).svg , mapsize = 275px , map_caption = Location ...

Sichuan Sichuan is a province in Southwestern China, occupying the Sichuan Basin and Tibetan Plateau—between the Jinsha River to the west, the Daba Mountains to the north, and the Yunnan–Guizhou Plateau to the south. Its capital city is Cheng ...

Yunnan Yunnan; is an inland Provinces of China, province in Southwestern China. The province spans approximately and has a population of 47.2 million (as of 2020). The capital of the province is Kunming. The province borders the Chinese provinces ...

, and

Zhejiang ) , translit_lang1_type2 = , translit_lang1_info2 = ( Hangzhounese) ( Ningbonese) (Wenzhounese) , image_skyline = 玉甑峰全貌 - panoramio.jpg , image_caption = View of the Yandang Mountains , image_map = Zhejiang i ...

;

Japan Japan is an island country in East Asia. Located in the Pacific Ocean off the northeast coast of the Asia, Asian mainland, it is bordered on the west by the Sea of Japan and extends from the Sea of Okhotsk in the north to the East China Sea ...

Korea Korea is a peninsular region in East Asia consisting of the Korean Peninsula, Jeju Island, and smaller islands. Since the end of World War II in 1945, it has been politically Division of Korea, divided at or near the 38th parallel north, 3 ...

, and the

Philippines The Philippines, officially the Republic of the Philippines, is an Archipelagic state, archipelagic country in Southeast Asia. Located in the western Pacific Ocean, it consists of List of islands of the Philippines, 7,641 islands, with a tot ...

in forests on mountain slopes and valleys at elevations of . Seeds are dispersed by

Japanese macaque The Japanese macaque (''Macaca fuscata''), also known as the snow monkey, is a terrestrial Old World monkey species that is native to Japan. Colloquially, they are referred to as "snow monkeys" because some live in areas where snow covers the g ...

s and Yakushima macaques.

Cultivation

Chinese cultivation is concentrated south of the

Yangtze River The Yangtze or Yangzi ( or ) is the longest river in Eurasia and the third-longest in the world. It rises at Jari Hill in the Tanggula Mountains of the Tibetan Plateau and flows including Dam Qu River the longest source of the Yangtze, i ...

, where it has considerable economic importance and has been grown for at least 2000 years. It tolerates poor

acid An acid is a molecule or ion capable of either donating a proton (i.e. Hydron, hydrogen cation, H+), known as a Brønsted–Lowry acid–base theory, Brønsted–Lowry acid, or forming a covalent bond with an electron pair, known as a Lewis ...

soil Soil, also commonly referred to as earth, is a mixture of organic matter, minerals, gases, water, and organisms that together support the life of plants and soil organisms. Some scientific definitions distinguish dirt from ''soil'' by re ...

s. The

root In vascular plants, the roots are the plant organ, organs of a plant that are modified to provide anchorage for the plant and take in water and nutrients into the plant body, which allows plants to grow taller and faster. They are most often bel ...

system is deep, with no obvious

taproot A taproot is a large, central, and dominant root from which other roots sprout laterally. Typically a taproot is somewhat straight and very thick, is tapering in shape, and grows directly downward. In some plants, such as the carrot, the taproot ...

. ''Myrica rubra'' was first introduced into the United States by Frank Nicholas Meyer from seed purchased from the Yokohama Nursery Co. in Japan and published in the Bulletin of Foreign Plant Introductions in 1918. Plants from the collection were grown and fruited in

Chico, California Chico ( ; Spanish language, Spanish for "little") is the most populous city in Butte County, California, United States. Located in the Sacramento Valley region of Northern California, the city had a population of 101,475 in the 2020 United Sta ...

and in

Brooksville, Florida Brooksville is a city in and the county seat of Hernando County, Florida, Hernando County, Florida, in the United States. At the 2010 United States Census, 2010 census it had a population of 7,719, up from 7,264 at the 2000 census. Brooksville is ...

David Fairchild David Grandison Fairchild (April 7, 1869 – August 6, 1954) was an American botanist and plant explorer. Fairchild was responsible for the introduction of more than 200,000 exotic plants and varieties of established crops into the United Stat ...

