Plant Associations
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A plant community is a collection or
association Association may refer to: *Club (organization), an association of two or more people united by a common interest or goal *Trade association, an organization founded and funded by businesses that operate in a specific industry *Voluntary associatio ...
of
plant species Flora (: floras or florae) is all the plant life present in a particular region or time, generally the naturally occurring ( indigenous) native plants. The corresponding term for animals is ''fauna'', and for fungi, it is '' funga''. Sometimes ...
within a designated geographical unit, which forms a relatively uniform patch, distinguishable from neighboring patches of different
vegetation type Vegetation classification is the process of classifying and mapping the vegetation over an area of the Earth's surface. Vegetation classification is often performed by state based agencies as part of land use, resource and environmental management ...
s. The components of each plant community are influenced by
soil type A soil type is a taxonomic unit in soil science. All soils that share a certain set of well-defined properties form a distinctive soil type. Soil type is a technical term of soil classification, the science that deals with the systematic categ ...
,
topography Topography is the study of the forms and features of land surfaces. The topography of an area may refer to the landforms and features themselves, or a description or depiction in maps. Topography is a field of geoscience and planetary sci ...
,
climate Climate is the long-term weather pattern in a region, typically averaged over 30 years. More rigorously, it is the mean and variability of meteorological variables over a time spanning from months to millions of years. Some of the meteoro ...
and human disturbance. In many cases there are several soil types present within a given plant community. This is because the soil type within an area is influenced by two factors, the rate at which water infiltrates or exits (via
evapotranspiration Evapotranspiration (ET) refers to the combined processes which move water from the Earth's surface (open water and ice surfaces, bare soil and vegetation) into the Atmosphere of Earth, atmosphere. It covers both water evaporation (movement of w ...
) the soil, as well as the rate at which organic matter (any carbon-based compound within the environment, such as decaying plant matter) enters or decays from the soil. Plant communities are studied substantially by ecologists, due to providing information on the effects of dispersal, tolerance to environmental conditions, and response to disturbance of a variety of plant species, information valuable to the comprehension of various plant community dynamics. Plant communities having a stable composition after a relatively long period free of disturbance represent the potential natural vegetation, or “climax” plant community and are often called "Plant Associations." A Plant Association can be conceptual, and gives an indication of the direction of succession. The USDA Forest Service collects field data, performs spatial statistics, and maps potential plant associations to assist in planting and restoration efforts. The US Bureau of Land Management also establishes plant communities using "Ecological Sites," which are roughly equivalent to plant associations.


Definition

A plant community can be described
floristically Floristics is the study of plants of geographical regions. It is a branch of phytogeography, which technically makes it a branch of botany, geography, and a subbranch of biogeography. Harvard University has a history of research with early contr ...
(the species of flowers or flora the plant community contains) and/or phytophysiognomically (the physical structure or appearance of the plant community). For example, a
forest A forest is an ecosystem characterized by a dense ecological community, community of trees. Hundreds of definitions of forest are used throughout the world, incorporating factors such as tree density, tree height, land use, legal standing, ...
(a community of trees) includes the
overstory In biology, the canopy is the aboveground portion of a plant cropping or crop, formed by the collection of individual plant crowns. In forest ecology, the canopy is the upper layer or habitat zone, formed by mature tree crowns and includ ...
, or upper tree layer of the
canopy Canopy may refer to: Plants * Canopy (biology), aboveground portion of plant community or crop (including forests) * Canopy (grape), aboveground portion of grapes Religion and ceremonies * Baldachin or canopy of state, typically placed over an a ...
, as well as the
understory In forestry and ecology, understory (American English), or understorey (English in the Commonwealth of Nations, Commonwealth English), also known as underbrush or undergrowth, includes plant life growing beneath the Canopy (biology), forest ca ...
, a layer consisting of trees and shrubs located beneath the canopy but above the forest floor. The understory can be further subdivided into the
shrub layer In ecology, stratification refers to the vertical layering of a habitat; the arrangement of vegetation in layers. It classifies the layers (sing. ''stratum'', pl. ''strata'') of vegetation largely according to the different heights to which their p ...
, composed of vegetation and trees between a height of approximately one to five meters, the
herbaceous layer In ecology, stratification refers to the vertical layering of a habitat; the arrangement of vegetation in layers. It classifies the layers (sing. ''stratum'', pl. ''strata'') of vegetation largely according to the different heights to which their p ...
, composed of vascular plants at a height of one meter or less, and sometimes also the moss layer, a layer of non-vascular bryophytes typically present at ground level (approximately 0.15 meters in height or less). In some cases of complex forests there is also a well-defined lower tree layer. A plant community is similar in concept to a
vegetation type Vegetation classification is the process of classifying and mapping the vegetation over an area of the Earth's surface. Vegetation classification is often performed by state based agencies as part of land use, resource and environmental management ...
, with the former having more of an emphasis on the ecological association of species within it, and the latter on overall appearance by which it is readily recognized by a layperson. A plant community can be rare even if none of the major species defining it are rare. This is because it is the association of species and relationship to their environment that may be rare.Introduction to California Plant Life,
Robert Ornduff Robert Ornduff (1932–2000) was an American botanist. He was Director of the University and Jepson Herbaria, Director of the University of California Botanical Garden, Executive Director of the Miller Institute for Basic Research in Science, an ...
,
Phyllis M. Faber Phyllis M. Faber (1928–2023) was an environmental activist and agriculture botanist from Marin County, California. Early life and education Originally from New York City, she grew up hearing about the West Coast because of her father, who w ...
, Todd Keeler-Wolf, California Natural History Guides No. 69, University of California Press, Ltd., 2003,
An example is the sycamore alluvial woodland in California dominated by the California sycamore ''
Platanus racemosa ''Platanus racemosa'' is a species of plane tree known by several common names, including California sycamore, western sycamore, California plane tree, and in North American Spanish . ''Platanus racemosa'' is native to California and Baja Califo ...
''. The community is rare, being localized to a small area of California and existing nowhere else, yet the California sycamore is not a rare tree in California.


