Ray-finned Fish Taxa
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Actinopterygii (; ), members of which are known as ray-finned fish or actinopterygians, is a
class Class, Classes, or The Class may refer to: Common uses not otherwise categorized * Class (biology), a taxonomic rank * Class (knowledge representation), a collection of individuals or objects * Class (philosophy), an analytical concept used d ...
of
bony fish Osteichthyes ( ; ), also known as osteichthyans or commonly referred to as the bony fish, is a Biodiversity, diverse clade of vertebrate animals that have endoskeletons primarily composed of bone tissue. They can be contrasted with the Chondricht ...
that comprise over 50% of living
vertebrate Vertebrates () are animals with a vertebral column (backbone or spine), and a cranium, or skull. The vertebral column surrounds and protects the spinal cord, while the cranium protects the brain. The vertebrates make up the subphylum Vertebra ...
species. They are so called because of their lightly built
fin A fin is a thin component or appendage attached to a larger body or structure. Fins typically function as foils that produce lift or thrust, or provide the ability to steer or stabilize motion while traveling in water, air, or other fluids. F ...
s made of webbings of
skin Skin is the layer of usually soft, flexible outer tissue covering the body of a vertebrate animal, with three main functions: protection, regulation, and sensation. Other animal coverings, such as the arthropod exoskeleton, have different ...
supported by radially extended thin bony
spine Spine or spinal may refer to: Science Biology * Spinal column, also known as the backbone * Dendritic spine, a small membranous protrusion from a neuron's dendrite * Thorns, spines, and prickles, needle-like structures in plants * Spine (zoology), ...
s called ''
lepidotrichia In a zoological context, spines are hard, needle-like anatomical structures found in both vertebrate and invertebrate species. The spines of most spiny mammals are modified hairs, with a spongy center covered in a thick, hard layer of keratin and ...
'', as opposed to the bulkier, fleshy lobed fins of the
sister A sister is a woman or a girl who shares parents or a parent with another individual; a female sibling. The male counterpart is a brother. Although the term typically refers to a familial relationship, it is sometimes used endearingly to ref ...
clade In biology, a clade (), also known as a Monophyly, monophyletic group or natural group, is a group of organisms that is composed of a common ancestor and all of its descendants. Clades are the fundamental unit of cladistics, a modern approach t ...
Sarcopterygii Sarcopterygii (; )—sometimes considered synonymous with Crossopterygii ()—is a clade (traditionally a class (biology), class or subclass) of vertebrate animals which includes a group of bony fish commonly referred to as lobe-finned fish. The ...
(lobe-finned fish). Resembling
folding fan A handheld fan, or simply hand fan, is a broad, flat surface that is waved back and forth to create an airflow. Generally, purpose-made handheld fans are folding fans, which are shaped like a sector of a circle and made of a thin material (such ...
s, the actinopterygian fins can easily change shape and
wetted area In fluid dynamics, the wetted area is the surface area that interacts with the working fluid or gas. * In maritime use, the wetted area is the area of the watercraft's hull which is immersed in water. This has a direct relationship on the overal ...
, providing superior
thrust-to-weight ratio Thrust-to-weight ratio is a dimensionless ratio of thrust to weight of a rocket, jet engine, propeller engine, or a vehicle propelled by such an engine that is an indicator of the performance of the engine or vehicle. The instantaneous thrust-to-w ...
s per movement compared to sarcopterygian and
chondrichthyian Chondrichthyes (; ) is a class of jawed fish that contains the cartilaginous fish or chondrichthyans, which all have skeletons primarily composed of cartilage. They can be contrasted with the Osteichthyes or ''bony fish'', which have skeletons p ...
fins. The fin rays attach directly to the proximal or basal skeletal elements, the radials, which represent the articulation between these fins and the internal skeleton (e.g., pelvic and pectoral girdles). The vast majority of actinopterygians are
teleost Teleostei (; Ancient Greek, Greek ''teleios'' "complete" + ''osteon'' "bone"), members of which are known as teleosts (), is, by far, the largest group of ray-finned fishes (class Actinopterygii), with 96% of all neontology, extant species of f ...
s. By
species A species () is often defined as the largest group of organisms in which any two individuals of the appropriate sexes or mating types can produce fertile offspring, typically by sexual reproduction. It is the basic unit of Taxonomy (biology), ...
count, they dominate the
subphylum In zoological nomenclature, a subphylum is a taxonomic rank below the rank of phylum. The taxonomic rank of " subdivision" in fungi and plant taxonomy is equivalent to "subphylum" in zoological taxonomy. Some plant taxonomists have also used th ...
Vertebrata Vertebrates () are animals with a vertebral column (backbone or spine), and a cranium, or skull. The vertebral column surrounds and protects the spinal cord, while the cranium protects the brain. The vertebrates make up the subphylum Verte ...
, and constitute nearly 99% of the over 30,000
extant Extant or Least-concern species, least concern is the opposite of the word extinct. It may refer to: * Extant hereditary titles * Extant literature, surviving literature, such as ''Beowulf'', the oldest extant manuscript written in English * Exta ...
species of
fish A fish (: fish or fishes) is an aquatic animal, aquatic, Anamniotes, anamniotic, gill-bearing vertebrate animal with swimming fish fin, fins and craniate, a hard skull, but lacking limb (anatomy), limbs with digit (anatomy), digits. Fish can ...
. They are the most abundant
nektonic Nekton or necton (from the ) is any aquatic organism that can actively and persistently propel itself through a water column (i.e. swimming) without touching the bottom. Nektons generally have powerful tails and appendages (e.g. fins, pleopods ...
aquatic animal An aquatic animal is any animal, whether vertebrate or invertebrate, that lives in a body of water for all or most of its lifetime. Aquatic animals generally conduct gas exchange in water by extracting dissolved oxygen via specialised respirato ...
s and are ubiquitous throughout
freshwater Fresh water or freshwater is any naturally occurring liquid or frozen water containing low concentrations of dissolved salts and other total dissolved solids. The term excludes seawater and brackish water, but it does include non-salty mi ...
and marine environments from the
deep sea The deep sea is broadly defined as the ocean depth where light begins to fade, at an approximate depth of or the point of transition from continental shelves to continental slopes. Conditions within the deep sea are a combination of low tempe ...
to subterranean waters to the highest
mountain stream A mountain stream is a Stream, brook or stream, usually with a steep gradient, flowing down a mountainside. Its swift flow rate often transports large quantities of rock (geology), rock, gravel, soil, wood or even entire Trunk (botany), logs with ...
s. Extant species can range in size from ''
Paedocypris ''Paedocypris'' is a genus of tiny cyprinid fish found in swamps and streams on the Southeast Asian islands of Borneo, Sumatra and Bintan. ''Paedocypris progenetica'' has been claimed to be the one of smallest known species of fish in the worl ...
'', at ; to the massive
ocean sunfish The ocean sunfish (''Mola mola''), also known as the common mola, is one of the largest bony fish in the world. It is the type species of the genus ''Mola'', and one of five extant species in the family Molidae. It was once misidentified as th ...
, at ; and to the giant
oarfish Oarfish are large and extremely long pelagic lampriform fish belonging to the small family (biology), family Regalecidae. Found in areas spanning from Temperate climate, temperate ocean zones to tropical ones, yet rarely seen by humans, the oa ...
, at . The largest ever known ray-finned fish, the extinct ''
Leedsichthys ''Leedsichthys'' is an extinct genus of pachycormid fish that lived in the oceans of the Middle to Late Jurassic.Liston, JJ (2004). An overview of the pachycormiform ''Leedsichthys''. In: Arratia G and Tintori A (eds) Mesozoic Fishes 3 - Systema ...
'' from the
Jurassic The Jurassic ( ) is a Geological period, geologic period and System (stratigraphy), stratigraphic system that spanned from the end of the Triassic Period million years ago (Mya) to the beginning of the Cretaceous Period, approximately 143.1 Mya. ...
, has been estimated to have grown to .


Characteristics

Ray-finned fishes occur in many variant forms. The main features of typical ray-finned fish are shown in the adjacent diagram. The
swim bladder The swim bladder, gas bladder, fish maw, or air bladder is an internal gas-filled organ (anatomy), organ in bony fish that functions to modulate buoyancy, and thus allowing the fish to stay at desired water depth without having to maintain lift ...
is a more derived structure and used for
buoyancy Buoyancy (), or upthrust, is the force exerted by a fluid opposing the weight of a partially or fully immersed object (which may be also be a parcel of fluid). In a column of fluid, pressure increases with depth as a result of the weight of t ...
. Except from the
bichir Bichirs and the reedfish comprise Polypteridae , a family (biology), family of archaic Actinopterygii, ray-finned fishes and the only family in the order (biology), order Polypteriformes .Helfman GS, Collette BB, Facey DE, Bowen BW. 2009. The D ...
s, which just like the
lung The lungs are the primary Organ (biology), organs of the respiratory system in many animals, including humans. In mammals and most other tetrapods, two lungs are located near the Vertebral column, backbone on either side of the heart. Their ...
s of
lobe-finned fish Sarcopterygii (; )—sometimes considered synonymous with Crossopterygii ()—is a clade (traditionally a class or subclass) of vertebrate animals which includes a group of bony fish commonly referred to as lobe-finned fish. These vertebrates ar ...
have retained the ancestral condition of ventral budding from the
foregut The foregut in humans is the anterior part of the alimentary canal, from the distal esophagus to the first half of the duodenum, at the entrance of the bile duct. Beyond the stomach, the foregut is attached to the abdominal walls by mesentery. ...
, the swim bladder in ray-finned fishes derives from a dorsal bud above the foregut. In early forms the swim bladder could still be used for breathing, a trait still present in
Holostei Holostei is a group of ray-finned bony fish. It is divided into two major clades, the Halecomorphi, represented by the single living genus, '' Amia'' with two species, the bowfins (''Amia calva'' and '' Amia ocellicauda''), as well as the Gin ...
