''Proconsul'' is an
extinct
Extinction is the termination of an organism by the death of its Endling, last member. A taxon may become Functional extinction, functionally extinct before the death of its last member if it loses the capacity to Reproduction, reproduce and ...
genus
Genus (; : genera ) is a taxonomic rank above species and below family (taxonomy), family as used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In bino ...
of
primate
Primates is an order (biology), order of mammals, which is further divided into the Strepsirrhini, strepsirrhines, which include lemurs, galagos, and Lorisidae, lorisids; and the Haplorhini, haplorhines, which include Tarsiiformes, tarsiers a ...
s that existed from 21 to 17 million years ago during the
Miocene
The Miocene ( ) is the first epoch (geology), geological epoch of the Neogene Period and extends from about (Ma). The Miocene was named by Scottish geologist Charles Lyell; the name comes from the Greek words (', "less") and (', "new") and mea ...
epoch.
Fossil
A fossil (from Classical Latin , ) is any preserved remains, impression, or trace of any once-living thing from a past geological age. Examples include bones, shells, exoskeletons, stone imprints of animals or microbes, objects preserve ...
remains are present in
Eastern Africa
East Africa, also known as Eastern Africa or the East of Africa, is a region at the eastern edge of the Africa, African continent, distinguished by its unique geographical, historical, and cultural landscape. Defined in varying scopes, the regi ...
, including
Kenya
Kenya, officially the Republic of Kenya, is a country located in East Africa. With an estimated population of more than 52.4 million as of mid-2024, Kenya is the 27th-most-populous country in the world and the 7th most populous in Africa. ...
and
Uganda
Uganda, officially the Republic of Uganda, is a landlocked country in East Africa. It is bordered to the east by Kenya, to the north by South Sudan, to the west by the Democratic Republic of the Congo, to the south-west by Rwanda, and to the ...
. Four species have been classified to date:
''P. africanus'', ''P. gitongai'', ''
P. major'' and ''P. meswae''. The four species differ mainly in body size. Environmental reconstructions for the Early Miocene ''Proconsul'' sites are still tentative and range from forested environments to more open, arid grasslands.
The
gibbon
Gibbons () are apes in the family Hylobatidae (). The family historically contained one genus, but now is split into four extant genera and 20 species. Gibbons live in subtropical and tropical forests from eastern Bangladesh and Northeast Indi ...
and
great apes
The Hominidae (), whose members are known as the great apes or hominids (), are a taxonomic family of primates that includes eight extant species in four genera: '' Pongo'' (the Bornean, Sumatran and Tapanuli orangutan); '' Gorilla'' (the ...
, including humans, are held in evolutionary biology to share a common ancestral lineage, which may have included ''Proconsul''. Its name, meaning "before Consul" (Consul being a certain chimpanzee that, at the time of the genus's discovery, was on display in
London
London is the Capital city, capital and List of urban areas in the United Kingdom, largest city of both England and the United Kingdom, with a population of in . London metropolitan area, Its wider metropolitan area is the largest in Wester ...
), implies that it is ancestral to the
chimpanzee
The chimpanzee (; ''Pan troglodytes''), also simply known as the chimp, is a species of Hominidae, great ape native to the forests and savannahs of tropical Africa. It has four confirmed subspecies and a fifth proposed one. When its close rel ...
. It might also be ancestral to the rest of the
ape
Apes (collectively Hominoidea ) are a superfamily of Old World simians native to sub-Saharan Africa and Southeast Asia (though they were more widespread in Africa, most of Asia, and Europe in prehistory, and counting humans are found global ...
s.
Description
The genus had a mixture of
Old World monkey
Old World monkeys are primates in the family Cercopithecidae (). Twenty-four genera and 138 species are recognized, making it the largest primate family. Old World monkey genera include baboons (genus '' Papio''), red colobus (genus '' Piliocolob ...
and
ape
Apes (collectively Hominoidea ) are a superfamily of Old World simians native to sub-Saharan Africa and Southeast Asia (though they were more widespread in Africa, most of Asia, and Europe in prehistory, and counting humans are found global ...
characteristics, so its placement in the ape superfamily
Hominoidea
Apes (collectively Hominoidea ) are a Family (biology), superfamily of Old World simians native to sub-Saharan Africa and Southeast Asia (though they were more widespread in Africa, most of Asia, and Europe in prehistory, and counting humans ...
is tentative, with some scientists placing ''Proconsul'' outside it, before the split of the
apes
Apes (collectively Hominoidea ) are a superfamily of Old World simians native to sub-Saharan Africa and Southeast Asia (though they were more widespread in Africa, most of Asia, and Europe in prehistory, and counting humans are found glo ...
and
Old World monkeys
Old World monkeys are primates in the family (biology), family Cercopithecidae (). Twenty-four genus, genera and 138 species are recognized, making it the largest primate family. Old World monkey genera include baboons (genus ''Papio''), red colo ...
.
''Proconsul's'' monkey-like features include
pronograde posture, indicated by a long flexible back, curved metacarpals, and an above-branch
arboreal
Arboreal locomotion is the locomotion of animals in trees. In habitats in which trees are present, animals have evolved to move in them. Some animals may scale trees only occasionally (scansorial), but others are exclusively arboreal. The hab ...
quadrupedal positional repertoire. The primary feature linking ''Proconsul'' with extant apes is its lack of a tail; other "ape-like" features include its enhanced grasping capabilities, stabilized elbow joint and facial structure. ''Proconsul'' could not
hang effortlessly from tree branches like gibbons and other nonhuman apes do today.