Examples

An example is a
grassland A grassland is an area where the vegetation is dominance (ecology), dominated by grasses (Poaceae). However, sedge (Cyperaceae) and rush (Juncaceae) can also be found along with variable proportions of legumes such as clover, and other Herbaceo ...
on the northern
Caucasus The Caucasus () or Caucasia (), is a region spanning Eastern Europe and Western Asia. It is situated between the Black Sea and the Caspian Sea, comprising parts of Southern Russia, Georgia, Armenia, and Azerbaijan. The Caucasus Mountains, i ...
steppes In physical geography, a steppe () is an ecoregion characterized by grassland plains without closed forests except near rivers and lakes. Steppe biomes may include: * the montane grasslands and shrublands biome * the tropical and subtropical gr ...
, where common grass species found are ''
Festuca sulcata ''Festuca rupicola'', the furrowed fescue, is a species of cool-season grass in the family Poaceae Poaceae ( ), also called Gramineae ( ), is a large and nearly ubiquitous family of monocotyledonous flowering plants commonly known as gra ...
'' and ''
Poa bulbosa ''Poa bulbosa'' is a species of grass known by the common names bulbous bluegrass or bulbous meadow-grass. It is native to Eurasia and North Africa, but it is present practically worldwide as an introduced species. It is widespread in the United ...
''. The most common species defining this grassland
phytocoenosis Phytosociology, also known as phytocoenology or simply plant sociology, is the study of groups of species of plant that are usually found together. Phytosociology aims to empirically describe the vegetative environment of a given territory. A spec ...
is ''
Carex shreberi ''Carex'' is a vast genus of over 2,000 species of grass-like plants in the family Cyperaceae, commonly known as sedges (or seg, in older books). Other members of the family Cyperaceae are also called sedges, however those of genus ''Carex'' may ...
''. Other representative
forb A forb or phorb is a herbaceous flowering plant that is not a graminoid (grass, sedge, or rush). The term is used in botany and in vegetation ecology especially in relation to grasslands and understory. Typically, these are eudicots without woo ...
s occurring in these steppe grasslands are ''
Artemisia austriaca ''Artemisia austriaca'' is a species of plant belonging to the family Asteraceae. Its native range is Europe to Western Siberia, and Iran Iran, officially the Islamic Republic of Iran (IRI) and also known as Persia, is a country in We ...
'' and ''
Polygonum aviculare ''Polygonum aviculare'' or common knotgrass is a plant related to buckwheat and dock. It is also called prostrate knotweed, birdweed, pigweed and lowgrass. It is an annual found in fields and wasteland, with white flowers from June to October. I ...
''. Other examples of different plant communities include the forests located on the granite peaks of the Huangshan Mountains in Eastern China. The deciduous broad-leaved forest, present from a height of 1,100 metres, is populated by trees such as '' Pinus hwangshanesis'', also known as the Huangshan pine. The Huangshan mountain also possesses an evergreen broad-leaved forest community, home to a variety of shrubs and small trees. Some examples of species present in the evergreen broad-leaved forest community include '' Castanopsis eyrei'', '' Eurya nitidia,
Rhododendron ovatum ''Rhododendron ovatum'' is an elepidote rhododendron species native to China and Taiwan. It is the type species In International_Code_of_Zoological_Nomenclature, zoological nomenclature, a type species (''species typica'') is the species nam ...
,
Pinus massoniana ''Pinus massoniana'' (English: Masson's pine, Chinese red pine, horsetail pine; Chinese: 馬尾松) is a species of pine, native to Taiwan, a wide area of central and southern China, and northern Vietnam. Description It is an evergreen tree ...
,'' as well as ''
Loropetalum chinense ''Loropetalum chinense'', commonly known as loropetalum,"Floridata Plan ...
''. An example of a three tiered plant community is in central Westland in the South Island, New Zealand. These forests are the most extensive continuous reaches of
