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Prodidominae is a family of
spider Spiders (order Araneae) are air-breathing arthropods that have eight legs, chelicerae with fangs generally able to inject venom, and spinnerets that extrude silk. They are the largest order of arachnids and rank seventh in total species d ...
, sometimes called long-spinneret ground spiders. It was formerly regarded as a subfamily of
Gnaphosidae Ground spiders comprise Gnaphosidae, the seventh largest spider family with over 2,000 described species in over 100 genera distributed worldwide. There are 105 species known to central Europe, and common genera include '' Gnaphosa'', '' Drassode ...
, but was raised to a family in 2022. Spiders in the family are easily identified by the greatly elongated base of the piriform gland spigots. At least parts of their body are covered with shiny scales or setae. The posterior median eyes are flat and silvery, with a triangular, egg-shaped or irregularly rectangular shape.Barbara Baehr
Prodidomidae
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Biology

Spiders in the Prodidominae are ground dwellers. Most species are nocturnal and hide during the day in litter, but ''Myandra'' species, which are probably mimicking ants, seem to be active during the day. The genus ''Zimiris'' is synanthropic and thus found throughout the tropics.


Distribution

Although ''
Theuma walteri Theuma is a municipality in the Vogtlandkreis district, in Saxony, Germany Germany, officially the Federal Republic of Germany (FRG),, is a country in Central Europe. It is the most populous member state of the European Union. Germany ...
'' was described from
Turkmenistan Turkmenistan ( or ; tk, Türkmenistan / Түркменистан, ) is a country located in Central Asia, bordered by Kazakhstan to the northwest, Uzbekistan to the north, east and northeast, Afghanistan to the southeast, Iran to the s ...
by
Eugène Simon Eugène Louis Simon (; 30 April 1848 – 17 November 1924) was a French naturalist who worked particularly on insects and spiders, but also on birds and plants. He is by far the most prolific spider taxonomist in history, describing over 4 ...
, it is suspected that Simon accidentally exchanged its locality with that of ''
Anagraphis ''Anagraphis'' is a genus of ground spiders that was first described by Eugène Simon in 1893. Originally placed in the now unrecognized family Prodidomidae, it was moved to the family Gnaphosidae in 2006. Species it contains seven species from ...
pallens'' (
Gnaphosidae Ground spiders comprise Gnaphosidae, the seventh largest spider family with over 2,000 described species in over 100 genera distributed worldwide. There are 105 species known to central Europe, and common genera include '' Gnaphosa'', '' Drassode ...
); then ''T. walteri'' would have been collected in the
Cape of Good Hope The Cape of Good Hope ( af, Kaap die Goeie Hoop ) ;''Kaap'' in isolation: pt, Cabo da Boa Esperança is a rocky headland on the Atlantic coast of the Cape Peninsula in South Africa. A common misconception is that the Cape of Good Hope is ...
, while ''A. pallens'' is from Turkmenistan.Platnick & Baehr 2006


