Ragnar Arthur Granit (30 October 1900 – 12 March 1991) was a Finnish-Swedish

scientist A scientist is a person who Scientific method, researches to advance knowledge in an Branches of science, area of the natural sciences. In classical antiquity, there was no real ancient analog of a modern scientist. Instead, philosophers engag ...

who was awarded the

Nobel Prize in Physiology or Medicine The Nobel Prize in Physiology or Medicine () is awarded yearly by the Nobel Assembly at the Karolinska Institute for outstanding discoveries in physiology or medicine. The Nobel Prize is not a single prize, but five separate prizes that, acco ...

in 1967 along with Haldan Keffer Hartline and George Wald "for their discoveries concerning the primary

physiological Physiology (; ) is the science, scientific study of function (biology), functions and mechanism (biology), mechanisms in a life, living system. As a branches of science, subdiscipline of biology, physiology focuses on how organisms, organ syst ...

and

chemical A chemical substance is a unique form of matter with constant chemical composition and characteristic properties. Chemical substances may take the form of a single element or chemical compounds. If two or more chemical substances can be combin ...

visual processes in the

eye An eye is a sensory organ that allows an organism to perceive visual information. It detects light and converts it into electro-chemical impulses in neurons (neurones). It is part of an organism's visual system. In higher organisms, the ey ...

Early life and education

Ragnar Arthur Granit was born on 30 October 1900 in

Riihimäki Riihimäki (; ) is a town and municipality in the south of Finland, about north of Helsinki and southeast of Tampere. An important railway junction is located in Riihimäki, since railway tracks from Riihimäki lead to Helsinki, Tampere and La ...

Finland Finland, officially the Republic of Finland, is a Nordic country in Northern Europe. It borders Sweden to the northwest, Norway to the north, and Russia to the east, with the Gulf of Bothnia to the west and the Gulf of Finland to the south, ...

, at the time part of the

Russian Empire The Russian Empire was an empire that spanned most of northern Eurasia from its establishment in November 1721 until the proclamation of the Russian Republic in September 1917. At its height in the late 19th century, it covered about , roughl ...

, into a Swedish-speaking Finnish family. Granit was raised in Oulunkylä, a suburb of the Finnish capital of

Helsinki Helsinki () is the Capital city, capital and most populous List of cities and towns in Finland, city in Finland. It is on the shore of the Gulf of Finland and is the seat of southern Finland's Uusimaa region. About people live in the municipali ...

, and attended the Svenska normallyceum in Helsinki. Granit graduated from the Faculty of Medicine at the

University of Helsinki The University of Helsinki (, ; UH) is a public university in Helsinki, Finland. The university was founded in Turku in 1640 as the Royal Academy of Åbo under the Swedish Empire, and moved to Helsinki in 1828 under the sponsorship of Alexander ...

in 1927.

Career and research

In 1940, when Finland became the target of a massive

Soviet The Union of Soviet Socialist Republics. (USSR), commonly known as the Soviet Union, was a List of former transcontinental countries#Since 1700, transcontinental country that spanned much of Eurasia from 1922 until Dissolution of the Soviet ...

attack during the

Winter War The Winter War was a war between the Soviet Union and Finland. It began with a Soviet invasion of Finland on 30 November 1939, three months after the outbreak of World War II, and ended three and a half months later with the Moscow Peac ...

, Granit sought refuge – and peaceful surroundings for his studies and research work – in

Stockholm Stockholm (; ) is the Capital city, capital and List of urban areas in Sweden by population, most populous city of Sweden, as well as the List of urban areas in the Nordic countries, largest urban area in the Nordic countries. Approximately ...

, the capital of neighbouring

Sweden Sweden, formally the Kingdom of Sweden, is a Nordic countries, Nordic country located on the Scandinavian Peninsula in Northern Europe. It borders Norway to the west and north, and Finland to the east. At , Sweden is the largest Nordic count ...

, at the age of 40. In 1941, Granit received Swedish citizenship, which made it possible for him to live and continue with his work without having to worry about the

Continuation War The Continuation War, also known as the Second Soviet–Finnish War, was a conflict fought by Finland and Nazi Germany against the Soviet Union during World War II. It began with a Finnish declaration of war on 25 June 1941 and ended on 19 ...

, which lasted in Finland until 1944. Granit was proud of his Finnish-Swedish roots and remained a patriotic Finnish-Swede throughout his life, maintaining homes in both Finland and Sweden after the

Moscow Armistice The Moscow Armistice was signed between Finland on one side and the Soviet Union and United Kingdom on the other side on 19 September 1944, ending the Continuation War. The Armistice restored the Moscow Peace Treaty of 1940, with a number of modi ...

ended the Continuation War and secured Finnish independence. Granit was professor of

neurophysiology Neurophysiology is a branch of physiology and neuroscience concerned with the functions of the nervous system and their mechanisms. The term ''neurophysiology'' originates from the Greek word ''νεῦρον'' ("nerve") and ''physiology'' (whic ...

at the

Karolinska Institute The Karolinska Institute (KI; ; sometimes known as the (Royal) Caroline Institute in English) is a research-led medical university in Solna within the Stockholm urban area of Sweden and one of the foremost medical research institutes globally ...

from 1946 to his retirement in 1967.

Awards and honors

Granit was elected an International Member of the

American Philosophical Society The American Philosophical Society (APS) is an American scholarly organization and learned society founded in 1743 in Philadelphia that promotes knowledge in the humanities and natural sciences through research, professional meetings, publicat ...

in 1954. In 1960, Granit was elected a Foreign Member of the

Royal Society The Royal Society, formally The Royal Society of London for Improving Natural Knowledge, is a learned society and the United Kingdom's national academy of sciences. The society fulfils a number of roles: promoting science and its benefits, re ...

(ForMemRS). In 1967 he was awarded the

. Granit said that he was a "fifty-fifty" Finnish and Swedish Nobel laureate. He was elected an International Member of the

United States National Academy of Sciences The National Academy of Sciences (NAS) is a United States nonprofit, non-governmental organization. NAS is part of the National Academies of Sciences, Engineering, and Medicine, along with the National Academy of Engineering (NAE) and the Nati ...

the following year. In 1971, he was elected an International Honorary Member of the

American Academy of Arts and Sciences The American Academy of Arts and Sciences (The Academy) is one of the oldest learned societies in the United States. It was founded in 1780 during the American Revolution by John Adams, John Hancock, James Bowdoin, Andrew Oliver, and other ...

Death

Granit died on 12 March 1991 in Stockholm at the age of 90. Granit and his wife Marguerite, who died the same year, were buried in a church cemetery on the Finnish island of

Korpo Korpo (; ) is an island located in the Turku archipelago. It is a former municipality of Finland. On 1 January 2009, it was consolidated with Houtskär, Iniö, Nagu and Pargas to form the new town of Väståboland. As of 1 January 2012, Vä ...

References

External links

* including the Nobel Lecture 12 December 1967 ''The Development of Retinal Neurophysiology'' * {{DEFAULTSORT:Granit, Ragnar 1900 births 1991 deaths Nobel laureates in Physiology or Medicine Finnish Nobel laureates Swedish Nobel laureates Academic staff of the Karolinska Institute Foreign members of the Royal Society Foreign associates of the National Academy of Sciences People from Vantaa Swedish-speaking Finns Finnish neuroscientists Swedish neuroscientists Finnish physiologists Finnish emigrants to Sweden 20th-century Finnish physicians Vitamin researchers International members of the American Philosophical Society