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Callicostaceae is a family of
pleurocarpous The Bryopsida constitute the largest class of mosses, containing 95% of all moss species. It consists of approximately 11,500 species, common throughout the whole world. The group is distinguished by having spore capsules with teeth that are '' ...
mosses in
Hookeriales Hookeriales is an order of Bryophyta or leafy mosses. Named for William Jackson Hooker Sir William Jackson Hooker (6 July 178512 August 1865) was an English botany, botanist and botanical illustrator, who became the first director of Kew ...
. It includes 21 genera. It is primarily tropical and
epiphytic An epiphyte is a plant or plant-like organism that grows on the surface of another plant and derives its moisture and nutrients from the air, rain, water (in marine environments) or from debris accumulating around it. The plants on which epiphyt ...
or
epiphyllous Foliicolous refers to the growth habit of certain lichens, algae, fungi, liverworts, and other bryophytes that prefer to grow on the leaves of vascular plants. ''Foliicolous'' simply means 'growing upon leaves' whilst epiphyllous derives from the ...
.


Description

Members of the family are characterized by a double
costa Costa may refer to: Biology * Rib (Latin: ''costa''), in vertebrate anatomy * Costa (botany), the central strand of a plant leaf or thallus * Costa (coral), a stony rib, part of the skeleton of a coral * Costa (entomology), the leading edge of th ...
, sometimes with hyalodermis, and no
alar Daminozide, also known as aminozide, Alar, Kylar, SADH, B-995, B-nine, and DMASA, is an organic compound which acts as a plant growth regulator. It was produced in the U.S. by the Uniroyal Chemical Company, Uniroyal Chemical Company, Inc., (no ...
differentiation. Leaves are usually complanate and may by asymmetrical along the dorsal/ventral axis. Leaf margins are dentate to serrate, bordered or not, and often reduced. Laminal cells are isodiametric to linear and smooth to papillose. The sporophyte seta is long, but many other characters vary genus to genus.


Nomenclatural history

The name Pilotrichaceae has long been, and continues to be, used for this group. However the type of the genus ''Pilotrichum'' is ''Leptodon smithii'', which is a member of
Neckeraceae Neckeraceae is a moss family in the order Hypnales. There are about 200 species native to temperate and tropical regions. Most grow on rocks, or other plants. Description Members of the family are usually large and glossy plants with creeping st ...
, and therefore Pilotrichaceae is not available under the ICNafp as a name for this group. The genus ''Callicosta'' was introduced for the ''Pilotrichum'' species belonging to this group, and was made the type genus.


Classification

As reported b
Tropicos
an
World Flora Online
*'' Actinodontium'' *'' Amblytropis'' *'' Brymela'' *'' Callicosta'' *'' Callicostella'' *'' Callicostellopsis'' *'' Cyclodictyon'' *'' Diploneuron'' *'' Helicoblepharum'' *'' Hemiragis'' *'' Hypnella'' *'' Lepidopilidium'' *'' Lepidopilum'' *'' Neohypnella'' *'' Philophyllum'' *'' Pilotrichidium'' *''
Stenodesmus Stenodesmus may refer to: * Stenodesmus (millipede), ''Stenodesmus'' (millipede), a genus of millipedes in the family Xystodesmidae * Stenodesmus (plant), ''Stenodesmus'' (plant), a genus of mosses in the family Daltoniaceae {{Genus disambiguation ...
'' *'' Stenodictyon'' *'' Tachypodium'' *'' Thamniopsis'' *'' Trachyxiphium''


References

Hookeriales Moss families {{bryidae-stub