(
bowfin The ruddy bowfin (''Amia calva'') is a ray-finned fish native to North America. Common names include mudfish, mud pike, dogfish, grindle, grinnel, swamp trout, and choupique. It is regarded as a relict, being one of only two surviving species ...
s and
gar Gars are an ancient group of ray-finned fish in the family Lepisosteidae. They comprise seven living species of fish in two genera that inhabit fresh, brackish, and occasionally marine waters of eastern North America, Central America and Cuba ...
s). In some fish like the
arapaima The arapaima, pirarucu, or paiche is any large species of bonytongue in the genus ''Arapaima'' native to the Amazon Basin, Amazon and Essequibo River, Essequibo basins of South America. ''Arapaima'' is the type genus of the subfamily Arapaiminae ...
, the swim bladder has been modified for breathing air again, and in other lineages it has been completely lost. The teleosts have urinary and reproductive tracts that are fully separated, while the Chondrostei have common urogenital ducts, and partially connected ducts are found in Cladistia and Holostei. Ray-finned fishes have many different types of scales; but all
teleost Teleostei (; Ancient Greek, Greek ''teleios'' "complete" + ''osteon'' "bone"), members of which are known as teleosts (), is, by far, the largest group of ray-finned fishes (class Actinopterygii), with 96% of all neontology, extant species of f ...
s have
leptoid scale A fish scale is a small rigid plate that grows out of the skin of a fish. The skin of most jawed fishes is covered with these protective scales, which can also provide effective camouflage through the use of reflection and colouration, as we ...
s. The outer part of these scales fan out with bony ridges, while the inner part is crossed with fibrous connective tissue. Leptoid scales are thinner and more transparent than other types of scales, and lack the hardened enamel- or
dentine Dentin ( ) (American English) or dentine ( or ) (British English) () is a calcified tissue of the body and, along with enamel, cementum, and pulp, is one of the four major components of teeth. It is usually covered by enamel on the crown ...
-like layers found in the scales of many other fish. Unlike
ganoid scale A fish scale is a small rigid plate that grows out of the skin of a fish. The skin of most jawed fishes is covered with these protective scales, which can also provide effective camouflage through the use of reflection and colouration, as w ...
s, which are found in non-teleost actinopterygians, new scales are added in concentric layers as the fish grows. Teleosts and chondrosteans (sturgeons and paddlefish) also differ from the bichirs and holosteans (bowfin and gars) in having gone through a whole-genome duplication (
paleopolyploidy Paleopolyploidy is the result of genome duplications which occurred at least several million years ago (MYA). Such an event could either double the genome of a single species (autopolyploidy) or combine those of two species (allopolyploidy). Bec ...
). The WGD is estimated to have happened about 320 million years ago in the teleosts, which on average has retained about 17% of the gene duplicates, and around 180 (124–225) million years ago in the chondrosteans. It has since happened again in some teleost lineages, like Salmonidae (80–100 million years ago) and several times independently within the
Cyprinidae Cyprinidae is a family of freshwater fish commonly called the carp or minnow family, including the carps, the true minnows, and their relatives the barbs and barbels, among others. Cyprinidae is the largest and most diverse fish family, and t ...
(in goldfish and common carp as recently as 14 million years ago).


Body shapes and fin arrangements

Ray-finned fish vary in size and shape, in their feeding specializations, and in the number and arrangement of their ray-fins.


Reproduction

In nearly all ray-finned fish, the sexes are separate, and in most species the females spawn eggs that are fertilized externally, typically with the male inseminating the eggs after they are laid. Development then proceeds with a free-swimming larval stage. However other patterns of
ontogeny Ontogeny (also ontogenesis) is the origination and development of an organism (both physical and psychological, e.g., moral development), usually from the time of fertilization of the ovum, egg to adult. The term can also be used to refer to t ...
exist, with one of the commonest being
sequential hermaphroditism Sequential hermaphroditism (called dichogamy in botany) is one of the two types of hermaphroditism, the other type being simultaneous hermaphroditism. It occurs when the organism's sex changes at some point in its life. A sequential hermaphrodit ...
. In most cases this involves
protogyny Sequential hermaphroditism (called dichogamy in botany) is one of the two types of hermaphroditism, the other type being simultaneous hermaphroditism. It occurs when the organism's sex changes at some point in its life. A sequential hermaphrodit ...
, fish starting life as females and converting to males at some stage, triggered by some internal or external factor.
Protandry Sequential hermaphroditism (called dichogamy in botany) is one of the two types of hermaphroditism, the other type being simultaneous hermaphroditism. It occurs when the organism's sex changes at some point in its life. A sequential hermaphrodit ...
, where a fish converts from male to female, is much less common than protogyny. Most families use external rather than
internal fertilization Internal fertilization is the union of an egg and sperm cell during sexual reproduction inside the female body. Internal fertilization, unlike its counterpart, external fertilization, brings more control to the female with reproduction. For inte ...
. Of the
oviparous Oviparous animals are animals that reproduce by depositing fertilized zygotes outside the body (i.e., by laying or spawning) in metabolically independent incubation organs known as eggs, which nurture the embryo into moving offsprings kno ...
teleosts, most (79%) do not provide parental care.
Viviparity In animals, viviparity is development of the embryo inside the body of the mother, with the maternal circulation providing for the metabolic needs of the embryo's development, until the mother gives birth to a fully or partially developed juv ...
,
ovoviviparity Ovoviviparity, ovovivipary, ovivipary, or aplacental viviparity is a "bridging" form of reproduction between egg-laying oviparous and live-bearing viviparous reproduction. Ovoviviparous animals possess embryos that develop inside eggs that r ...
, or some form of parental care for eggs, whether by the male, the female, or both parents is seen in a significant fraction (21%) of the 422 teleost families; no care is likely the ancestral condition. The oldest case of viviparity in ray-finned fish is found in
Middle Triassic In the geologic timescale, the Middle Triassic is the second of three epoch (geology), epochs of the Triassic period (geology), period or the middle of three series (stratigraphy), series in which the Triassic system (stratigraphy), system is di ...
species of ''
Saurichthys ''Saurichthys'' (from , 'lizard' and 'fish') is an extinct genus of predatory Actinopterygii, ray-finned fish from the Triassic period (geology), Period. It is the type genus of the family (taxonomy), family Saurichthyidae (Changhsingian-Ju ...
''. Viviparity is relatively rare and is found in about 6% of living teleost species; male care is far more common than female care. Male territoriality "preadapts" a species for evolving male parental care. There are a few examples of fish that self-fertilise. The
mangrove rivulus The mangrove rivulus or mangrove killifish, ''Kryptolebias marmoratus'' (syn. ''Rivulus marmoratus''), is a species of killifish in the Family (biology), family Rivulidae. It lives in brackish and marine waters (less frequently in fresh water) a ...
is an amphibious, simultaneous hermaphrodite, producing both eggs and spawn and having internal fertilisation. This mode of reproduction may be related to the fish's habit of spending long periods out of water in the mangrove forests it inhabits. Males are occasionally produced at temperatures below and can fertilise eggs that are then spawned by the female. This maintains genetic variability in a species that is otherwise highly inbred.


Classification and fossil record

Actinopterygii is divided into the subclasses
Cladistia Cladistia is a subclass of bony fishes whose only living members are the bichirs of tropical Africa. Their major synapomorphies are a heterocercal tail in which the dorsal fin has independent rays, and a posteriorly elongated parasphenoid. ...
,
Chondrostei Chondrostei is a subclass of non- neopterygian ray-finned fish. While the term originally referred to the paraphyletic grouping of all non-neopterygian ray-finned fish, it was redefined by Patterson in 1982 to be a clade comprising the Acipenser ...
and
Neopterygii Neopterygii (from Greek νέος ''neos'' 'new' and πτέρυξ ''pteryx'' 'fin') is a subclass of ray-finned fish (Actinopterygii). Neopterygii includes the Holostei and the Teleostei, of which the latter comprise the vast majority of extant ...
. The
Neopterygii Neopterygii (from Greek νέος ''neos'' 'new' and πτέρυξ ''pteryx'' 'fin') is a subclass of ray-finned fish (Actinopterygii). Neopterygii includes the Holostei and the Teleostei, of which the latter comprise the vast majority of extant ...
, in turn, is divided into the infraclasses
Holostei Holostei is a group of ray-finned bony fish. It is divided into two major clades, the Halecomorphi, represented by the single living genus, '' Amia'' with two species, the bowfins (''Amia calva'' and '' Amia ocellicauda''), as well as the Gin ...
and
Teleostei Teleostei (; Ancient Greek, Greek ''teleios'' "complete" + ''osteon'' "bone"), members of which are known as teleosts (), is, by far, the largest group of ray-finned fishes (class Actinopterygii), with 96% of all neontology, extant species of f ...
. During the
Mesozoic The Mesozoic Era is the Era (geology), era of Earth's Geologic time scale, geological history, lasting from about , comprising the Triassic, Jurassic and Cretaceous Period (geology), Periods. It is characterized by the dominance of archosaurian r ...
(
Triassic The Triassic ( ; sometimes symbolized 🝈) is a geologic period and system which spans 50.5 million years from the end of the Permian Period 251.902 million years ago ( Mya), to the beginning of the Jurassic Period 201.4 Mya. The Triassic is t ...
,
Jurassic The Jurassic ( ) is a Geological period, geologic period and System (stratigraphy), stratigraphic system that spanned from the end of the Triassic Period million years ago (Mya) to the beginning of the Cretaceous Period, approximately 143.1 Mya. ...
,
Cretaceous The Cretaceous ( ) is a geological period that lasted from about 143.1 to 66 mya (unit), million years ago (Mya). It is the third and final period of the Mesozoic Era (geology), Era, as well as the longest. At around 77.1 million years, it is the ...
) and
Cenozoic The Cenozoic Era ( ; ) is Earth's current geological era, representing the last 66million years of Earth's history. It is characterized by the dominance of mammals, insects, birds and angiosperms (flowering plants). It is the latest of three g ...
the teleosts in particular diversified widely. As a result, 96% of living fish species are teleosts (40% of all fish species belong to the teleost subgroup
Acanthomorpha Acanthomorpha (meaning "thorn-shaped") is an extraordinarily diverse taxon of teleost fishes with spiny fin rays. The clade contains about one-third of the world's modern species of vertebrates: over 14,000 species. A key anatomical innovatio ...
