Porpidia Degelii
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''Porpidia'' is a
genus Genus (; : genera ) is a taxonomic rank above species and below family (taxonomy), family as used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In bino ...
of
crustose lichen Crustose lichens are lichens that form a crust which strongly adheres to the Substrate (biology), substrate (soil, rock, tree bark, etc.), making separation from the substrate impossible without destruction. The basic structure of crustose lichen ...
s in the family
Lecideaceae The Lecideaceae are a family (biology), family of lichen-forming fungi in the order Lecideales. It contains about 30 genus, genera and roughly 250 species. A major distinguishing characteristic of the family is the form of the ascomata, fruit ...
. ''Porpidia'' species primarily inhabit
siliceous rock Siliceous rocks are sedimentary rocks that have silica (SiO2) as the principal constituent. The most common siliceous rock is chert; other types include diatomite. They commonly form from silica-secreting organisms such as radiolarians, diatoms, o ...
s, pebbles, and stonework, with rare occurrences on bark, wood, and compacted soil. The
thallus Thallus (: thalli), from Latinized Greek (), meaning "a green shoot" or "twig", is the vegetative tissue of some organisms in diverse groups such as algae, fungi, some liverworts, lichens, and the Myxogastria. A thallus usually names the entir ...
, or body of the lichen, varies in appearance from thick and crusty to barely visible. It may form a continuous layer or develop cracks resulting in a segmented, structure. The colour of the thallus ranges from grey and white to orange.


Taxonomy

''Porpidia'' was
circumscribed In geometry, a circumscribed circle for a set of points is a circle passing through each of them. Such a circle is said to ''circumscribe'' the points or a polygon formed from them; such a polygon is said to be ''inscribed'' in the circle. * Circum ...
by the German lichenologist
Gustav Wilhelm Körber Gustav Wilhelm Körber (10 January 1817, Jelenia Góra, Hirschberg – 27 January 1885, Breslau) was a Silesian-German lichenologist and a professor at the University of Wrocław, University of Breslau. He specialized in the flora of Central Europ ...
in 1855, with '' Porpidia trullisata'' designated as the
type species In International_Code_of_Zoological_Nomenclature, zoological nomenclature, a type species (''species typica'') is the species name with which the name of a genus or subgenus is considered to be permanently taxonomically associated, i.e., the spe ...
.


Description

Genus ''Porpidia'' consists of
crustose lichen Crustose lichens are lichens that form a crust which strongly adheres to the Substrate (biology), substrate (soil, rock, tree bark, etc.), making separation from the substrate impossible without destruction. The basic structure of crustose lichen ...
s, meaning the
thallus Thallus (: thalli), from Latinized Greek (), meaning "a green shoot" or "twig", is the vegetative tissue of some organisms in diverse groups such as algae, fungi, some liverworts, lichens, and the Myxogastria. A thallus usually names the entir ...
(the main body of the lichen) is closely attached to the surface it grows on. The thallus can vary in appearance, from thick and scaly to almost invisible, and may be solid or cracked into small pieces called . Its colour typically ranges from grey to white, though some species may show orange hues. A , a thin, often black or orange border between neighbouring lichens, may be present but is not always visible. The lichen's inner structure, or
medulla Medulla (Latin for "marrow") or medullary may refer to: Science * Medulla oblongata, a part of the brain stem * Renal medulla, a part of the kidney * Adrenal medulla, a part of the adrenal gland * Medulla of ovary, a stroma in the center of the ...
, may or may not react with
iodine Iodine is a chemical element; it has symbol I and atomic number 53. The heaviest of the stable halogens, it exists at standard conditions as a semi-lustrous, non-metallic solid that melts to form a deep violet liquid at , and boils to a vi ...
(I+ or I–), and some species produce powdery
soredia Soredia are common reproduction, reproductive structures of lichens. Lichens asexual reproduction, reproduce asexually by employing simple fragmentation and production of soredia and isidia. Soredia are powdery propagules composed of fungus, fung ...
for reproduction, while others do not. One species of ''Porpidia'' even has
isidia An isidium (plural: isidia) is a tiny, wart- or finger-like outgrowth on the thallus surface of certain lichen species. It is one of two principal types of vegetative reproduction, vegetative reproductive structures in lichens, the other being ...
—small, finger-like structures that aid in reproduction. The lichen's
green algae The green algae (: green alga) are a group of chlorophyll-containing autotrophic eukaryotes consisting of the phylum Prasinodermophyta and its unnamed sister group that contains the Chlorophyta and Charophyta/ Streptophyta. The land plants ...
partner is usually from the genera ''
Trebouxia ''Trebouxia'' is a unicellular green alga. It is a photosynthetic organism that can exist in almost all habitats found in polar, tropical, and temperate regions.Erokhina, L. G., Shatilovich, A. V., Kaminskaya, O. P., & Gilichinskii, D. A. (2004 ...
'' or ''
Asterochloris ''Asterochloris'' is a genus of green algae in the family Trebouxiophyceae. It is a common in lichen, occurring in the thalli of more than 20 lichen genera worldwide. ''Asterochloris'' is distinguishable from the morphologically similar genus ...
''. The
apothecia An ascocarp, or ascoma (: ascomata), is the fruiting body ( sporocarp) of an ascomycete phylum fungus. It consists of very tightly interwoven hyphae and millions of embedded asci, each of which typically contains four to eight ascospores. As ...
(fruiting bodies) are common and can appear either scattered or in clusters. These round, dark brown to black are often relatively large, ranging from 0.5 to 3 mm in diameter, and may be coated with a frosty layer (a ). The apothecia have a distinct outer margin, which may fade as the lichen ages, and are seated directly on the surface of the thallus without a (a rim formed by the thallus itself). The rim surrounding the apothecia is usually dark blue-black or brown, and made of tightly packed fungal filaments. Under the surface, the reproductive layer (the
hymenium The hymenium is the tissue layer on the hymenophore of a fungal fruiting body where the cells develop into basidia or asci, which produce spores. In some species all of the cells of the hymenium develop into basidia or asci, while in oth ...
) reacts with iodine (I+) to turn blue, a useful trait for identification. The asci, the spore-producing sacs, each hold eight spores and have a distinctive shape called the "''Porpidia''-type". The are typically
ellipsoid An ellipsoid is a surface that can be obtained from a sphere by deforming it by means of directional Scaling (geometry), scalings, or more generally, of an affine transformation. An ellipsoid is a quadric surface;  that is, a Surface (mathemat ...
and measure 15–22 μm long, with a thick outer layer. In some species, other reproductive structures, known as
conidiomata Conidiomata (singular: Conidioma) are blister-like fruiting structures produced by a specific type of fungus called a coelomycete. They are formed as a means of dispersing asexual spores call conidia, which they accomplish by creating the blister- ...
, are present and often have a rough surface.


