The natterjack toad (''Epidalea calamita'') is a

toad Toad (also known as a hoptoad) is a common name for certain frogs, especially of the family Bufonidae, that are characterized by dry, leathery skin, short legs, and large bumps covering the parotoid glands. In popular culture (folk taxonomy ...

native to sandy and

heathland A heath () is a shrubland habitat found mainly on free-draining infertile, acidic soils and is characterised by open, low-growing woody vegetation. Moorland is generally related to high-ground heaths with—especially in Great Britain—a coole ...

areas of

Europe Europe is a continent located entirely in the Northern Hemisphere and mostly in the Eastern Hemisphere. It is bordered by the Arctic Ocean to the north, the Atlantic Ocean to the west, the Mediterranean Sea to the south, and Asia to the east ...

and the United Kingdom. Adults are in length, and are distinguished from

common toad The common toad, European toad, or in Anglophone parts of Europe, simply the toad (''Bufo bufo'', from Latin ''bufo'' "toad"), is a toad found throughout most of Europe (with the exception of Ireland, Iceland, parts of Scandinavia, and some List ...

s by a yellow line down the middle of the back and parallel paratoid glands. They have relatively short legs, which gives them a distinctive gait, contrasting with the hopping movement of many other toad species. Natterjacks have a very loud and distinctive

mating call A mating call is the auditory signal used by animals to attract mates. It can occur in males or females, but literature is abundantly favored toward researching mating calls in females. In addition, mating calls are often the subject of mate choi ...

amplified by the single vocal sac found under the chin of the male.

Life history

Natterjacks live for up to 15 years, and feed mainly on

insects Insects (from Latin ') are hexapod invertebrates of the class Insecta. They are the largest group within the arthropod phylum. Insects have a chitinous exoskeleton, a three-part body (head, thorax and abdomen), three pairs of jointed ...

, especially beetles. At night, they move around open terrain with sparse vegetation, and their tracks can often be seen in loose sand. They move considerable distances each night, enabling the species to colonize new habitats very quickly.

Reproduction

The natterjack toad spawns between the end of April and July, laying strings of eggs in shallow, warm pools. Because the natterjack toad is often present in low numbers, its loud mating calls are important so that the sexes can find each other. Natterjacks can only spawn in pools that have a very slightly sloping bottom, and tolerate only sparse vegetation on the banks and in the water. Since pools of this description are often temporary, sometimes the tadpoles die when the pools dry out. The natterjack compensates for that risk by mating over an extended period each summer. Thus, in September, the age of the juveniles can vary from one to three months. Early breeders seldom breed again later in the season, though some females do spawn twice in a year.

Distribution

Populations of the natterjack extend through 17 European countries. In the UK, the toad is now almost completely confined to coastal sites. The natterjack is the only species of toad native to Ireland. It is found in

County Kerry County Kerry () is a Counties of Ireland, county on the southwest coast of Republic of Ireland, Ireland, within the Provinces of Ireland, province of Munster and the Southern Region, Ireland, Southern Region. It is bordered by two other countie ...

( Dingle Peninsula and Derrynane) and also in

County Wexford County Wexford () is a Counties of Ireland, county in Republic of Ireland, Ireland. It is in the Provinces of Ireland, province of Leinster and is part of the Southern Region, Ireland, Southern Region. Named after the town of Wexford, it was ba ...

, where it was introduced to a dune site. In mainland Europe, particularly in the southern part of its range, it lives inland in a variety of habitats.

Conservation

UK

In the UK, the threatened status of the species resulted in a national Biodiversity Action Plan designating it as one of three protected amphibians. Reasons for its threatened status include: * Loss of habitat from

human overpopulation Human overpopulation (or human population overshoot) is the idea that human populations may become too large to be sustainability, sustained by their environment or resources in the long term. The topic is usually discussed in the context of wor ...

* Deterioration and loss of lowland heaths * Reduction in habitable coast from construction of dykes and seawalls * Acidification of aquatic habitat from

acid rain Acid rain is rain or any other form of Precipitation (meteorology), precipitation that is unusually acidic, meaning that it has elevated levels of hydrogen ions (low pH). Most water, including drinking water, has a neutral pH that exists b ...

and other

pollution Pollution is the introduction of contaminants into the natural environment that cause harm. Pollution can take the form of any substance (solid, liquid, or gas) or energy (such as radioactivity, heat, sound, or light). Pollutants, the component ...

In England, the toad's

sand dune A dune is a landform composed of wind- or water-driven sand. It typically takes the form of a mound, ridge, or hill. An area with dunes is called a dune system or a dune complex. A large dune complex is called a dune field, while broad, flat ...

habitat is protected by a number of national nature reserves. For example, in the north-west, reserves are at

Hoylake Hoylake () is a coast, seaside town in the Metropolitan Borough of Wirral, Merseyside, England. It is at the north west of the Wirral Peninsula, near West Kirby and where the River Dee, Wales, River Dee meets the Irish Sea. At the 2021 United K ...

, Ainsdale Sand Dunes, North Walney and Sandscale Haws. In Scotland, where the species is confined to the

Solway Firth The Solway Firth is an inlet on the west coast of Great Britain, forming part of the border between England and Scotland. The firth (a Scottish term for an inlet of the sea) divides Cumbria (including the Solway Plain) from Dumfries and Gallow ...

, a reserve is at Caerlaverock. In Wales, the species became extinct in the 20th century, but has been reintroduced as part of the Biodiversity Action Plan.

Ireland

To reverse habitat loss, the National Parks and Wildlife Service has created ponds for the species with some funding from the Heritage Council. The natterjack is considered endangered in Ireland. , a project involving Fota Wildlife Park and the National Parks and Wildlife Service has released 6,000 toadlets into their natural range in County Kerry. The natterjack toad is used as a symbol for Dooks Golf club in Kerry.

References

External links

Amphibians and Reptiles of EuropeFauna Europaea
West

Palaearctic The Palearctic or Palaearctic is a biogeographic realm of the Earth, the largest of eight. Confined almost entirely to the Eastern Hemisphere, it stretches across Europe and Asia, north of the foothills of the Himalayas, and North Africa. Th ...

distribution.
Natterjack Toad Species Action Plan
(pdf). Amphibian & Reptile Conservation (July 2009) — via Internet Archive.
Natterjack Toad V 2.1. (Species Action Plan)
(pdf) Cumbria Biodiversity Data Centre (April 2010).
Competencies for Species Survey: Natterjack Toad
(pdf). Chartered Institute of Ecology and Environmental Management – CIEEM (25 April 2013). * * * * * Last updated 2015-05-12. {{Authority control Bufonidae Amphibians of Europe Amphibians described in 1768 Articles containing video clips Taxa named by Josephus Nicolaus Laurenti Habitats Directive species