), while all other groups of actinopterygians represent depauperate lineages. The classification of ray-finned fishes can be summarized as follows: * Cladistia, which include bichirs and reedfish * Actinopteri, which include: ** Chondrostei, which include Acipenseriformes (paddlefishes and sturgeons) ** Neopterygii, which include: ***Teleostei (most living fishes) ***Holostei, which include: ****Lepisosteiformes (gars) ****Amiiformes (bowfin) The
cladogram A cladogram (from Greek language, Greek ''clados'' "branch" and ''gramma'' "character") is a diagram used in cladistics to show relations among organisms. A cladogram is not, however, an Phylogenetic tree, evolutionary tree because it does not s ...
below shows the main clades of living actinopterygians and their evolutionary relationships to other
extant Extant or Least-concern species, least concern is the opposite of the word extinct. It may refer to: * Extant hereditary titles * Extant literature, surviving literature, such as ''Beowulf'', the oldest extant manuscript written in English * Exta ...
groups of
fishes A fish (: fish or fishes) is an aquatic, anamniotic, gill-bearing vertebrate animal with swimming fins and a hard skull, but lacking limbs with digits. Fish can be grouped into the more basal jawless fish and the more common jawed ...
and the four-limbed vertebrates (
tetrapods A tetrapod (; from Ancient Greek τετρα- ''(tetra-)'' 'four' and πούς ''(poús)'' 'foot') is any four- limbed vertebrate animal of the clade Tetrapoda (). Tetrapods include all extant and extinct amphibians and amniotes, with the lat ...
). The latter include mostly terrestrial
species A species () is often defined as the largest group of organisms in which any two individuals of the appropriate sexes or mating types can produce fertile offspring, typically by sexual reproduction. It is the basic unit of Taxonomy (biology), ...
but also groups that became secondarily aquatic (e.g.
whales and dolphins Cetacea (; , ) is an infraorder of aquatic mammals belonging to the order Artiodactyla that includes whales, dolphins and porpoises. Key characteristics are their fully aquatic lifestyle, streamlined body shape, often large size and exclusively c ...
). Tetrapods
evolved Evolution is the change in the heritable Phenotypic trait, characteristics of biological populations over successive generations. It occurs when evolutionary processes such as natural selection and genetic drift act on genetic variation, re ...
from a group of
bony fish Osteichthyes ( ; ), also known as osteichthyans or commonly referred to as the bony fish, is a Biodiversity, diverse clade of vertebrate animals that have endoskeletons primarily composed of bone tissue. They can be contrasted with the Chondricht ...
during the
Devonian The Devonian ( ) is a period (geology), geologic period and system (stratigraphy), system of the Paleozoic era (geology), era during the Phanerozoic eon (geology), eon, spanning 60.3 million years from the end of the preceding Silurian per ...
period Period may refer to: Common uses * Period (punctuation) * Era, a length or span of time *Menstruation, commonly referred to as a "period" Arts, entertainment, and media * Period (music), a concept in musical composition * Periodic sentence (o ...
. Approximate
divergence In vector calculus, divergence is a vector operator that operates on a vector field, producing a scalar field giving the rate that the vector field alters the volume in an infinitesimal neighborhood of each point. (In 2D this "volume" refers to ...
dates for the different actinopterygian clades (in millions of years, mya) are from Near et al., 2012. The polypterids (bichirs and reedfish) are the
sister lineage In phylogenetics, a sister group or sister taxon, also called an adelphotaxon, comprises the closest relative(s) of another given unit in an evolutionary tree. Definition The expression is most easily illustrated by a cladogram: Taxon A and t ...
of all other actinopterygians, the Acipenseriformes (sturgeons and paddlefishes) are the sister lineage of Neopterygii, and Holostei (bowfin and gars) are the sister lineage of teleosts. The
Elopomorpha The superorder Elopomorpha contains a variety of types of fishes that range from typical silvery-colored species, such as the tarpons and ladyfishes of the Elopiformes and the bonefishes of the Albuliformes, to the long and slender, smooth-bodie ...
(
eel Eels are ray-finned fish belonging to the order Anguilliformes (), which consists of eight suborders, 20 families, 164 genera, and about 1000 species. Eels undergo considerable development from the early larval stage to the eventual adult stage ...
s and
tarpon Tarpon are fish of the genus ''Megalops''. They are the only members of the family Megalopidae. Of the two species, one (''M. atlanticus'') is native to the Atlantic, and the other (''M. cyprinoides'') to the Indo-Pacific Oceans. Species and ...
s) appear to be the most basal teleosts. The earliest known
fossil A fossil (from Classical Latin , ) is any preserved remains, impression, or trace of any once-living thing from a past geological age. Examples include bones, shells, exoskeletons, stone imprints of animals or microbes, objects preserve ...
actinopterygian is '' Andreolepis hedei'', dating back 420 million years (
Late Silurian The Silurian ( ) is a geologic period and system spanning 23.5 million years from the end of the Ordovician Period, at million years ago ( Mya), to the beginning of the Devonian Period, Mya. The Silurian is the third and shortest period of t ...
), remains of which have been found in
Russia Russia, or the Russian Federation, is a country spanning Eastern Europe and North Asia. It is the list of countries and dependencies by area, largest country in the world, and extends across Time in Russia, eleven time zones, sharing Borders ...
,
Sweden Sweden, formally the Kingdom of Sweden, is a Nordic countries, Nordic country located on the Scandinavian Peninsula in Northern Europe. It borders Norway to the west and north, and Finland to the east. At , Sweden is the largest Nordic count ...
, and
Estonia Estonia, officially the Republic of Estonia, is a country in Northern Europe. It is bordered to the north by the Gulf of Finland across from Finland, to the west by the Baltic Sea across from Sweden, to the south by Latvia, and to the east by Ru ...
. Crown group actinopterygians most likely originated near the Devonian-Carboniferous boundary. The earliest fossil relatives of modern teleosts are from the
Triassic The Triassic ( ; sometimes symbolized 🝈) is a geologic period and system which spans 50.5 million years from the end of the Permian Period 251.902 million years ago ( Mya), to the beginning of the Jurassic Period 201.4 Mya. The Triassic is t ...
period Period may refer to: Common uses * Period (punctuation) * Era, a length or span of time *Menstruation, commonly referred to as a "period" Arts, entertainment, and media * Period (music), a concept in musical composition * Periodic sentence (o ...
(''
Prohalecites ''Prohalecites'' is an extinct genus of ray-finned fish from the Ladinian and possibly Carnian (Triassic) of Italy. It is the oldest known teleosteomorph, a group that includes extant teleosts and their close fossil relatives. The type and on ...
'', ''
Pholidophorus ''Pholidophorus'' (from , 'horny scale' and , 'to bear') is an extinct genus of Stem group, stem-teleost fish. Numerous species were assigned to this genus in the past, but only the type species ''Pholidophorus latiusculus'', from the Late Tria ...
''), although it is suspected that teleosts originated already during the
Paleozoic The Paleozoic ( , , ; or Palaeozoic) Era is the first of three Era (geology), geological eras of the Phanerozoic Eon. Beginning 538.8 million years ago (Ma), it succeeds the Neoproterozoic (the last era of the Proterozoic Eon) and ends 251.9 Ma a ...
Era An era is a span of time. Era or ERA may also refer to: * Era (geology), a subdivision of geologic time * Calendar era Education * Academy of European Law (German: '), an international law school * ERA School, in Melbourne, Australia * E ...
.


Taxonomy

The listing below is a summary of all
extinct Extinction is the termination of an organism by the death of its Endling, last member. A taxon may become Functional extinction, functionally extinct before the death of its last member if it loses the capacity to Reproduction, reproduce and ...
(indicated by a
dagger A dagger is a fighting knife with a very sharp point and usually one or two sharp edges, typically designed or capable of being used as a cutting or stabbing, thrusting weapon.State v. Martin, 633 S.W.2d 80 (Mo. 1982): This is the dictionary or ...
, †) and living groups of Actinopterygii with their respective
taxonomic rank In biology, taxonomic rank (which some authors prefer to call nomenclatural rank because ranking is part of nomenclature rather than taxonomy proper, according to some definitions of these terms) is the relative or absolute level of a group of or ...
. The
taxonomy image:Hierarchical clustering diagram.png, 280px, Generalized scheme of taxonomy Taxonomy is a practice and science concerned with classification or categorization. Typically, there are two parts to it: the development of an underlying scheme o ...
follows ''
Eschmeyer's Catalog of Fishes Catalog of Fishes is a comprehensive on-line database and reference work on the scientific names of fish species and genera. It is global in its scope and is hosted by the California Academy of Sciences. It has been compiled and is continuously up ...
'' and Phylogenetic Classification of Bony Fishes with notes when this differs from Nelson,
ITIS The Integrated Taxonomic Information System (ITIS) is an American partnership of federal agencies designed to provide consistent and reliable information on the taxonomy of biological species. ITIS was originally formed in 1996 as an interagenc ...
and
FishBase FishBase is a global species database of fish species (specifically finfish). It is the largest and most extensively accessed online database on adult finfish on the web.
and extinct groups from Van der Laan 2016 and Xu 2021. * Order †?
Asarotiformes ''Asarotus'' is an extinct genus of prehistoric marine ray-finned fish from the Niobrara Formation of Nebraska, during the Campanian The Campanian is the fifth of six ages of the Late Cretaceous epoch on the geologic timescale of the Intern ...
Schaeffer 1968 * Order †? Discordichthyiformes Minikh 1998 * Order †? Paphosisciformes Grogan & Lund 2015 * Order †?
Scanilepiformes Cladistia is a subclass of bony fishes whose only living members are the bichirs of tropical Africa. Their major synapomorphies are a heterocercal tail in which the dorsal fin has independent rays, and a posteriorly elongated parasphenoid. Cl ...
Selezneya 1985 * Order † Cheirolepidiformes Kazantseva-Selezneva 1977 * Order † Paramblypteriformes Heyler 1969 * Order † Rhadinichthyiformes * Order †
Palaeonisciformes The Palaeonisciformes, commonly known as "palaeoniscoids" (also spelled "paleoniscoid", or alternatively "paleoniscids") are an extinct grouping of primitive ray-finned fish (Actinopterygii), spanning from the Silurian/Devonian to the Cretaceous. ...