Genera

Genera included: * '' Anagrina'' Berland, 1920
Africa Africa is the world's second-largest and second-most populous continent, after Asia in both cases. At about 30.3 million km2 (11.7 million square miles) including adjacent islands, it covers 6% of Earth's total surface area ...
* '' Austrodomus'' Lawrence, 1947
South Africa South Africa, officially the Republic of South Africa (RSA), is the southernmost country in Africa. It is bounded to the south by of coastline that stretch along the South Atlantic and Indian Oceans; to the north by the neighbouring count ...
* '' Brasilomma'' Brescovit, Ferreira & Rheims, 2012 * '' Caudalia'' Alayón, 1980
Cuba Cuba ( , ), officially the Republic of Cuba ( es, República de Cuba, links=no ), is an island country comprising the island of Cuba, as well as Isla de la Juventud and several minor archipelagos. Cuba is located where the northern Caribbea ...
* '' Chileomma'' Platnick, Shadab & Sorkin, 2005
Chile Chile, officially the Republic of Chile, is a country in the western part of South America. It is the southernmost country in the world, and the closest to Antarctica, occupying a long and narrow strip of land between the Andes to the eas ...
* '' Chileuma'' Platnick, Shadab & Sorkin, 2005 — Chile * '' Chilongius'' Platnick, Shadab & Sorkin, 2005 — Chile * '' Cryptoerithus''
Rainbow A rainbow is a meteorological phenomenon that is caused by reflection, refraction and dispersion of light in water droplets resulting in a spectrum of light appearing in the sky. It takes the form of a multicoloured circular arc. Rainbows ...
, 1915
Australia * '' Eleleis'' Simon, 1893 — South Africa * ''
Encoptarthria ''Encoptarthria'' is a genus of Australian ground spiders that was first described by Barbara York Main in 1954. Originally placed in the no longer recognized family Prodidomidae, it was moved to the family Gnaphosidae in 2007. Species it cont ...
'' Main, 1954 — Australia * '' Indiani'' Rodrigues, Cizauskas & Lemos, 2020 — Brazil * ''
Katumbea ''Katumbea'' is a monotypic genus of Tanzanian ground spiders containing the single species, ''Katumbea oxoniensis''. It was first described by J. A. L. Cooke in 1964, and is only found in Tanzania. See also * List of Gnaphosidae species T ...
'' Cooke, 1964
Tanzania Tanzania (; ), officially the United Republic of Tanzania ( sw, Jamhuri ya Muungano wa Tanzania), is a country in East Africa within the African Great Lakes region. It borders Uganda to the north; Kenya to the northeast; Comoro Islands ...
* '' Lygromma'' Simon, 1893Costa Rica to
Brazil Brazil ( pt, Brasil; ), officially the Federative Republic of Brazil (Portuguese: ), is the largest country in both South America and Latin America. At and with over 217 million people, Brazil is the world's fifth-largest country by area ...
, Galapagos * '' Lygrommatoides'' Strand, 1918Japan * ''
Moreno Moreno may refer to: Places Argentina *Moreno (Buenos Aires Metro), a station on Line C of the Buenos Aires Metro *Moreno, Buenos Aires, a city in Buenos Aires Province, Argentina *Moreno Department, a depatnent of Santiago del Estero Province, A ...
'' Mello-Leitão, 1940 — Chile,
Argentina Argentina (), officially the Argentine Republic ( es, link=no, República Argentina), is a country in the southern half of South America. Argentina covers an area of , making it the second-largest country in South America after Brazil, t ...
* ''
Myandra ''Myandra'' is a genus of Oceania, South Pacific Gnaphosidae, ground spiders that was first described by Eugène Simon, Eugène Louis Simon in 1887. Species it contains four species, found only in Australia: *''Myandra bicincta'' Simon, 1908 – ...
'' Simon, 1887 — Australia * '' Namundra'' Platnick & Bird, 2007 — Africa * '' Neozimiris'' Simon, 1903 — USA,
Mexico Mexico ( Spanish: México), officially the United Mexican States, is a country in the southern portion of North America. It is bordered to the north by the United States; to the south and west by the Pacific Ocean; to the southeast by Guate ...
,
Panama Panama ( , ; es, link=no, Panamá ), officially the Republic of Panama ( es, República de Panamá), is a transcontinental country spanning the southern part of North America and the northern part of South America. It is bordered by Co ...
, Galapagos, Bahamas * ''
Nomindra ''Nomindra'' is a genus of Australia, Australian Gnaphosidae, ground spiders that was first described by Norman I. Platnick & Barbara Baehr in 2006. Originally placed with the Prodidomidae, long-spinneret ground spiders, it was transferred to the ...
'' Platnick & Baehr, 2006 — Australia * '' Nopyllus'' Ott, 2014 — Brazil * '' Oltacloea'' Mello-Leitão, 1940 — Brazil, Argentina * '' Paracymbiomma'' Rodrigues, Cizauskas & Rheims, 2018 * '' Plutonodomus'' Cooke, 1964 — Tanzania * '' Prodida'' Dalmas, 1919
Philippines The Philippines (; fil, Pilipinas, links=no), officially the Republic of the Philippines ( fil, Republika ng Pilipinas, links=no), * bik, Republika kan Filipinas * ceb, Republika sa Pilipinas * cbk, República de Filipinas * hil, Republ ...
,
Seychelles Seychelles (, ; ), officially the Republic of Seychelles (french: link=no, République des Seychelles; Creole: ''La Repiblik Sesel''), is an archipelagic state consisting of 115 islands in the Indian Ocean. Its capital and largest city, ...
* '' Prodidomus'' Hentz, 1847
Mediterranean The Mediterranean Sea is a sea connected to the Atlantic Ocean, surrounded by the Mediterranean Basin and almost completely enclosed by land: on the north by Western and Southern Europe and Anatolia, on the south by North Africa, and on th ...
, Africa, Australia,