Species

,
Species Fungorum ''Index Fungorum'' is an international project to index all formal names (Binomial nomenclature, scientific names) in the fungus Kingdom (biology), kingdom. As of 2015, the project is based at the Royal Botanic Gardens, Kew, one of three partn ...
accepts 41 species of ''Porpidia'': *'' Porpidia albocaerulescens'' *'' Porpidia cinereoatra'' *'' Porpidia circumnigrata'' *'' Porpidia contraponenda'' *'' Porpidia crustulata'' *'' Porpidia degelii'' *'' Porpidia flavicunda'' *'' Porpidia flavocruenta'' - Austria, the British Isles, Scandinavia, Iceland, and North America (Alaska) *'' Porpidia grisea'' *'' Porpidia hydrophila'' *'' Porpidia hypostictica'' – China *'' Porpidia irrigua'' *'' Porpidia islandica'' - Scotland and Iceland *'' Porpidia kashmirensis'' – Pakistan *'' Porpidia littoralis'' – Australia *'' Porpidia lowiana'' *'' Porpidia macrocarpa'' *'' Porpidia melinodes'' *'' Porpidia nadvornikiana'' *'' Porpidia navarina'' – Chile *'' Porpidia ochrolemma'' *'' Porpidia pachythallina'' - British Isles *'' Porpidia platycarpoides'' *'' Porpidia rugosa'' *'' Porpidia seakensis''
Alaska Alaska ( ) is a non-contiguous U.S. state on the northwest extremity of North America. Part of the Western United States region, it is one of the two non-contiguous U.S. states, alongside Hawaii. Alaska is also considered to be the north ...
*'' Porpidia shangrila'' – China *'' Porpidia soredizodes'' *'' Porpidia speirea'' *'' Porpidia squamosa'' – China *'' Porpidia stephanodes'' *'' Porpidia striata'' - British Isles *'' Porpidia submelinodes'' – Antarctica *'' Porpidia subsimplex'' *'' Porpidia superba'' *'' Porpidia thomsonii'' *'' Porpidia trullisata'' *'' Porpidia tuberculosa'' *'' Porpidia ulleungdoensis''
Ulleungdo Ulleungdo (), also spelled Ulreungdo, is a South Korean island east of the Korean Peninsula in the Sea of Japan. It was formerly known as Dagelet Island or Argonaut Island in Europe. Volcanic in origin, the rocky steep-sided island is the top o ...
, South Korea *'' Porpidia umbonifera'' *'' Porpidia vulcanoides'' *'' Porpidia zeoroides''


References

{{Taxonbar, from=Q7230357 Lecideales Lecideales genera Lichen genera Taxa named by Gustav Wilhelm Körber Taxa described in 1855