Hay 1902 * Order †
Tarrasiiformes Tarasiiformes is an extinct order of prehistoric Actinopterygii, ray-finned fish. Taxonomy * Order †Tarrasiiformes sensu Lund & Poplin 2002 aplistia** Family †Tarrasiidae Traquair 1881 emend. Woodward 1891 *** Genus †''Apholidotos'' Lu ...
sensu Lund & Poplin 2002 * Order †
Ptycholepiformes Ptycholepiformes are an extinct order (biology), order of prehistoric Actinopterygii, ray-finned fish that existed during the Triassic period (geology), period and the Early Jurassic epoch (geology), epoch. The order includes the genera ''Acrorha ...
Andrews et al. 1967 * Order † Haplolepidiformes Westoll 1944 * Order † Aeduelliformes Heyler 1969 * Order † Platysomiformes Aldinger 1937 * Order † Dorypteriformes Cope 1871 * Order † Eurynotiformes Sallan & Coates 2013 * Subclass
Cladistia Cladistia is a subclass of bony fishes whose only living members are the bichirs of tropical Africa. Their major synapomorphies are a heterocercal tail in which the dorsal fin has independent rays, and a posteriorly elongated parasphenoid. ...
Pander 1860 ** Order †
Guildayichthyiformes Guildayichthyidae is a prehistoric family of marine fish from the Mississippian Bear Gulch Limestone of Montana Montana ( ) is a landlocked U.S. state, state in the Mountain states, Mountain West subregion of the Western United States. It ...
Lund 2000 ** Order
Polypteriformes Bichirs and the reedfish comprise Polypteridae , a family of archaic ray-finned fishes and the only family in the order Polypteriformes .Helfman GS, Collette BB, Facey DE, Bowen BW. 2009. The Diversity of Fishes. West Sussex, UK: Blackwell Pu ...
Bleeker 1859 (
bichir Bichirs and the reedfish comprise Polypteridae , a family (biology), family of archaic Actinopterygii, ray-finned fishes and the only family in the order (biology), order Polypteriformes .Helfman GS, Collette BB, Facey DE, Bowen BW. 2009. The D ...
s and
reedfish The reedfish, ropefish (more commonly used in the United States), or snakefish, ''Erpetoichthys calabaricus'', is a species of fish in the family Polypteridae alongside the bichirs. It is the only member of the genus ''Erpetoichthys''. It is n ...
es) ** Subclass
Actinopteri Actinopteri () is the sister group of Cladistia (bichirs) in the class Actinopterygii (ray-finned fish). Dating back to the Permian period, the Actinopteri comprise the Chondrostei (sturgeons and paddlefish), the Holostei ( bowfins and gars), ...
Cope 1972 s.s. *** Order † Elonichthyiformes Kazantseva-Selezneva 1977 *** Order † Phanerorhynchiformes *** Order †
Bobasatraniiformes Bobasatraniiformes is an extinct order of durophagous ray-finned fish that existed from the late Permian to the Middle Triassic in both marine and freshwater environments. The order includes two families: Bobasatraniidae, with the genera ''Bo ...
Berg 1940 *** Order †
Saurichthyiformes Saurichthyiformes is an extinct order (biology), order of Actinopterygii, ray-finned fish which existed in Asia, Africa, Australia (continent), Australia, Europe and North America, during the late Permian to early Middle Jurassic. Saurichthy ...
Aldinger 1937 ** Subclass
Chondrostei Chondrostei is a subclass of non- neopterygian ray-finned fish. While the term originally referred to the paraphyletic grouping of all non-neopterygian ray-finned fish, it was redefined by Patterson in 1982 to be a clade comprising the Acipenser ...
Müller, 1844 *** Order † Birgeriiformes Heyler 1969 *** Order †
Chondrosteiformes Chondrosteidae is a family of extinct marine actinopterygian fishes, known from the Early Jurassic of Europe. They are closely related to modern sturgeons and paddlefish of the order Acipenseriformes, and are either placed as part of that orde ...
Aldinger, 1937 *** Order
Acipenseriformes Acipenseriformes is an order (biology), order of basal (phylogenetics), basal Actinopterygii, ray-finned fishes that includes living and fossil sturgeons and paddlefishes (Acipenseroidei), as well as the extinct family (biology), families C ...
Berg 1940 (includes
sturgeon Sturgeon (from Old English ultimately from Proto-Indo-European language, Proto-Indo-European *''str̥(Hx)yón''-) is the common name for the 27 species of fish belonging to the family Acipenseridae. The earliest sturgeon fossils date to the ...
s and
paddlefish Paddlefish (family Polyodontidae) are a family of ray-finned fish belonging to order Acipenseriformes, and one of two living groups of the order alongside sturgeons (Acipenseridae). They are distinguished from other fish by their elongated rost ...
es) ** Subclass
Neopterygii Neopterygii (from Greek νέος ''neos'' 'new' and πτέρυξ ''pteryx'' 'fin') is a subclass of ray-finned fish (Actinopterygii). Neopterygii includes the Holostei and the Teleostei, of which the latter comprise the vast majority of extant ...
Regan 1923 sensu Xu & Wu 2012 *** Order †
Pholidopleuriformes Pholidopleuriformes is an extinct order of Actinopterygii, ray-finned fish containing a single family, Pholidopleuridae. Classification ** Family †Pholidopleuridae Othenio Abel, Abel, 1919 / Wade, 1932 *** Genus †''Arctosomus'' Berg 1941 ' ...
Berg 1937 *** Order †Redfieldiiformes Berg 1940 *** Order † Redfieldiiformes">' ...
Berg 1937 *** Order †Redfieldiiformes Berg 1940 *** Order †Platysiagiformes Brough 1939 *** Order †Polzbergiiformes">Platysiagiformes">Redfieldiiformes">' ...
Berg 1937 *** Order †Redfieldiiformes Berg 1940 *** Order †Platysiagiformes Brough 1939 *** Order †Polzbergiiformes Griffith 1977 *** Order †Perleidiformes Berg 1937 *** Order †Louwoichthyiformes Xu 2021 *** Order †Peltopleuriformes Lehman 1966 *** Order †Luganoiiformes Lehman 1958 *** Order †Pycnodontiformes Berg 1937 *** Infraclass
Holostei Holostei is a group of ray-finned bony fish. It is divided into two major clades, the Halecomorphi, represented by the single living genus, '' Amia'' with two species, the bowfins (''Amia calva'' and '' Amia ocellicauda''), as well as the Gin ...
Müller 1844 **** Division
Halecomorphi Halecomorphi is a taxon of Actinopterygii, ray-finned bony fish in the clade Neopterygii. The only extant Halecomorph species are the bowfin (''Amia calva'') and eyespot bowfin (''Amia ocellicauda''), but the group contains many extinct species i ...
Cope 1872 sensu Grande & Bemis 1998 ***** Order †
Parasemionotiformes Parasemionotiformes is an extinct order of neopterygian ray-finned fish that existed globally during the Triassic period. It comprises the families Parasemionotidae and Promecosominidae. Many of the included genera are monotypic and most specie ...
Lehman 1966 ***** Order † Ionoscopiformes Grande & Bemis 1998 ***** Order
Amiiformes The Amiiformes order (biology), order of fish has only two extant taxa, extant species, the bowfins: ''Amia calva'' and ''Amia ocellicauda'', the latter recognized as a separate species in 2022. These Amiiformes are found in the freshwater syste ...
Huxley 1861 sensu Grande & Bemis 1998 (
bowfin The ruddy bowfin (''Amia calva'') is a ray-finned fish native to North America. Common names include mudfish, mud pike, dogfish, grindle, grinnel, swamp trout, and choupique. It is regarded as a relict, being one of only two surviving species ...
s) **** Division
Ginglymodi Ginglymodi is a clade of ray-finned fish containing modern-day gars (Lepisosteidae) and their extinct relatives (including the family (biology), family Lepidotidae) in the Order (biology), order Lepisosteiformes, the extinct orders Semionotiforme ...
Cope 1871 ***** Order †
Dapediiformes Dapediidae is an extinct family of neopterygian ray-finned fish that lived from the Middle Triassic to Late Jurassic (Ladinian to Tithonian). It is the only family of the order Dapediiformes. Its members were historically placed within the gingly ...
Thies & Waschkewitz 2015 ***** Order †
Semionotiformes Semionotiformes is an order of ray-finned fish known from the Middle Triassic (Anisian) to the Late Cretaceous (Maastrichtian). Their closest living relatives are gars (Lepisosteidae), with both groups belonging to the clade Ginglymodi within th ...
Arambourg & Bertin 1958 ***** Order
Lepisosteiformes Lepisosteiformes is an order of ray-finned fish and the only living members of the clade Ginglymodi. Its only extant representatives are the gar (family Lepisosteidae), and it is defined as all members of Ginglymodi that are more closely rela ...
Hay 1929 (
gar Gars are an ancient group of ray-finned fish in the family Lepisosteidae. They comprise seven living species of fish in two genera that inhabit fresh, brackish, and occasionally marine waters of eastern North America, Central America and Cuba ...
s) *** Clade
Teleosteomorpha Teleosteomorpha is an infraclass of ray-finned fishes containing all teleost fish and their closest extinct relatives. Also in this group are two diverse Mesozoic fish orders, the Aspidorhynchiformes and the Pachycormiformes. Several other non- ...
Arratia 2000 sensu Arratia 2013 **** Order †
Prohaleciteiformes ''Prohalecites'' is an extinct genus of ray-finned fish from the Ladinian and possibly Carnian (Triassic) of Italy. It is the oldest known teleosteomorph, a group that includes extant teleosts and their close fossil relatives. The type and onl ...
Arratia 2017 **** Division
Aspidorhynchei Teleosteomorpha is an Class (biology), infraclass of Actinopterygii, ray-finned fishes containing all teleost fish and their closest extinct relatives. Also in this group are two diverse Mesozoic fish orders, the Aspidorhynchiformes and the Pachy ...
Nelson, Grand & Wilson 2016 ***** Order †
Aspidorhynchiformes Aspidorhynchidae (from Neo-Latin "shield-snouts") is an extinct Family (biology), family of Actinopterygii, ray-finned fish from the Mesozoic, Mesozoic Era. It is the only member of the Monotypic taxon, monotypic Order (biology), order Aspidorhync ...
Bleeker 1859 ***** Order †
Pachycormiformes Pachycormiformes is an extinct order of marine ray-finned fish known from the Early Jurassic to the end of the Cretaceous. It only includes a single family, Pachycormidae. They were characterized by having serrated pectoral fins (though more rece ...
