Panthera Onca Augusta
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''Panthera onca augusta'' is an
extinct Extinction is the termination of an organism by the death of its Endling, last member. A taxon may become Functional extinction, functionally extinct before the death of its last member if it loses the capacity to Reproduction, reproduce and ...
subspecies In Taxonomy (biology), biological classification, subspecies (: subspecies) is a rank below species, used for populations that live in different areas and vary in size, shape, or other physical characteristics (Morphology (biology), morpholog ...
of the
jaguar The jaguar (''Panthera onca'') is a large felidae, cat species and the only extant taxon, living member of the genus ''Panthera'' that is native to the Americas. With a body length of up to and a weight of up to , it is the biggest cat spe ...
that was endemic to
North America North America is a continent in the Northern Hemisphere, Northern and Western Hemisphere, Western hemispheres. North America is bordered to the north by the Arctic Ocean, to the east by the Atlantic Ocean, to the southeast by South Ameri ...
from
Middle Middle or The Middle may refer to: * Centre (geometry), the point equally distant from the outer limits. Places * Middle (sheading), a subdivision of the Isle of Man * Middle Bay (disambiguation) * Middle Brook (disambiguation) * Middle Creek ...
to
Late Pleistocene The Late Pleistocene is an unofficial Age (geology), age in the international geologic timescale in chronostratigraphy, also known as the Upper Pleistocene from a Stratigraphy, stratigraphic perspective. It is intended to be the fourth division ...
epoch.


Description

''P. o. augusta'' was 15-20% larger than most Jaguar populations. During the
Irvingtonian The Irvingtonian North American Land Mammal Age on the geologic timescale is the North American faunal stage according to the North American Land Mammal Ages chronology (NALMA), spanning from 1.8 million – 250,000 years Before Present, BP.
, this subspecies could’ve averaged , with the largest individual weighing , but the subspecies went through a downsize with
Rancholabrean The Rancholabrean North American Land Mammal Age on the geologic timescale is a North American faunal stage in the North American Land Mammal Ages chronology (NALMA),Sanders, A.E., R.E. Weems, and L.B. Albright III (2009) Formalization of the mid- ...
populations being smaller than Irvingtonian populations.