. ''M. rubra'' is being commercialized in California by Calmei, a California corporation. Trees are prolific producers, with a single tree yielding some of fruit. As of 2007, 865,000

acres The acre ( ) is a unit of land area used in the British imperial and the United States customary systems. It is traditionally defined as the area of one chain by one furlong (66 by 660 feet), which is exactly equal to 10 square chains, ...

were devoted to yangmei production in

double the amount of acres utilized in

apple An apple is a round, edible fruit produced by an apple tree (''Malus'' spp.). Fruit trees of the orchard or domestic apple (''Malus domestica''), the most widely grown in the genus, are agriculture, cultivated worldwide. The tree originated ...

production in the

United States The United States of America (USA), also known as the United States (U.S.) or America, is a country primarily located in North America. It is a federal republic of 50 U.S. state, states and a federal capital district, Washington, D.C. The 48 ...

. The tree is used as ornaments for parks and streets. It is also a traditional tree used in composing classical East Asian gardens.

Products

Some

s with large fruit, up to in diameter, have been developed. Besides fresh consumption, the fruits may be dried, canned, soaked in ''

baijiu ''Baijiu'' (), or ''shaojiu'' (), is a colorless Chinese liquor typically coming in between 35% and 60% alcohol by volume (ABV). Each type of baijiu uses its own type of ''qū'' for fermentation to create a distinct and characteristic flavo ...

'' (Chinese

liquor Liquor ( , sometimes hard liquor), spirits, distilled spirits, or spiritous liquor are alcoholic drinks produced by the distillation of grains, fruits, vegetables, or sugar that have already gone through ethanol fermentation, alcoholic ferm ...

), or fermented into

alcoholic beverage Drinks containing alcohol (drug), alcohol are typically divided into three classes—beers, wines, and Distilled beverage, spirits—with alcohol content typically between 3% and 50%. Drinks with less than 0.5% are sometimes considered Non-al ...

s, such as wine,

beer Beer is an alcoholic beverage produced by the brewing and fermentation of starches from cereal grain—most commonly malted barley, although wheat, maize (corn), rice, and oats are also used. The grain is mashed to convert starch in the ...

, or

cocktail A cocktail is a mixed drink, usually alcoholic beverage, alcoholic. Most commonly, a cocktail is a combination of one or more liquor, spirits mixed with other ingredients, such as juices, flavored syrups, tonic water, Shrub (drink), shrubs, and ...

s. Dried fruits are often prepared in the manner of dry '' huamei'' (''

Prunus mume ''Prunus mume'', the Chinese plum or Japanese apricot, is a tree species in the family Rosaceae. Along with bamboo, the plant is intimately associated with art, literature, and everyday life in China, from where it was then introduced to Kor ...

'' with flavorings such as

licorice Liquorice (Commonwealth English) or licorice (American English; see spelling differences; ) is the common name of ''Glycyrrhiza glabra'', a flowering plant of the bean family Fabaceae, from the root of which a sweet, aromatic flavouring is ...

or salty licorice). The juice has been commercialised under the brand name "Yumberry" under which name it is trademarked in the EU. In

Yunnan Province Yunnan; is an inland Provinces of China, province in Southwestern China. The province spans approximately and has a population of 47.2 million (as of 2020). The capital of the province is Kunming. The province borders the Chinese provinces ...

in China, there are two main types of yangmei, a sour type used for making dried fruit and a sweet type used for juice and fresh eating. In the

, they are dried and preserved in brine and vinegar and made into champóy, the local version of the Chinese '' huamei''. Other uses include * bottled

pasteurize In food processing, pasteurization ( also pasteurisation) is a process of food preservation in which packaged foods (e.g., milk and fruit juices) are treated with mild heat, usually to less than , to eliminate pathogens and extend shelf life ...

juice Juice is a drink made from the extraction or Cold-pressed juice, pressing of the natural liquid contained in fruit and vegetables. It can also refer to liquids that are flavored with concentrate or other biological food sources, such as meat ...

or juice blends *

dye Juan de Guillebon, better known by his stage name DyE, is a French musician. He is known for the music video of the single "Fantasy Fantasy is a genre of speculative fiction that involves supernatural or Magic (supernatural), magical ele ...