Berg 1937 **** Infraclass
Teleostei Teleostei (; Ancient Greek, Greek ''teleios'' "complete" + ''osteon'' "bone"), members of which are known as teleosts (), is, by far, the largest group of ray-finned fishes (class Actinopterygii), with 96% of all neontology, extant species of f ...
Müller 1844 sensu Arratia 2013 ***** Order †?
Araripichthyiformes ''Araripichthys'' is an extinct genus of marine ray-finned fish that lived from the Aptian to Coniacian stages of the Cretaceous period. The genus is named after the Araripe Basin, where it was found in the Crato and Santana Formations. Other f ...
***** Order †? Ligulelliiformes Taverne 2011 ***** Order †?
Tselfatiiformes Tselfatiiformes is an extinct order of bony fishes from the infraclass Teleostei. The order represents the most important radiation of marine teleosts during the Cretaceous period. Fossils of tselfatiiforms are known from Europe, North America, c ...
Nelson 1994 ***** Order † Pholidophoriformes Berg 1940 ***** Order †
Dorsetichthyiformes ''Dorsetichthys'' is an extinct genus of marine stem-teleost ray-finned fish from the Early Jurassic period of Europe. It contains a single species, ''D. bechei'', known from the Sinemurian-aged Lower Lias of England. Description ''Dorsetichthys ...
Nelson, Grand & Wilson 2016 ***** Order †
Leptolepidiformes Leptolepidae (also spelt as Leptolepididae) is an extinct family of herring-like stem-teleost fish found throughout the world during the Jurassic. They were among the first fish to possess certain teleost synapomorphies, such as cycloid scales an ...
***** Order †
Crossognathiformes Crossognathiformes is an extinct order of ray-finned fish that lived from the Late Jurassic to the Eocene. Its phylogenetic placement is disputed; some authors have recovered it as part of the teleost stem group, while others place it in a basal ...
Taverne 1989 ***** Order †
Ichthyodectiformes Ichthyodectiformes is an Extinction, extinct order of marine stem-teleost ray-finned fish. The order is named after the genus ''Ichthyodectes'', established by Edward Drinker Cope in 1870. Ichthyodectiforms are usually considered to be some of th ...
Bardeck & Sprinkle 1969 ***** Teleocephala de Pinna 1996 s.s. ******Megacohort Elopocephalai Patterson 1977 sensu Arratia 1999 (
Elopomorpha The superorder Elopomorpha contains a variety of types of fishes that range from typical silvery-colored species, such as the tarpons and ladyfishes of the Elopiformes and the bonefishes of the Albuliformes, to the long and slender, smooth-bodie ...
Greenwood et al. 1966) ******* Order
Elopiformes The Elopiformes are the order of ray-finned fish including the tarpons, tenpounders, and ladyfish, as well as a number of extinct types. They have a long fossil record, easily distinguished from other fishes by the presence of an additional s ...
Gosline 1960 (
ladyfish The Elopidae are a family of ray-finned fish containing a single living genus ''Elops''. They are commonly known as ladyfish, skipjacks, jack-rashes, or tenpounders. The ladyfish are a coastal-dwelling fish found throughout the tropical and su ...
es and
tarpon Tarpon are fish of the genus ''Megalops''. They are the only members of the family Megalopidae. Of the two species, one (''M. atlanticus'') is native to the Atlantic, and the other (''M. cyprinoides'') to the Indo-Pacific Oceans. Species and ...
) ******* Order
Albuliformes Albulidae is a family of fish, commonly known as the bonefishes, that are popular as game fish in Florida, select locations in the South Pacific and the Bahamas (where two bonefish are featured on the 10-cent coin) and elsewhere. The family is s ...
Greenwood et al. 1966 sensu Forey et al. 1996 (bonefishes) ******* Order
Notacanthiformes The Notacanthiformes are an order of deep-sea ray-finned fishes, consisting of the families Halosauridae and Notacanthidae ( spiny eels). ''Fishes of the World'' (2006) lists it as the suborder Notacanthoidei of Albuliformes. The notacanthif ...
Goodrich 1909 ( halosaurs and
spiny eel The name spiny eel is used to describe members of two different families of fish: the freshwater Mastacembelidae of Asia and Africa, and the marine (and generally deep sea) Notacanthidae. Both are so-named because of their eel-like shape and sturdy ...
s) ******* Order
Anguilliformes Eels are ray-finned fish belonging to the order (biology), order Anguilliformes (), which consists of eight suborders, 20 Family (biology), families, 164 genus, genera, and about 1000 species. Eels undergo considerable development from the earl ...
Jarocki 1822 sensu Goodrich 1909 (true
eel Eels are ray-finned fish belonging to the order Anguilliformes (), which consists of eight suborders, 20 families, 164 genera, and about 1000 species. Eels undergo considerable development from the early larval stage to the eventual adult stage ...
s) ****** Megacohort Osteoglossocephalai sensu Arratia 1999 *******Supercohort Osteoglossocephala sensu Arratia 1999 (
Osteoglossomorpha Osteoglossomorpha is a group of bony fish in the Teleostei. Notable members A notable member is the arapaima (''Arapaima gigas''), the largest freshwater fish in South America and one of the largest bony fishes alive. Other notable members inclu ...
Greenwood et al. 1966) ******** Order † Lycopteriformes Chang & Chou 1977 ******** Order
Hiodontiformes Hiodontiformes is an order of ray-finned fish consisting of the two living species of the mooneye family, Hiodontidae, and three extinct genera. These are traditionally classified within the order Osteoglossiformes, a placement some authori ...
McAllister 1968 sensu Taverne 1979 (
mooneye Hiodontidae, commonly called mooneyes, is a family of ray-finned fish with a single included genus ''Hiodon''. The genus comprise two extant species native to North America and three to five extinct species recorded from Paleocene to Eocene age ...
and
goldeye The goldeye (''Hiodon alosoides'') is a freshwater fish found in Canada and the northern United States. It is one of only two extant species in the family Hiodontidae, the other species being ''Hiodon tergisus''. The species name ''alosoides' ...
) ******** Order
Osteoglossiformes Osteoglossiformes , meaning "bony tongues" in Ancient Greek, is a relatively primitive order of ray-finned fish that contains two sub-orders, the Osteoglossoidei and the Notopteroidei. All of at least 245 living species inhabit freshwater. They ...
Regan 1909 sensu Zhang 2004 (bony-tongued fishes) ******* Supercohort Clupeocephala Patterson & Rosen 1977 sensu Arratia 2010 ********Cohort Otomorpha Wiley & Johnson 2010 (
Otocephala Otocephala is a clade of ray-finned fishes within the infraclass Teleostei that evolved some 230 million years ago. It is named for the presence of a hearing (otophysic) link from the swimbladder to the inner ear. Other names proposed for the gro ...
; Ostarioclupeomorpha) ********* Subcohort Clupei Wiley & Johnson 2010 (
Clupeomorpha Clupeomorpha is a superorder of ray-finned fish which belongs to the clade Otocephala. Represented today only by the diverse, economically-important order Clupeiformes (containing herrings, anchovies and allies), it was formerly even more diverse ...
Greenwood et al. 1966) ********** Order †
Ellimmichthyiformes The Ellimmichthyiformes, also known as double-armored herrings, are an Extinction, extinct Order (biology), order of Actinopterygii, ray-finned fish known from the Early Cretaceous to the Oligocene. They were the sister group to the extant true h ...
Grande 1982 ********** Order
Clupeiformes Clupeiformes is the order of ray-finned fish that includes the herring family, Clupeidae, and the anchovy family, Engraulidae and sardines. The group includes many of the most important forage and food fish. Clupeiformes are physostomes, ...
Bleeker 1859 (
herring Herring are various species of forage fish, belonging to the Order (biology), order Clupeiformes. Herring often move in large Shoaling and schooling, schools around fishing banks and near the coast, found particularly in shallow, temperate wate ...
s and
anchovies An anchovy is a small, common forage fish of the family Engraulidae. Most species are found in marine waters, but several will enter brackish water, and some in South America are restricted to fresh water. More than 140 species are placed in 1 ...
) ********* Subcohort Alepocephali ********** Order
Alepocephaliformes Alepocephaliformes is an order of marine deep-sea teleost fishes. It was previously classified as the suborder Alepocephaloidei of the order Argentiniformes. As an adaptation to a life in the deep-sea, there is no swim bladder, and the ossifica ...
Marshall 1962 ********* Subcohort
Ostariophysi Ostariophysi is the second-largest superorder of fish. Members of this superorder are called ostariophysians. This diverse group contains 10,758 species, about 28% of known fish species in the world and 68% of freshwater species, and are present ...
Sagemehl 1885 ********** Section Anotophysa (Rosen & Greenwood 1970) Sagemehl 1885 *********** Order † Sorbininardiformes Taverne 1999 *********** Order
Gonorynchiformes The Gonorynchiformes are an order of ray-finned fish that includes the important food source, the milkfish (''Chanos chanos'', family Chanidae), and a number of lesser-known types, both marine and freshwater. The alternate spelling "Gonorhy ...
Regan 1909 (
milkfish The milkfish (''Chanos chanos'') is a widespread species of ray-finned fish found throughout the Indo-Pacific. It is the sole living species in the family Chanidae, and the only living member of the genus ''Chanos''. The repeating scientific na ...
es) ********** Section Otophysa Garstang 1931 *********** Order
Cypriniformes Cypriniformes is an order of ray-finned fish, which includes many families and genera of cyprinid ( carps and their kin) fish, such as barbs, loaches, botias, and minnows (among others). Cypriniformes is an "order-within-an-order", placed ...
Bleeker 1859 sensu Goodrich 1909 ( barbs,
carp The term carp (: carp) is a generic common name for numerous species of freshwater fish from the family (biology), family Cyprinidae, a very large clade of ray-finned fish mostly native to Eurasia. While carp are prized game fish, quarries and a ...
,
danios ''Danio'' is a genus of small freshwater fish in the family Cyprinidae found in South and Southeast Asia, commonly kept in aquaria. They are generally characterised by a pattern of horizontal stripes, rows of spots or vertical bars. Some species ...