History and distribution

Although ''P. onca'' still resides in the southern United States and several Native American tales possibly about ''P. onca augusta'' exist, the first published remains were described in 1872. The fossils were found by
Ferdinand Vandeveer Hayden Ferdinand Vandeveer Hayden (September 7, 1829 – December 22, 1887) was an American geologist noted for his pioneering surveying expeditions of the Rocky Mountains in the late 19th century. He was also a physician who served with the Union A ...
on the
Platte River The Platte River () is a major American river, in the state of Nebraska. It is about long; measured to its farthest source via its tributary, the North Platte River, it flows for over . The Platte River is a tributary of the Missouri River, w ...
of Nebraska and sent to
Joseph Leidy Joseph Mellick Leidy (September 9, 1823 – April 30, 1891) was an American paleontologist, parasitologist and anatomist. Leidy was professor of anatomy at the University of Pennsylvania, later becoming a professor of natural history at Swarth ...
at the Academy of Natural Sciences in Philadelphia, who named them ''Felis augustus'' in 1872. The fossils consisted of a
premolar The premolars, also called premolar Tooth (human), teeth, or bicuspids, are transitional teeth located between the Canine tooth, canine and Molar (tooth), molar teeth. In humans, there are two premolars per dental terminology#Quadrant, quadrant in ...
IV and
maxilla In vertebrates, the maxilla (: maxillae ) is the upper fixed (not fixed in Neopterygii) bone of the jaw formed from the fusion of two maxillary bones. In humans, the upper jaw includes the hard palate in the front of the mouth. The two maxil ...
fragment, the type ( USNM 1004), as well as another tooth and a distal humerus, though the humerus is likely that of '' Panthera atrox'' or ''
Smilodon ''Smilodon'' is an extinct genus of Felidae, felids. It is one of the best known saber-toothed predators and prehistoric mammals. Although commonly known as the saber-toothed tiger, it was not closely related to the tiger or other modern cats ...
fatalis''. Leidy also mistakenly believed the fossils were from the
Pliocene The Pliocene ( ; also Pleiocene) is the epoch (geology), epoch in the geologic time scale that extends from 5.33 to 2.58Pleistocene The Pleistocene ( ; referred to colloquially as the ''ice age, Ice Age'') is the geological epoch (geology), epoch that lasted from to 11,700 years ago, spanning the Earth's most recent period of repeated glaciations. Before a change was fin ...
. Later in 1919,
Oliver Perry Hay Oliver Perry Hay (May 22, 1846 – November 2, 1930) was an American herpetologist, ichthyologist, and paleontologist. Hay was born in Jefferson County, Indiana, to Robert and Margaret Hay. In 1870, Hay graduated with a bachelor of arts from ...
described a left canine from Vero, Florida that he named ''Felis veronis'', though it is now seen as a synonym of ''P. onca augusta.'' In 1929, George Simpson referred several teeth from Seminole Field, Florida to ''F. veronis'' and the same year a right mandible and several teeth from
Melbourne, Florida Melbourne ( ) is a city in Brevard County, Florida, United States. It is located southeast of Orlando, Florida, Orlando along Florida's Space Coast, named because of the region's proximity to Cape Canaveral and the Kennedy Space Center. The city ...
were collected by Dr. J. W. Gidley. All material found has been referred to ''P. onca augusta'', the mandible notably was the first complete one found of the taxon. Yet another discovery came in 1938, with the discovery of 2 fragmentary postcranial skeletons of ''P. onca augusta'' in Cumberland Cave, Maryland intermingled with that of a puma. The most productive discoveries outside of California came in 1939 and 1944, with the discoveries of footprints and several fossils in Craighead Caverns, Tennessee during the former year. 1944 saw the collection of 2, well preserved skeletons in Salt River Cave, also in Tennessee, that included 2 partial skulls and many axial elements that clearly demonstrated the subspecies’ clear differences with other taxa. Many more fossils were later collected by the
American Museum of Natural History The American Museum of Natural History (AMNH) is a natural history museum on the Upper West Side of Manhattan in New York City. Located in Theodore Roosevelt Park, across the street from Central Park, the museum complex comprises 21 interconn ...
at Craighead Caverns, including several mandibles and partial craniums. Additional fossils referable to ''P. onca augusta'' have been found in Oregon and, most notably, the
La Brea Tar Pits La Brea Tar Pits comprise an active Paleontological site, paleontological research site in urban Los Angeles. Hancock Park was formed around a group of tar pits where natural Bitumen, asphalt (also called asphaltum, bitumen, or pitch; ''brea'' ...
of California. ''P. onca augusta'' is most frequently found in Florida’s localities, as there are many fossil-bearing sites from the Pleistocene throughout Florida. In 2021, a partial mandible was referred to ''P. onca augusta'' from Chapala, Mexico, extending the range south to southwestern Mexico. A possible specimen of ''P. onca augusta'' is also known from Georgia, and mitogenomic analysis suggests that the specimen certainly belongs to ''P. onca'', though its subspecific assignment remains unresolved.


See also

*''
Panthera onca mesembrina ''Panthera onca mesembrina'', also known as the Patagonian panther, is an extinct subspecies of jaguar (''Panthera onca'') that was endemic to southern Patagonia during the late Pleistocene epoch. It is known from several fragmentary specimens, ...
'' * ''
Panthera gombaszoegensis ''Panthera gombaszoegensis'', also known as the European jaguar, is a ''Panthera'' species that lived from about 2.0 to 0.3 million years ago in Europe, as well as likely elsewhere in Eurasia. The first fossils were Excavation (archaeology), exca ...
''


References

{{Taxonbar, from=Q2710152 Jaguars onca augusta Pleistocene carnivorans Pleistocene mammals of North America Pleistocene mammals of South America Prehistoric mammals of North America Pleistocene first appearances Pleistocene extinctions