prepared from the bark *

yogurt Yogurt (; , from , ; also spelled yoghurt, yogourt or yoghourt) is a food produced by bacterial Fermentation (food), fermentation of milk. Fermentation of sugars in the milk by these bacteria produces lactic acid, which acts on milk protein to ...

flavoring * blended jam and preserves

Research and phytochemicals

Various species of ''Myrica'' have been studied scientifically for

horticultural Horticulture (from ) is the art and science of growing fruits, vegetables, flowers, trees, shrubs and ornamental plants. Horticulture is commonly associated with the more professional and technical aspects of plant cultivation on a smaller and mo ...

characteristics or

phytochemicals Phytochemicals are naturally-occurring chemicals present in or extract, extracted from plants. Some phytochemicals are nutrients for the plant, while others are metabolites produced to enhance plant survivability and reproduction. The field ...

implicated with health benefits. Dating to 1951, the horticultural literature includes studies on *

nitrogen Nitrogen is a chemical element; it has Symbol (chemistry), symbol N and atomic number 7. Nitrogen is a Nonmetal (chemistry), nonmetal and the lightest member of pnictogen, group 15 of the periodic table, often called the Pnictogen, pnictogens. ...

-fixing ability of the

root nodule Root nodules are found on the roots of plants, primarily legumes, that form a symbiosis with nitrogen-fixing bacteria. Under nitrogen-limiting conditions, capable plants form a symbiotic relationship with a host-specific strain of bacteria known ...

s system * presence of ''

Frankia ''Frankia'' is a genus of nitrogen-fixing bacteria that live in symbiosis with actinorhizal plants, similar to the '' Rhizobium'' bacteria found in the root nodules of legumes in the family Fabaceae. ''Frankia'' also initiate the forming of ro ...

bacteria Bacteria (; : bacterium) are ubiquitous, mostly free-living organisms often consisting of one Cell (biology), biological cell. They constitute a large domain (biology), domain of Prokaryote, prokaryotic microorganisms. Typically a few micr ...

having nitrogen-fixing properties in root nodules *

microbial A microorganism, or microbe, is an organism of microscopic size, which may exist in its single-celled form or as a colony of cells. The possible existence of unseen microbial life was suspected from antiquity, with an early attestation in ...

characteristics of the subcanopy soil * niche characteristics in the

forest A forest is an ecosystem characterized by a dense ecological community, community of trees. Hundreds of definitions of forest are used throughout the world, incorporating factors such as tree density, tree height, land use, legal standing, ...

environment * growth of

pollen Pollen is a powdery substance produced by most types of flowers of seed plants for the purpose of sexual reproduction. It consists of pollen grains (highly reduced Gametophyte#Heterospory, microgametophytes), which produce male gametes (sperm ...

tubes

Cultural significance

Archaeological and written evidence suggest that yangmei cultivation first took place in China over 2,000 years ago during the

Han dynasty The Han dynasty was an Dynasties of China, imperial dynasty of China (202 BC9 AD, 25–220 AD) established by Liu Bang and ruled by the House of Liu. The dynasty was preceded by the short-lived Qin dynasty (221–206 BC ...

. Yangmei is mentioned throughout

Chinese literature The history of Chinese literature extends thousands of years, and begins with the earliest recorded inscriptions, court archives, building to the major works of philosophy and history written during the Axial Age. The Han dynasty, Han (202 ...

, including several appearances in

Li Bai Li Bai (, 701–762), Literary and colloquial readings, also pronounced Li Bo, courtesy name Taibai (), was a Chinese poet acclaimed as one of the greatest and most important poets of the Tang dynasty and in Chinese history as a whole. He and hi ...

's poems. In Japan, it is the prefectural flower of Kōchi and the prefectural tree of

Tokushima is a Prefectures of Japan, prefecture of Japan located on the island of Shikoku. Tokushima Prefecture has a population of 682,439 (1 February 2025) and has a geographic area of 4,146 Square kilometre, km2 (1,601 sq mi). Tokushima Prefecture b ...

. The plant's name appears in many old Japanese poems.

References

External links

* * {{Authority control rubra Berries Endemic flora of China Trees of China Flora of South-Central China Garden plants of Asia Ornamental trees Dioecious plants Plants described in 1846 Taxa named by Philipp Franz von Siebold Taxa named by Joseph Gerhard Zuccarini