Asia Asia (, ) is one of the world's most notable geographical regions, which is either considered a continent in its own right or a subcontinent of Eurasia, which shares the continental landmass of Afro-Eurasia with Africa. Asia covers an ...
,
Venezuela Venezuela (; ), officially the Bolivarian Republic of Venezuela ( es, link=no, República Bolivariana de Venezuela), is a country on the northern coast of South America, consisting of a continental landmass and many islands and islets in ...
,
Hawaii Hawaii ( ; haw, Hawaii or ) is a state in the Western United States, located in the Pacific Ocean about from the U.S. mainland. It is the only U.S. state outside North America, the only state that is an archipelago, and the only ...
* '' Purcelliana'' Cooke, 1964 — South Africa * ''
Theuma Theuma is a municipality in the Vogtlandkreis The Vogtlandkreis () is a ''Landkreis'' (rural district) in the southwest of Saxony, Germany, at the borders to Thuringia, Bavaria, and the Czech Republic. Neighboring districts are (from south cloc ...
'' Simon, 1893 — Africa,
Turkmenistan Turkmenistan ( or ; tk, Türkmenistan / Түркменистан, ) is a country located in Central Asia, bordered by Kazakhstan to the northwest, Uzbekistan to the north, east and northeast, Afghanistan to the southeast, Iran to the s ...
? * '' Theumella'' Strand, 1906
Ethiopia Ethiopia, , om, Itiyoophiyaa, so, Itoobiya, ti, ኢትዮጵያ, Ítiyop'iya, aa, Itiyoppiya officially the Federal Democratic Republic of Ethiopia, is a landlocked country in the Horn of Africa. It shares borders with Eritrea to the Er ...
* '' Tivodrassus'' Chamberlin & Ivie, 1936 — Mexico * '' Tricongius'' Simon, 1892
South America South America is a continent entirely in the Western Hemisphere and mostly in the Southern Hemisphere, with a relatively small portion in the Northern Hemisphere at the northern tip of the continent. It can also be described as the souther ...
* '' Zimirina'' Dalmas, 1919
Spain , image_flag = Bandera de España.svg , image_coat = Escudo de España (mazonado).svg , national_motto = '' Plus ultra'' ( Latin)(English: "Further Beyond") , national_anthem = (English: "Royal March") , ...
,
Algeria ) , image_map = Algeria (centered orthographic projection).svg , map_caption = , image_map2 = , capital = Algiers , coordinates = , largest_city = capital , religi ...
,
Canary Islands The Canary Islands (; es, Canarias, ), also known informally as the Canaries, are a Spanish autonomous community and archipelago in the Atlantic Ocean, in Macaronesia. At their closest point to the African mainland, they are west of Mo ...
, South Africa * ''
Zimiris ''Zimiris'' is a genus of long-spinneret ground spiders that was first described by Eugène Louis Simon in 1882. it contains only two species, found in the Caribbean, Africa, South America, Asia, Germany, Mexico, and on Saint Helena: '' Z. di ...
'' Simon, 1882Circumtropical A 2020
phylogenetic analysis In biology Biology is the scientific study of life. It is a natural science with a broad scope but has several unifying themes that tie it together as a single, coherent field. For instance, all organisms are made up of cells that pro ...
involving 59 species of Prodidominae and 32 outgroup species did not recover Propidominae as
monophyletic In cladistics for a group of organisms, monophyly is the condition of being a clade—that is, a group of taxa composed only of a common ancestor (or more precisely an ancestral population) and all of its lineal descendants. Monophyletic ...
because '' Anagrina'' did not arise within the subfamily. The study re-established Molycriinae (including genera '' Cryptoerithus'', ''
Molycria ''Molycria'' is a genus of Australian Gnaphosidae, ground spiders that was first described by Eugène Simon, Eugène Louis Simon in 1887. Species it contains thirty-six species, found in the Northern Territory, Western Australia, South Australi ...
'', ''
Nomindra ''Nomindra'' is a genus of Australia, Australian Gnaphosidae, ground spiders that was first described by Norman I. Platnick & Barbara Baehr in 2006. Originally placed with the Prodidomidae, long-spinneret ground spiders, it was transferred to the ...
'', ''
Wesmaldra ''Wesmaldra'' is a genus of Australia, Australian Gnaphosidae, ground spiders that was first described by Norman I. Platnick & Barbara Baehr in 2006. Originally placed with the Prodidomidae, long-spinneret ground spiders, it was transferred to the ...
'', and ''
Wydundra ''Wydundra'' is a genus of Gnaphosidae, ground spiders that was first described by Norman I. Platnick & Barbara Baehr in 2006. Originally placed with the Prodidomidae, long-spinneret ground spiders, it was transferred to the ground spiders in 2018 ...
'') as a distinct subfamily in
Gnaphosidae Ground spiders comprise Gnaphosidae, the seventh largest spider family with over 2,000 described species in over 100 genera distributed worldwide. There are 105 species known to central Europe, and common genera include '' Gnaphosa'', '' Drassode ...
, a sister to Prodidominae.


References


Bibliography

* Platnick, N.I. & Penney, D. (2004)
A Revision of the Widespread Spider Genus ''Zimiris'' (Araneae, Prodidomidae)
''American Museum Novitates'' 3450. * Platnick, Norman I. & Baehr, Barbara C. (2006)
A revision of the Australasian ground spiders of the family Prodidomidae (Araneae, Gnaphosoidea)
''Bulletin of the American Museum of Natural History'' 298: 1-287. (with keys to subfamilies and genera, and picture) * Platnick, Norman I. (2008)

version 8.5. ''American Museum of Natural History''.


External links


Picture of ''Molycria flavipes''
Gnaphosidae Spider subfamilies {{Gnaphosidae-stub