podocarp Podocarpaceae is a large family of mainly southern hemisphere conifers, known in English as podocarps, comprising about 156 species of evergreen trees and shrubs.James E. Eckenwalder. 2009. ''Conifers of the World''. Portland, Oregon: Timber Pres ...
/broadleaf forests in that country. The canopy includes ''
Prumnopitys ferruginea ''Pectinopitys ferruginea'', commonly known as miro or toromiro, is an evergreen coniferous tree which is endemic to New Zealand. Description It grows up to high, with a trunk up to 1.3 m diameter. The leaves are linear to sickle-shaped, 15 ...
'',
rimu ''Dacrydium cupressinum'', commonly known as rimu, is a species of tree in the family Podocarpaceae. It is a dioecious evergreen conifer, reaching heights of up to , and can have a stout trunk (botany), trunk up to in diameter. It is endemis ...
and mountain totara. The mid-story includes tree ferns such as ''
Cyathea smithii ''Alsophila smithii'', synonym ''Cyathea smithii'', commonly known as the soft tree fern or kātote, is a species of tree fern from New Zealand. Distribution and ecology The species' natural distribution covers all major islands of New Zealand: ...
'' and ''
Dicksonia squarrosa ''Dicksonia squarrosa'', the New Zealand tree fern, whekī or rough tree fern, is a common tree fern endemic to New Zealand. It has a slender black trunk that is usually surrounded by many dead brown fronds. Description This species has a fas ...
'', whilst the lowest tier and
epiphytic An epiphyte is a plant or plant-like organism that grows on the surface of another plant and derives its moisture and nutrients from the air, rain, water (in marine environments) or from debris accumulating around it. The plants on which epiphyt ...
associates include '' Asplenium polyodon'', ''
Tmesipteris tannensis ''Tmesipteris'', the hanging fork ferns, is a genus of ferns, one of two genera in the family Psilotaceae, order Psilotales (the other being ''Psilotum''). ''Tmesipteris'' is restricted to certain lands in the Southern Pacific, notably Australia, ...
'', '' Astelia solandri'' and ''
Lomaria discolor ''Lomaria discolor'', synonym ''Blechnum discolor'', commonly called crown fern (Māori: piupiu), is a species of fern in the family Blechnaceae. This species is endemic to New Zealand New Zealand () is an island country in the southw ...
''.C. Michael Hogan. 2009
''Crown Fern: Blechnum discolor'', Globaltwitcher.com, ed. N. Stromberg
/ref>


See also

*
Community (ecology) In ecology, a community is a group or association (ecology), association of Population ecology, populations of two or more different species occupying the same geographical area at the same time, also known as a biocoenosis, biotic community, ...
* Size-asymmetric competition *
Ecosystem An ecosystem (or ecological system) is a system formed by Organism, organisms in interaction with their Biophysical environment, environment. The Biotic material, biotic and abiotic components are linked together through nutrient cycles and en ...
*
Habitat In ecology, habitat refers to the array of resources, biotic factors that are present in an area, such as to support the survival and reproduction of a particular species. A species' habitat can be seen as the physical manifestation of its ...
*
Phytosociology Phytosociology, also known as phytocoenology or simply plant sociology, is the study of groups of species of plant that are usually found together. Phytosociology aims to Empirical evidence, empirically describe the vegetative environment of a giv ...
*
Stand level modelling A forest stand is a contiguous Plant community, community of trees sufficiently uniform in composition, structure, Forest inventory#Timber metrics, age, size, class, distribution, spatial arrangement, condition, or location on a Site quality (for ...
*
Vegetation classification Vegetation classification is the process of classifying and mapping the vegetation over an area of the Earth's surface. Vegetation classification is often performed by state based agencies as part of land use, resource management, resource and envi ...


References

{{Authority control Community ecology Plant ecology Biogeography Habitats Ecology terminology