Discovery and classification

The first specimen, a partial jaw discovered in 1909 by a gold prospector at
Koru
The is a spiral shape evoking a newly Circinate vernation, unfurling frond from a Cyathea dealbata, silver fern frond. It is an integral symbol in Māori people, Māori art, bone carving, carving and Tā moko, tattooing, where it symbolise ...
, near
Kisumu
Kisumu ( ) is the third-largest city in Kenya located in the Lake Victoria area in the former Nyanza Province. It is the second-largest city after Kampala in the Lake Victoria Basin. The city has a population of slightly over 600,000. The ...
in western Kenya, was also the oldest fossil hominoid known until recently, and the first fossil mammal ever found in sub-Saharan Africa. The name, ''Proconsul'', was devised by Arthur Hopwood in 1933 and means "before Consul" – the name of a famous captive chimpanzee in London.
At the time Consul was being used as a circus name for performing chimpanzees. The
Folies Bergère
150px, Stanisław Julian Ignacy Ostroróg">Walery, 1927
The Folies Bergère () is a cabaret music hall in Paris, France. Located at 32 Rue Richer in the 9th Arrondissement, the Folies Bergère was built as an opera house by the arc ...
of 1903 in
Paris
Paris () is the Capital city, capital and List of communes in France with over 20,000 inhabitants, largest city of France. With an estimated population of 2,048,472 residents in January 2025 in an area of more than , Paris is the List of ci ...
had a popular performing
chimpanzee
The chimpanzee (; ''Pan troglodytes''), also simply known as the chimp, is a species of Hominidae, great ape native to the forests and savannahs of tropical Africa. It has four confirmed subspecies and a fifth proposed one. When its close rel ...
named Consul, and so did the
Belle Vue Zoological Gardens in
Manchester
Manchester () is a city and the metropolitan borough of Greater Manchester, England. It had an estimated population of in . Greater Manchester is the third-most populous metropolitan area in the United Kingdom, with a population of 2.92&nbs ...
, England, in 1894. On the latter's death in that year
Ben Brierley wrote a commemorative poem wondering where the "
Missing Link" between chimpanzees and men was.

Hopwood in 1931 had discovered the fossils of three individuals while expeditioning with
Louis Leakey
Louis Seymour Bazett Leakey (7 August 1903 – 1 October 1972) was a Kenyan-British palaeoanthropologist and archaeologist whose work was important in demonstrating that humans evolved in Africa, particularly through discoveries made at Olduvai ...
in the vicinity of
Lake Victoria
Lake Victoria is one of the African Great Lakes. With a surface area of approximately , Lake Victoria is Africa's largest lake by area, the world's largest tropics, tropical lake, and the world's second-largest fresh water lake by surface are ...
. The Consul that he selected to use in the name was neither of the ones mentioned above, but another located in the
London Zoo
London Zoo, previously known as ZSL London Zoo or London Zoological Gardens and sometimes called Regent's Park Zoo, is the world's oldest scientific zoo. It was opened in London on 27 April 1828 and was originally intended to be used as a colle ...
. Consul is being used Linnaean-style to symbolize the
chimpanzee
The chimpanzee (; ''Pan troglodytes''), also simply known as the chimp, is a species of Hominidae, great ape native to the forests and savannahs of tropical Africa. It has four confirmed subspecies and a fifth proposed one. When its close rel ...
. ''Proconsul'' is therefore "ancestral to the Chimpanzee" in Hopwood's words. He also added ''africanus'' as the specific name.
Other fossils discovered later were initially classified as ''africanus'' and subsequently reclassified; that is, the total pool of fossils originally considered ''africanus'' was split and the fragments lumped with other finds to create a new species. For example,
Mary Leakey
Mary Douglas Leakey, Fellow of the British Academy, FBA (née Nicol, 6 February 1913 – 9 December 1996) was a British paleoanthropologist who discovered the first fossilised ''Proconsul (mammal), Proconsul'' skull, an extinct ape which is now ...
's famous find of 1948 began as ''africanus'' and was split from it to be lumped with Thomas Whitworth's finds of 1951 as ''heseloni'' by
Alan Walker
Alan Olav Walker (born 24 August 1997) is a Norwegian DJ and record producer. His songs "Faded (Alan Walker song), Faded", "Sing Me to Sleep", "Alone (Alan Walker song), Alone", "All Falls Down (Alan Walker song), All Falls Down" (with Noah Cy ...
in 1993. This process creates some confusion for the public, which is told that ''africanus'' became ''heseloni''. The finds from Koru and Songhor are still considered ''africanus''. Four species are still defined even though many fossils have jumped species.
The family of
Proconsulidae
Proconsulidae is an early family of primates that lived during the Miocene epoch in Kenya, and was restricted to Africa. Members of the family have a mixture of Old World monkey and ape characteristics, so the placement in the ape superfamily Hom ...
was first proposed by Louis Leakey in 1963, a decade after he and
Wilfrid Le Gros Clark had defined ''africanus'', ''nyanzae'' and ''major''. It was not immediately accepted but ultimately prevailed.
The history of
hominoid classification in the second half of the 20th century is sufficiently complex to warrant a few books itself. Most of the palaeoanthropologists have changed their minds at least once as new fossils have come to light and new observations have been made, and will probably continue to do so. The classifications found in the literature of one decade are not generally the same as those of another. For example, in 1987 Peter Andrews and
Lawrence Martin, established palaeontologists, took the point of view that ''Proconsul'' is not a hominoid, but is a sister taxon to it.
Reassigned species
The species ''Proconsul heseloni'' and ''P. nyanzae'' have been reclassified in the new genus ''
Ekembo''.
Notes
References
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{{Taxonbar, from=Q131791
Proconsulidae
Miocene primates of Africa
Fossil taxa described in 1933
Prehistoric primate genera