,
goldfish The goldfish (''Carassius auratus'') is a freshwater fish in the family Cyprinidae of the order Cypriniformes. It is commonly kept as a pet in indoor aquariums, and is one of the most popular aquarium fish. Goldfish released into the w ...
es,
loaches Loaches are ray-finned fishes of the suborder Cobitoidei. They are freshwater, benthic (bottom-dwelling) fish found in rivers and creeks throughout Eurasia and northern Africa. Loaches are among the most diverse groups of fish; the 1249 known sp ...
,
minnow Minnow is the common name for a number of species of small freshwater fish, belonging to several genus, genera of the family Cyprinidae and in particular the subfamily Leuciscinae. They are also known in Ireland as wikt:pinkeen, pinkeens. While ...
s,
rasbora ''Rasbora'' is a genus of fish in the family Danionidae. They are native to freshwater habitats in South and Southeast Asia, as well as southeast China. A single species, ''R. gerlachi'', is only known from an old specimen that reputedly origina ...
s) *********** Order
Characiformes Characiformes is an order of ray-finned fish, comprising the characins and their allies. Grouped in 18 recognized families, more than 2000 different species are described, including the well-known piranha and tetras.; Buckup P.A.: "Relationsh ...
Goodrich 1909 (
characin Characiformes is an order of ray-finned fish, comprising the characins and their allies. Grouped in 18 recognized families, more than 2000 different species are described, including the well-known piranha and tetras.; Buckup P.A.: "Relationsh ...
s, pencilfishes, hatchetfishes,
piranha A piranha (, or ; ) is any of a number of freshwater fish species in the subfamily Serrasalminae, of the family Serrasalmidae, in the order Characiformes. These fish inhabit South American rivers, floodplains, lakes and reservoirs. Although ...
s,
tetra Tetra is the common name of many small freshwater characiform fishes. Tetras come from Africa, Central America, and South America, belonging to the biological families Characidae, Alestidae (the "African tetras"), Lepidarchidae, Lebiasi ...
s, dourado / golden (genus ''Salminus'') and
pacu Pacu () is a common name used to refer to several species of omnivorous South American freshwater Serrasalmidae, serrasalmid fish related to piranhas. Pacu and piranha do not have similar teeth, the main difference being jaw alignment; piranha ...
) *********** Order
Gymnotiformes The Gymnotiformes are an order of teleost bony fishes commonly known as Neotropical knifefish or South American knifefish. They have long bodies and swim using undulations of their elongated anal fin. Found almost exclusively in fresh water (the ...
Berg 1940 (
electric eel The electric eels are a genus, ''Electrophorus'', of neotropical freshwater fish from South America in the family Gymnotidae, of which they are the only members of the subfamily Electrophorinae. They are known for their electric fish, ability ...
s and knifefishes) *********** Order
Siluriformes Catfish (or catfishes; order Siluriformes or Nematognathi) are a diverse group of ray-finned fish. Catfish are named for their prominent barbels, which resemble a cat's whiskers, though not all catfish have prominent barbels or "whis ...
Cuvier 1817 sensu Hay 1929 (
catfish Catfish (or catfishes; order (biology), order Siluriformes or Nematognathi) are a diverse group of ray-finned fish. Catfish are common name, named for their prominent barbel (anatomy), barbels, which resemble a cat's whiskers, though not ...
es) ******** Cohort Euteleosteomorpha (Greenwood et al. 1966) (
Euteleostei Euteleostei, whose members are known as euteleosts, is a clade of bony fishes within Teleostei that evolved some 240 million years ago, although the oldest known fossil remains are only from the Early Cretaceous. It is divided into Protacanthopt ...
Greenwood 1967 sensu Johnson & Patterson 1996) *********Subcohort Lepidogalaxii ********** Order Lepidogalaxiiformes Betancur-Rodriguez et al. 2013 ( salamanderfish) ********* Subcohort
Protacanthopterygii Protacanthopterygii is a superorder of ray-finned fish. They inhabit both Marine (ocean), marine and freshwater habitats. They appear to have evolved in the Cretaceous or perhaps late Jurassic, originating probably roughly 150 million years ago; ...
Greenwood et al. 1966 sensu Johnson & Patterson 1996 ********** Order
Argentiniformes The Argentiniformes is an order of marine ray-finned fish whose distinctness was recognized only fairly recently. In former times, they were included in the Osmeriformes (typical smelt and allies) as suborder Argentinoidei. That term refers onl ...
(
barreleye Barreleyes, also known as spook fish (a name also applied to several species of chimaera (fish), chimaera), are small deep-sea Argentiniformes, argentiniform fish comprising the family (biology), family Opisthoproctidae found in tropical-to-temp ...
s and
slickhead Slickheads, also known as nakedheads or smoothheads, are deep water fishes that belong to the family Alepocephalidae. They are most commonly found in the bathypelagic layer, which is approximately 3000m below the surface. They get their name from ...
s) (formerly in Osmeriformes) ********** Order
Galaxiiformes The Galaxiidae are a family of mostly small freshwater fish in the Southern Hemisphere. The majority live in Southern Australia or New Zealand, but some are found in South Africa, southern South America, Lord Howe Island, New Caledonia, and th ...
********** Order
Salmoniformes Salmoniformes (, lit. "salmon-shaped") is an Order (biology), order of Actinopterygii, ray-finned fishes native to the temperate and subarctic Northern Hemisphere. It contains two suborders: Salmonoidei (containing only the Salmonidae) and Esocoi ...
Bleeker 1859 sensu Nelson 1994 (
salmon Salmon (; : salmon) are any of several list of commercially important fish species, commercially important species of euryhaline ray-finned fish from the genera ''Salmo'' and ''Oncorhynchus'' of the family (biology), family Salmonidae, native ...
,
trout Trout (: trout) is a generic common name for numerous species of carnivorous freshwater ray-finned fishes belonging to the genera '' Oncorhynchus'', ''Salmo'' and ''Salvelinus'', all of which are members of the subfamily Salmoninae in the ...
and
pike Pike, Pikes or The Pike may refer to: Fish * Blue pike or blue walleye, an extinct color morph of the yellow walleye ''Sander vitreus'' * Ctenoluciidae, the "pike characins", some species of which are commonly known as pikes * ''Esox'', genus of ...
) ********* Subcohort Stomiati ********** Order
Osmeriformes The Osmeriformes are an order (biology), order of ray-finned fish that includes the true or freshwater smelts and allies, such as the noodlefishes; they are also collectively called osmeriforms. They belong to the teleost superorder Protacanthopt ...
(
smelts Smelts are a family of small fish, the Osmeridae, found in the North Atlantic and North Pacific oceans, as well as rivers, streams and lakes in Europe, North America and Northeast Asia. They are also known as freshwater smelts or typical smelts ...
) ********** Order
Stomiiformes Stomiiformes is an order (biology), order of deep-sea ray-finned fishes of very diverse Morphology (biology), morphology. It includes, for example, Barbeled dragonfish, dragonfishes, lightfishes (Gonostomatidae and Phosichthyidae), loosejaws, ma ...
Regan 1909 (
bristlemouth The Gonostomatidae are a family of mesopelagic marine fish, commonly named bristlemouths, lightfishes, or anglemouths. It is a relatively small family, containing only eight known genera and 32 species. However, bristlemouths make up for their l ...
s and
marine hatchetfish Marine hatchetfishes or deep-sea hatchetfishes are small deep-sea mesopelagic ray-finned fish of the stomiiform subfamily Sternoptychinae. They should not be confused with the freshwater hatchetfishes, which are not particularly closely relate ...
es) ********* Subcohort
Neoteleostei The Neoteleostei is a large clade of teleostei, bony fish mostly consisting of marine clades. Only three lineages have freshwater species: Percopsiformes (Troutperches), which lives exclusively in freshwater, Gadiformes (cods), which is largely m ...
Nelson 1969 **********Infracohort Ateleopodia *********** Order
Ateleopodiformes The jellynose fishes or tadpole fishes are the small order Ateleopodiformes. This group of ray-finned fish is monotypic, containing a single family Ateleopodidae. It has about a dozen species in four genera, but these enigmatic fishes are in nee ...
(
jellynose fish The jellynose fishes or tadpole fishes are the small order Ateleopodiformes. This group of ray-finned fish is monotypic, containing a single family Ateleopodidae. It has about a dozen species in four genera, but these enigmatic fishes are in nee ...
) ********** Infracohort Eurypterygia Rosen 1973 ***********Section Aulopa yclosquamata Rosen 1973************ Order
Aulopiformes Aulopiformes is a diverse order (biology), order of Marine (ocean), marine ray-finned fish consisting of some 15 Extant taxon, extant and several prehistoric family (biology), families with about 45 genera and over 230 species. The common names ...
Rosen 1973 (
Bombay duck ''Harpadon nehereus'', commonly known as the Bombay duck or bummalo, is a species of lizardfish. Adults may reach a maximum length of , but the usual size is around . Etymology The small bomelon fish caught in Bombay (now Mumbai) was tradit ...
and
lancetfish Lancetfishes are large oceanic predatory ray-finned fishes in the genus ''Alepisaurus'' ("scaleless lizard") in the monogeneric family Alepisauridae. Lancetfishes grow up to in length. Very little is known about their biology, though they are ...
es) *********** Section Ctenosquamata Rosen 1973 ************Subsection Myctophata copelomorpha************* Order
Myctophiformes The Myctophiformes are an order of ray-finned fishes consisting of two families of deep-sea marine fish, most notably the highly abundant lanternfishes (Myctophidae). The blackchins (Neoscopelidae) contain six species in three genera, whi ...
Regan 1911 (
lanternfish Lanternfish (or myctophids, from the Greek language, Greek μυκτήρ ''myktḗr'', "nose" and ''ophis'', "serpent") are small mesopelagic fish of the large family (biology), family Myctophidae. One of two families in the order Myctophiformes, ...
es) ************ Subsection
Acanthomorpha Acanthomorpha (meaning "thorn-shaped") is an extraordinarily diverse taxon of teleost fishes with spiny fin rays. The clade contains about one-third of the world's modern species of vertebrates: over 14,000 species. A key anatomical innovatio ...
Betancur-Rodriguez et al. 2013 *************Division Lampridacea Betancur-Rodriguez et al. 2013 ampridomorpha; Lampripterygii************** Order
Lampriformes Lampriformes is an order (biology), order of Actinopterygii, ray-finned fish. Members are collectively called lamprids (which is more properly used for the Opah, Lampridae) or lampriforms, and unite such open-ocean and partially Deep sea, deep-s ...
