Pinus Sabineana
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''Pinus sabiniana'' (sometimes spelled ''P. sabineana'') is a
pine A pine is any conifer tree or shrub in the genus ''Pinus'' () of the family Pinaceae. ''Pinus'' is the sole genus in the subfamily Pinoideae. ''World Flora Online'' accepts 134 species-rank taxa (119 species and 15 nothospecies) of pines as cu ...
endemic Endemism is the state of a species being found only in a single defined geographic location, such as an island, state, nation, country or other defined zone; organisms that are indigenous to a place are not endemic to it if they are also foun ...
to
California California () is a U.S. state, state in the Western United States that lies on the West Coast of the United States, Pacific Coast. It borders Oregon to the north, Nevada and Arizona to the east, and shares Mexico–United States border, an ...
in the United States. Its
vernacular name Vernacular is the ordinary, informal, spoken form of language, particularly when perceived as having lower social status or less prestige than standard language, which is more codified, institutionally promoted, literary, or formal. More n ...
s include towani pine, foothill pine, gray pine, ghost pine, and bull pine. The name digger pine was historically used but includes a racial slur.


Description

''Pinus sabiniana'' trees typically grow to , but can reach . The pine needles are in fascicles (bundles) of three, distinctively pale gray-green, sparse and drooping, and grow to in length. The seed
cones In geometry, a cone is a three-dimensional figure that tapers smoothly from a flat base (typically a circle) to a point not contained in the base, called the ''apex'' or '' vertex''. A cone is formed by a set of line segments, half-lines, ...
are large and heavy, in length and almost as wide as they are long. When fresh, they weigh from , rarely over . The male cones grow at the base of shoots on the lower branches. File:Pinus sabineana 00061.JPG, Bark File:Pinus sabiniana (Gray Pine) - foliage (30485878313).jpg, Foliage File:Pinus sabiniana pollen cones Pinnacles, California.jpg, Pollen cones File:J20161101-0079—Gray pine cone, pine nuts, and resin—RPBG (30547385050).jpg, Cone, seeds, and resin


Taxonomy


Common name

The name digger pine supposedly came from the observation that the
Paiute Paiute (; also Piute) refers to three non-contiguous groups of Indigenous peoples of the Great Basin. Although their languages are related within the Numic group of Uto-Aztecan languages, these three languages do not form a single subgroup and th ...
foraged for its seeds by digging around the base of the tree. It is more likely that the term was first applied to the people; "Digger Indians" was in common use in California literature from the 1800s. The historically more common name ''digger pine'' is still in widespread use. The
Jepson Manual ''The Jepson Manual'' is a flora of the vascular plants that are either native to or naturalized in California. Botanists often refer to the book simply as ''Jepson''. It is produced by the University and Jepson Herbaria, of the University of Cal ...
advises avoiding this name as the authors believe "digger" is pejorative in origin. The tree is also sometimes thought of as a
pinyon pine The pinyon or piñon pine group grows in southwestern North America, especially in New Mexico, Colorado, Arizona, and Utah, with the single-leaf pinyon pine just reaching into southern Idaho. The trees yield edible Pine nut, nuts, which are a sta ...
, though it does not belong to that group.