Regan 1909 (
oarfish Oarfish are large and extremely long pelagic lampriform fish belonging to the small family (biology), family Regalecidae. Found in areas spanning from Temperate climate, temperate ocean zones to tropical ones, yet rarely seen by humans, the oa ...
,
opah The opah, also commonly known as moonfish, sunfish (not to be confused with Molidae), cowfish, kingfish, and redfin ocean pan are large, colorful, deep-bodied Pelagic zone, pelagic lampriform Fish, fishes comprising the genus ''Lampris'', of ...
and ribbonfishes) ************* Division Paracanthomorphacea sensu Grande et al. 2013 (
Paracanthopterygii Paracanthopterygii is a superorder of fishes. Members of this group are called paracanthopterygians. The oldest members are known as fossils from the Cenomanian. It includes five orders: * † Sphenocephaliformes * Percopsiformes ( trout-perche ...
Greenwood 1937) ************** Order
Percopsiformes The Percopsiformes are a small order of freshwater teleost fishes measuring less than 20 cm in length, comprising the trout-perch and its allies. It contains just ten extant taxon, extant species, grouped into seven genus, genera and three ...
Berg 1937 (
cavefishes Cavefish or cave fish is a generic term for fresh and brackish water fish adapted to life in caves and other underground habitats. Related terms are subterranean fish, Troglomorphism, troglomorphic fish, troglobitic fish, stygobitic fish, phreat ...
and
trout-perch ''Percopsis omiscomaycus'', also known as the trout-perch, the grounder or the sand minnow, is one of two species in the family Percopsidae. Its name comes from the Greek root words ''perc'', meaning perch and ''opsi'' meaning appearance. The spe ...
es) ************** Order † Sphenocephaliformes Rosen & Patterson 1969 ************** Order
Zeiformes The Zeiformes are a small order of exclusively marine ray-finned fishes most notable for the dories, a group of common food fish. The order consists of about 33 species in six extant families, mostly deep-sea types. The boarfishes ( Caproidae) ...
Regan 1909 ( dories) ************** Order
Gadiformes Gadiformes , also called the Anacanthini, are an order of ray-finned fish that include the cod, hakes, pollock, haddock, burbot, rocklings and moras, many of which are food fish of major commercial value. They are mostly marine fish found thr ...
Goodrich 1909 (
cod Cod (: cod) is the common name for the demersal fish genus ''Gadus'', belonging to the family (biology), family Gadidae. Cod is also used as part of the common name for a number of other fish species, and one species that belongs to genus ''Gad ...
s) ************* Division Polymixiacea Betancur-Rodriguez et al. 2013 (Polymyxiomorpha; Polymixiipterygii) ************** Order † Pattersonichthyiformes Gaudant 1976 ************** Order † Ctenothrissiformes Berg 1937 ************** Order
Polymixiiformes The beardfishes consist of a single extant genus, ''Polymixia'', of deep-sea marine ray-finned fish named for their pair of long hyoid barbels. They are classified in their own order Polymixiiformes . But as Nelson says, "few groups have been sh ...
Lowe 1838 (
beardfish The beardfishes consist of a single extant genus, ''Polymixia'', of deep-sea marine ray-finned fish named for their pair of long hyoid barbels. They are classified in their own order Polymixiiformes . But as Nelson says, "few groups have been sh ...
es) ************* Division Euacanthomorphacea Betancur-Rodriguez et al. 2013 (Euacanthomorpha sensu Johnson & Patterson 1993;
Acanthopterygii Acanthopterygii (meaning "spiny-finned one") is a superorder of teleost, bony fishes in the class Actinopterygii. Members of this superorder are sometimes called ray-finned fishes for the characteristic sharp, bony rays in their fins; however th ...
Gouan 1770 sensu]) **************Order
Trachichthyiformes The Trachichthyiformes are an order of ray-finned fishes in the superorder Acanthopterygii. Fossils of this group date back to the Cenomanian. Member genera include the flashlight fishes, fangtooth fishes, spinyfins, pineconefishes, redfishes, ...
(
fangtooth Fangtooths are beryciform fish of the family (biology), family Anoplogastridae (sometimes spelled "Anoplogasteridae") that live in the deep sea. The name is from the Greek language, Greek , meaning "unarmed", and (), meaning "stomach". With a wo ...
s and
pineconefish Pinecone fishes are small and unusual marine fish of the family Monocentridae. The family contains just five species in two genera, one of which is monotypic. Their distribution is limited to tropical and subtropical waters of the Indo-Pacific. ...
es) **************Subdivision Berycimorphaceae Betancur-Rodriguez et al. 2013 *************** Order
Beryciformes The Beryciformes are a poorly-understood Order (biology), order of carnivorous ray-finned fishes consisting of 7 families, 30 genera, and 161 species. They feed on small fish and invertebrates. Beyond this, little is known about the biology of m ...
( alfonsinos and holocentrids) (incl.
Holocentriformes Holocentridae is a family of ray-finned fish, the only family of the suborder Holocentroidei and the only family of the order Holocentriformes. The members of the subfamily Holocentrinae are typically known as squirrelfish, while the members of ...
,
Stephanoberyciformes The Stephanoberycoidei is a suborder of marine ray-finned fishes, consisting of about 68 species, the majority (61) of which belong to the ridgehead family (biology), family (Melamphaidae). They were formerly placed as their own order, the Stepha ...
;
Cetomimiformes The Cetomimoidea or whalefishes are a superfamily of small, deep-sea ray-finned fish. Formerly treated as either their own order (Cetomimiformes) due to their unusual anatomy, more recent studies incorporating genetic data confirm them to be de ...
) ************** Subdivision Percomorphaceae Betancur-Rodriguez et al. 2013 (
Percomorpha Percomorpha () is an extremely large and diverse clade of ray-finned fish. With more than 17,000 known species (including Scombroidei, tuna, Syngnathiformes, seahorses, gobies, Cichlidae, cichlids, flatfish, Labridae, wrasse, Perciformes, perches ...
sensu Miya et al. 2003;
Acanthopteri Perciformes (), also called the Acanthopteri, is an order or superorder of ray-finned fish in the clade Percomorpha. ''Perciformes'' means "perch-like". Among the well-known members of this group are perches and darters (Percidae), and also sea ...
) ***************Series Ophidiimopharia Betancur-Rodriguez et al. 2013 **************** Order
Ophidiiformes Ophidiiformes is an order of ray-finned fish that includes the cusk-eels (family Ophidiidae), pearlfishes (family Carapidae), viviparous brotulas (family Bythitidae), and others. Members of this order have small heads and long slender bodies. ...
(
pearlfish Pearlfish are marine fish in the ray-finned fish family Carapidae. Pearlfishes inhabit the tropical waters of the Atlantic, Indian, and Pacific Oceans at depths to , along oceanic shelves and slopes. They are slender, elongated fish with no s ...
es) *************** Series Batrachoidimopharia Betancur-Rodriguez et al. 2013 **************** Order
Batrachoidiformes Batrachoididae is the only family in the ray-finned fish order Batrachoidiformes . Members of this family are usually called toadfish or frogfish: both the English common name and scientific name refer to their toad-like appearance (''batrakhos ...
(
toadfishes {{short description, Common name used for fish Toadfish is the common name for a variety of species from several different families of fish, usually because of their toad-like appearance or calls using their swimbladder. File:OysterToadfish.jpeg, ...
) *************** Series Gobiomopharia Betancur-Rodriguez et al. 2013 **************** Order
Gobiiformes Gobiiformes (meaning "goby-like") is an order of percomorph fish containing three suborders: Apogonoidei, Trichonotoidei, and Gobioidei. The order was formerly defined as containing only the gobies (now placed within the Gobioidei). However, ...
(
cardinalfish Cardinalfishes are a family, Apogonidae, of ray-finned fishes found in the Atlantic, Indian, and Pacific Oceans; they are chiefly marine, but some species are found in brackish water and a few (notably '' Glossamia'') are found in fresh water. A ...
es, sleepers and
gobies Gobiidae or gobies is a family of bony fish in the order Gobiiformes, one of the largest fish families comprising over 2,000 species in more than 200 genera. Most of gobiid fish are relatively small, typically less than in length, and the fam ...
) *************** Series Scombrimopharia Betancur-Rodriguez et al. 2013 **************** Order
Syngnathiformes The Syngnathiformes are an order of ray-finned fishes that includes the leafy seadragons, sea moths, trumpetfishes and seahorses, among others.FishBase (2005)Order Summary for Syngnathiformes Version of 2005-FEB-15. Retrieved 19 Aug 2008. Th ...
( seahorses,
pipefish Pipefishes or pipe-fishes (Syngnathinae) are a subfamily of small fishes, which, together with the seahorses and seadragons ('' Phycodurus'' and '' Phyllopteryx''), form the family Syngnathidae. Description Pipefish look like straight-bodied s ...
es,
sea moth The seamoths make up a family of ray-finned fishes, the Pegasidae, within the order Syngnathiformes. They are named for Pegasus, a creature from Greek mythology. Seamoths are notable for their unusual appearance, including flattened bodies, the ...
s,
cornetfish The cornetfishes or flutemouths are a small family, the Fistulariidae, of extremely elongated fish in the order Syngnathiformes. The family consists of a single genus, ''Fistularia'', with four species, found worldwide in tropical and subtropical ...
es and flying gurnardsIn Nelson and ITIS,
Syngnathiformes The Syngnathiformes are an order of ray-finned fishes that includes the leafy seadragons, sea moths, trumpetfishes and seahorses, among others.FishBase (2005)Order Summary for Syngnathiformes Version of 2005-FEB-15. Retrieved 19 Aug 2008. Th ...
is placed as the suborder Syngnathoidei of the order
Gasterosteiformes Gasterosteoidei is a suborder of ray-finned fishes that includes the sticklebacks and relatives, the 5th edition of ''Fishes of the World'' classifies this suborder within the order Scorpaeniformes. Systematics Gasterosteoidei is treated as a ...
.
) **************** Order
Scombriformes Scombriformes, also known as Pelagia and Pelagiaria, is an order of ray-finned fish within the clade Percomorpha. It contains 287 extant species in 16 families, most of which were previously classified under the suborders Scombroidei and Stroma ...
(
Tuna A tuna (: tunas or tuna) is a saltwater fish that belongs to the tribe Thunnini, a subgrouping of the Scombridae ( mackerel) family. The Thunnini comprise 15 species across five genera, the sizes of which vary greatly, ranging from the bul ...
s and (
mackerel Mackerel is a common name applied to a number of different species of pelagic fish, mostly from the family Scombridae. They are found in both temperate and tropical seas, mostly living along the coast or offshore in the oceanic environment. ...
s) *************** Series Carangimopharia Betancur-Rodriguez et al. 2013 **************** Subseries Anabantaria Betancur-Rodriguez et al. 2014 ***************** Order
Synbranchiformes Synbranchiformes, often called swamp eels, though that name can also refer specifically to Synbranchidae, is an order of ray-finned fishes that are eel-like but have spiny rays, indicating that they belong to the superorder Acanthopterygii. ...
(
swamp eel The swamp eels (also written "swamp-eels") are a family (biology), family (Synbranchidae) of freshwater eel-like fishes of the tropics and subtropics. Most species are able to breathe air and typically live in marshes, ponds and damp places, som ...
s) ***************** Order
Anabantiformes The Anabantiformes , is an Order (biology), order of bony fish (Teleostei) proposed in 2009. They are collectively known as labyrinth fish, are an Order (biology), order of air-breathing freshwater Actinopterygii, ray-finned fish with three sub ...
(Labyrinthici) (
gourami Gouramis, or gouramies , are a group of fresh water, freshwater Anabantiformes, anabantiform fish that comprise the family (biology), family Osphronemidae. The fish are native to Asia—from the Indian Subcontinent to Southeast Asia and northeas ...
es, snakeheads, ) **************** Subseries Carangaria Betancur-Rodriguez et al. 2014 ***************** Order
Carangiformes Carangiformes is a large, diverse order of ray-finned fishes within the clade Percomorpha. It is part of a sister clade to the Ovalentaria, alongside its sister group, the Anabantaria (including Anabantiformes and Synbranchiformes). The Cara ...
(
Jack mackerel Jack mackerels or saurels are marine ray-finned fish in the genus ''Trachurus'' of the family Carangidae. The name of the genus derives from the Greek (language), Greek words ''trachys'' ("rough") and ''oura'' ("tail"). Some species, such as ''T ...
s,
pompano Pompanos ( ) are saltwater fish, marine fish in the genus ''Trachinotus'' in the family (biology), family Carangidae (better known as "slabs"). Pompano may also refer to various other, similarly shaped members of the Carangidae, or the Order (bi ...
s,
flatfish A flatfish is a member of the Ray-finned fish, ray-finned demersal fish Order (biology), suborder Pleuronectoidei, also called the Heterosomata. In many species, both eyes lie on one side of the head, one or the other migrating through or around ...
es, billfishes) **************** Subseries Ovalentaria Smith & Near 2012 (
Stiassnyiformes Stiassnyiformes is an order of bony fish (Teleostei) proposed in 2009.Blaise Li, Agnès Dettaï, Corinne Cruaud, Arnaud Couloux, Martine Desoutter-Meniger, Guillaume Lecointre: ''RNF213, a new nuclear marker for acanthomorph phylogeny.'' Molecula ...
sensu Li et al. 2009) ***************** Order
Atheriniformes The Atheriniformes, also known as the silversides, are an order of ray-finned fishes that includes the Old & New World silversides, the rainbowfishes, and several less-familiar families, including the unusual Phallostethidae. The order incl ...
Rosen 1964 ( silversides and
rainbowfish Rainbowfishes are small, colourful freshwater fishes belonging to the Family (biology), family Melanotaeniidae, found in northern and eastern Australia, New Guinea (including islands in Cenderawasih Bay and Raja Ampat Islands in Indonesia), Sula ...
es) ***************** Order
Cyprinodontiformes Cyprinodontiformes is an order (biology), order of Actinopterygii, ray-finned fish, comprising mostly small, freshwater fish. Many popular aquarium fish, such as killifish and Poeciliidae, live-bearers, are included. They are closely related t ...
Berg 1940 (
livebearers Livebearers are fish that retain their eggs inside the body and give birth to live, free-swimming young. They are especially prized by aquarium owners. Among aquarium fish, livebearers are nearly all members of the family Poeciliidae and include: ...
,
killifish A killifish is any of various oviparous (egg-laying) cyprinodontiform fish, including families Aplocheilidae, Pantanodontidae, Cyprinodontidae, Fundulidae, Nothobranchiidae, Profundulidae, Aphaniidae and Valenciidae. All together, there ar ...
es) ***************** Order
Beloniformes Beloniformes is an order composed of six families (and about 264 species) of freshwater and marine ray-finned fish: * Adrianichthyidae (ricefish and medakas) * Belonidae (needlefish) * Exocoetidae (flyingfishes) * Hemiramphidae (halfb ...
Berg 1940 ( flyingfishes and
ricefish The ricefishes are a family (biology), family (Adrianichthyidae) of small ray-finned fish that are found in fresh and brackish waters from India to Japan and out into the Malay Archipelago, most notably Sulawesi (where the Lake Poso and Lore Lin ...
es) ***************** Order
Cichliformes Cichliformes is an order of fishes. Its members were previously classified under the order Perciformes, but now many authorities consider it to be an independent order within the subseries Ovalentaria. The earliest fossils are known from the Eoc ...
Betancur-Rodriguez et al. 2013 (
Cichlid Cichlids () are a large, diverse, and widespread family of percomorph fish in the family Cichlidae, order Cichliformes. At least 1,760 species have been scientifically described, making it one of the largest vertebrate families, with on ...
s, Convict blenny, leaf fishes) ***************** Order
Mugiliformes Mugiliformes is a small order of percomorph fish in the clade Ovalentaria. This order is recognized by the Catalog of Fishes (2025). It contains only two families: * Mugilidae Jarocki, 1822 - mullets (57 species) * Ambassidae Klunzinger, 1870 ...
Berg 1940 ( mullets) ***************** Order
Blenniiformes Blenniiformes is an order of percomorph fish in the clade Ovalentaria, of which it is the most diverse group. This order contains several well-known fish groups such as blennies and damselfish. The term is derived from the Latin word '' blenn ...
Springer 1993 (
Blennies Blennies (from the Greek and , mucus, slime) are a diverse clade of ray-finned fish in the suborder Blennioidei of the percomorph order Blenniiformes. They inhabit marine, brackish, and occasionally freshwater habitats, and generally share simi ...
,
damselfish Damselfish are those fish within the subfamilies Abudefdufinae, Chrominae, Lepidozyginae, Pomacentrinae, and Stegastinae within the family Pomacentridae. Most species within this group are relatively small, although the four largest speci ...
,
Clingfish Clingfishes are ray-finned fishes of the family Gobiesocidae, the only family in the suborder Gobiesocoidei of the order Blenniiformes. These fairly small to very small fishes are widespread in tropical and temperate regions, mostly near the coa ...
es) *************** Series Eupercaria Betancur-Rodriguez et al. 2014 (Percomorpharia Betancur-Rodriguez et al. 2013) ****************Order
Perciformes Perciformes (), also called the Acanthopteri, is an order or superorder of ray-finned fish in the clade Percomorpha. ''Perciformes'' means " perch-like". Among the well-known members of this group are perches and darters ( Percidae), and als ...
Bleeker 1859 ****************Order
Centrarchiformes Centrarchiformes is an order of ray-finned fish, previously included amongst the perciformes. Froese, R. and D. Pauly. Editors. (2023). FishBase. Centrarchiformes. Accessed through: World Register of Marine Species at: https://www.marinespecies. ...
Bleeker 1859 ( Sunfishes and mandarin fishes) ****************Order
Labriformes Labriformes is an Order (biology), order of ray-finned fishes within the clade Percomorpha. Some authors include the Labriformes as the clade Labroidei within the Perciformes while others include more Family (biology), families within the Labrif ...
(
Wrasse The wrasses are a family, Labridae, of marine ray-finned fish, many of which are brightly colored. The family is large and diverse, with over 600 species in 81 genera, which are divided into nine subgroups or tribes. They are typically small, ...
s and
Parrotfish Parrotfish (named for their mouths, which resemble a parrot's beak) are a clade of fish placed in the tribe Scarini of the wrasse family (Labridae). Traditionally treated as their own family (Scaridae), genetic studies have found them to be dee ...
es) ****************Order
Acropomatiformes The Acropomatiformes or Pempheriformes are an order of fish from the group of perch relatives Percomorpha. The relationship of the families assigned to the Acropomatiformes is based on molecular biological studies and is not yet supported by mor ...
****************Order
Acanthuriformes Acanthuriformes is a large, diverse order of mostly marine ray-finned fishes, part of the Percomorpha clade. In the past, members of this clade were placed in the suborders Acanthuroidea and Percoidea of the order Perciformes, but this treatment ...
**************** Order
Lophiiformes The anglerfish are ray-finned fish in the order Lophiiformes (). Both the order's common and scientific name comes from the characteristic mode of predation, in which a modified dorsal fin ray acts as a lure for prey (akin to a human angler, ...
Garman 1899 (
Anglerfish The anglerfish are ray-finned fish in the order Lophiiformes (). Both the order's common name, common and scientific name comes from the characteristic mode of predation, in which a modified dorsal Fish fin#Ray-fins, fin ray acts as a Aggressiv ...
es) **************** Order
Tetraodontiformes Tetraodontiformes (), also known as the Plectognathi, is an order of ray-finned fishes which includes the pufferfishes and related taxa. This order has been classified as a suborder of the order Perciformes, although recent studies have found ...
Regan 1929 (
Filefish The filefish (Monacanthidae) are a diverse family of tropical to subtropical tetraodontiform marine fish, which are also known as foolfish, leatherjackets, or shingles. They live in the Atlantic, Pacific, and Indian Oceans. Filefish are closely ...
es and
pufferfish Tetraodontidae is a family of marine and freshwater fish in the order Tetraodontiformes. The family includes many familiar species variously called pufferfish, puffers, balloonfish, blowfish, blowers, blowies, bubblefish, globefish, swellfis ...
)


References


External links

* * {{Authority control Fish classes Extant Silurian first appearances