Botanical name

The scientific
botanical name A botanical name is a formal scientific name conforming to the ''International Code of Nomenclature for algae, fungi, and plants'' (ICN) and, if it concerns a plant cultigen, the additional cultivar or cultivar group, Group epithets must conform t ...
with the standard spelling ''sabiniana'' commemorates
Joseph Sabine Joseph Sabine FRS ( ; 6 June 1770 – 24 January 1837) was an English lawyer, naturalist and writer on horticulture. Life and work Sabine was born into a prominent Anglo-Irish family in Tewin, Hertfordshire, the eldest son of Joseph Sabine. ...
, secretary of the
Horticultural Society of London The Royal Horticultural Society (RHS), founded in 1804 as the Horticultural Society of London, is the UK's leading gardening charity. The RHS promotes horticulture through its five gardens at Wisley (Surrey), Hyde Hall (Essex), Harlow Carr (Nor ...
. Some botanists proposed a new spelling ''sabineana'', because they were confused with Latin grammar. The proposal has not been accepted by the relevant authorities (i.e.
United States Department of Agriculture The United States Department of Agriculture (USDA) is an executive department of the United States federal government that aims to meet the needs of commercial farming and livestock food production, promotes agricultural trade and producti ...
, The Jepson Manual or
Germplasm Resources Information Network Germplasm Resources Information Network or GRIN is an online USDA National Genetic Resources Program software project to comprehensively manage the computer database for the holdings of all plant germplasm collected by the National Plant Germpla ...
(GRIN). The GRIN notes that the spelling ''sabiniana'' agrees with a provision in the Vienna Code of the
International Code of Botanical Nomenclature The ''International Code of Nomenclature for algae, fungi, and plants'' (ICN or ICNafp) is the set of rules and recommendations dealing with the formal botanical names that are given to plants, fungi and a few other groups of organisms, all tho ...
, the governing body of botanical nomenclature. In that code, recommendation 60.2C states that personal names can be Latinized in species epithets: 'Sabine' is Latinised to ''sabinius'', with the addition of the suffix "-anus" (pertaining to) the word becomes ''sabiniana'' (In Latin, trees are feminine, irrespective if the word ends with a masculine suffix, i.e. ''pinus'').International Code of Botanical Nomenclature. 2006
Recommendation 60C.2
. Accessed online: 1 October 2010.
The GRIN database notes that Sabine's last name is not correctable and therefore ''Pinus sabiniana'' is the proper name for the species.


Distribution and habitat

''Pinus sabiniana'' grows at elevations between sea level and and is common in the northern and interior portions of the
California Floristic Province The California Floristic Province (CFP) is a floristic province with a Mediterranean-type climate located on the Pacific Coast of North America with a distinctive flora similar to other regions with a winter rainfall and summer drought climat ...
. It is found throughout the
Sierra Nevada The Sierra Nevada ( ) is a mountain range in the Western United States, between the Central Valley of California and the Great Basin. The vast majority of the range lies in the state of California, although the Carson Range spur lies primari ...
and
Coast Ranges The Pacific Coast Ranges (officially gazetted as the Pacific Mountain System in the United States; ; ) are the series of mountain ranges that stretch along the West Coast of North America from Alaska south to Northern and Central Mexico. Althoug ...
foothills that ring the Central, San Joaquin and interior valleys; the
Transverse Transverse may refer to: *Transverse engine, an engine in which the crankshaft is oriented side-to-side relative to the wheels of the vehicle *Transverse flute, a flute that is held horizontally * Transverse force (or ''Euler force''), the tangen ...
and
Peninsular Ranges The Peninsular Ranges (also called the Lower California province) are a group of mountain ranges that stretch from Southern California to the southern tip of the Baja California peninsula; they are part of the North American Pacific Coast Range ...
; and
Mojave Desert The Mojave Desert (; ; ) is a desert in the rain shadow of the southern Sierra Nevada mountains and Transverse Ranges in the Southwestern United States. Named for the Indigenous peoples of the Americas, indigenous Mohave people, it is located pr ...
sky island Sky islands are isolated mountains surrounded by radically different lowland environments. The term originally referred to those found on the Mexican Plateau and has extended to similarly isolated high-elevation forests. The isolation has s ...
s. Multiple specimens have also been found in
Southern Oregon Southern Oregon is a region of the U.S. state of Oregon south of Lane County and generally west of the Cascade Range The Cascade Range or Cascades is a major mountain range of western North America, extending from southern British Columbia thr ...
as well. It is adapted to long, hot, dry summers and is found in areas with an unusually wide range of precipitation: from an average of per year at the edge of the Mojave to in parts of the Sierra Nevada. It prefers rocky, well drained soil, but also grows in serpentine soil and heavy, poorly drained clay soils. It commonly occurs in association with ''
Quercus douglasii ''Quercus douglasii'', known as blue oak, is a species of oak endemic to California, common in the Coast Ranges and the foothills of the Sierra Nevada. It is California's most drought-tolerant deciduous oak, and is a dominant species in the b ...
'', and "Oak/Foothill Pine vegetation" (also known as "Oak/Gray Pine vegetation") is used as a description of a type of
habitat In ecology, habitat refers to the array of resources, biotic factors that are present in an area, such as to support the survival and reproduction of a particular species. A species' habitat can be seen as the physical manifestation of its ...
characteristic within the
California chaparral and woodlands The California chaparral and woodlands is a terrestrial ecoregion of southwestern Oregon, northern, central, and southern California (United States) and northwestern Baja California (Mexico), located on the west coast of North America. It is a ...
ecoregion An ecoregion (ecological region) is an ecological and geographic area that exists on multiple different levels, defined by type, quality, and quantity of environmental resources. Ecoregions cover relatively large areas of land or water, and c ...
in California, providing a sparse
overstory In biology, the canopy is the aboveground portion of a plant cropping or crop, formed by the collection of individual plant crowns. In forest ecology, the canopy is the upper layer or habitat zone, formed by mature tree crowns and includ ...
above a canopy of the
oak woodland An oak woodland is a plant community with a tree canopy dominated by oaks (''Quercus spp.''). In terms of canopy closure, oak woodlands are intermediate between oak savanna, which is more open, and oak forest, which is more closed. Although the c ...
.


Ecology

''Pinus sabiniana'' needles are a food of the
caterpillar Caterpillars ( ) are the larval stage of members of the order Lepidoptera (the insect order comprising butterflies and moths). As with most common names, the application of the word is arbitrary, since the larvae of sawflies (suborder ...
s of the Gelechiid moth '' Chionodes sabinianus''. Fossil evidence suggests that it has only recently become adapted to the Mediterranean climate as its closest relatives are part of the Madrean pine-oak woodlands found at higher elevations in the southwest US and Mexico. Animals help spread the seeds, including birds such as the
scrub jay The passerine birds of the genus ''Aphelocoma'' include the scrub jays and their relatives. They are New World jays found in Mexico, western Central America and the western United States, with an outlying population in Florida. This genus belon ...
and
acorn woodpecker The acorn woodpecker (''Melanerpes formicivorus'') is a medium-sized woodpecker with a length of around , and an average weight of . It is found across Central America, as well as North into the western United States and South into parts of Colo ...
.


Uses

Some Native American groups relied heavily on sweet pine nuts for food and are thought to have contributed to the current distribution pattern, including the large gap in distribution in
Tulare County Tulare County ( ) is a county located in the U.S. state of California. As of the 2020 census, the population was 473,117. The county seat is Visalia. The county is named for Tulare Lake, once the largest freshwater lake west of the Great Lake ...
. Native Americans also consumed the roots. Protein and fat nutritional value of the seed are similar to ''
Pinus pinea The Italian stone pine, botanical name ''Pinus pinea'', also known as the Mediterranean stone pine is a tree from the pine family (''Pinaceae''). The tree is native to the Mediterranean region, occurring in Southern Europe and the Levant. The ...
'' seeds and figured in the local indigenous diet. Wood uses historically were determined by its particular characteristics, e.g., 0.43 mean
specific gravity Relative density, also called specific gravity, is a dimensionless quantity defined as the ratio of the density (mass of a unit volume) of a substance to the density of a given reference material. Specific gravity for solids and liquids is nea ...
nearly equal to Douglas-fir (
Pseudotsuga menziesii The Douglas fir (''Pseudotsuga menziesii'') is an evergreen conifer species in the pine family, Pinaceae. It is the tallest tree in the Pinaceae family. It is native plant, native to western North America and is also known as Douglas-fir, Dougl ...
); strength properties similar to
ponderosa pine ''Pinus ponderosa'', commonly known as the ponderosa pine, bull pine, blackjack pine, western yellow-pine, or filipinus pine, is a very large pine tree species of variable habitat native to mountainous regions of western North America. It is t ...
; Kraft pulps high in
bursting Bursting, or burst firing, is an extremely diverse general phenomenon of the activation patterns of neurons in the central nervous system where periods of rapid action potential spiking are followed by quiescent periods much longer than typical in ...
with
tensile strength Ultimate tensile strength (also called UTS, tensile strength, TS, ultimate strength or F_\text in notation) is the maximum stress that a material can withstand while being stretched or pulled before breaking. In brittle materials, the ultimate ...
comparable to some northern
conifer Conifers () are a group of conifer cone, cone-bearing Spermatophyte, seed plants, a subset of gymnosperms. Scientifically, they make up the phylum, division Pinophyta (), also known as Coniferophyta () or Coniferae. The division contains a sin ...
pulps; and foothill stands loggable in winter, when higher-altitude species were inaccessible. However, the high amounts of
resin A resin is a solid or highly viscous liquid that can be converted into a polymer. Resins may be biological or synthetic in origin, but are typically harvested from plants. Resins are mixtures of organic compounds, predominantly terpenes. Commo ...
and compression wood, the often crooked form, heavy weight, and low stand density, made it expensive otherwise to log, transport and process. Commercial value decreased by the 1960s, to limited use for
railroad tie A railroad tie, crosstie (American English), railway tie (Canadian English) or railway sleeper ( Australian and British English) is a rectangular support for the rails in railroad tracks. Generally laid perpendicular to the rails, ties trans ...
s, box "shook",
pallet A pallet (also called a skid) is a flat transport structure, which supports goods in a stable fashion while being lifted by a forklift, a pallet jack, a Loader (equipment), front loader, a Jack (mechanical), jacking device, or an erect cra ...
stock, and
chips ''CHiPs'' is an American crime drama television series created by Rick Rosner and originally aired on NBC from September 15, 1977, to May 1, 1983. After the final first-run telecast on NBC in May 1983, the series went into reruns on Sundays fr ...
. It may still offer potential as windbreak
shelterbelt A windbreak (shelterbelt) is a planting usually made up of one or more rows of trees or shrubs planted in such a manner as to provide shelter from the wind and to protect soil from erosion. They are commonly planted in hedgerows around the edg ...
plantings. The main turpentine constituent,
heptane Heptane or ''n''-heptane is the straight-chain alkane with the chemical formula H3C(CH2)5CH3 or C7H16. When used as a test fuel component in anti-knock test engines, a 100% heptane fuel is the zero point of the octane rating scale (the 100 poi ...
, an alkane hydrocarbon, at about 3 percent of needle and twig oil, is unusual in botany; the only other source in nature perhaps being the ''
Pittosporum resiniferum ''Pittosporum'' ( or The first pronunciation is that expected for Anglo-Latin; the second is common in nurseries. ''Sunset Western Garden Book,'' 1995:606–607) is a genus of about 250 species of flowering plants in the family Pittosporaceae. P ...
'', known as "petroleum nut" or kerosene tree.


Notes


References

* * A. Farjon (2005). ''Pines: Drawings and descriptions of the genus Pinus''. Brill. * * Discovery Channel (2010), ''
MythBusters ''MythBusters'' is a science entertainment television series created by Peter Rees (producer), Peter Rees and produced by Beyond International in Australia. The series premiered on the Discovery Channel on January 23, 2003. It was broadcast in ...
''
Episode 138
* *


Further reading

*


External links

* * *
USDA PLANTS Treatment for ''Pinus sabiniana'' (California foothill pine)
* {{Authority control sabiniana Endemic flora of California Trees of Northern America Flora of the Sierra Nevada (United States) Flora of the California desert regions Natural history of the California chaparral and woodlands Garden plants of North America Ornamental trees Drought-tolerant trees Butterfly food plants Edible nuts and seeds