''Polyscias racemosa'', or false 'ohe, is a

species A species () is often defined as the largest group of organisms in which any two individuals of the appropriate sexes or mating types can produce fertile offspring, typically by sexual reproduction. It is the basic unit of Taxonomy (biology), ...

flowering plant Flowering plants are plants that bear flowers and fruits, and form the clade Angiospermae (). The term angiosperm is derived from the Ancient Greek, Greek words (; 'container, vessel') and (; 'seed'), meaning that the seeds are enclosed with ...

in the

family Family (from ) is a Social group, group of people related either by consanguinity (by recognized birth) or Affinity (law), affinity (by marriage or other relationship). It forms the basis for social order. Ideally, families offer predictabili ...

Araliaceae The Araliaceae are a family of flowering plants composed of about 43 genera and around 1500 species consisting of primarily woody plants and some herbaceous plants commonly called the ginseng family. The morphology of Araliaceae varies widely ...

. As ''Munroidendron racemosum'', the species was until recently considered to be the only species in the

monotypic In biology, a monotypic taxon is a taxonomic group (taxon) that contains only one immediately subordinate taxon. A monotypic species is one that does not include subspecies or smaller, infraspecific taxa. In the case of genera, the term "unisp ...

genus ''Munroidendron''. With the change in classification, ''Munroidendron'' is now obsolete. ''Polyscias racemosa'' is

endemic Endemism is the state of a species being found only in a single defined geographic location, such as an island, state, nation, country or other defined zone; organisms that are indigenous to a place are not endemic to it if they are also foun ...

to the

Hawaii Hawaii ( ; ) is an island U.S. state, state of the United States, in the Pacific Ocean about southwest of the U.S. mainland. One of the two Non-contiguous United States, non-contiguous U.S. states (along with Alaska), it is the only sta ...

island An island or isle is a piece of land, distinct from a continent, completely surrounded by water. There are continental islands, which were formed by being split from a continent by plate tectonics, and oceanic islands, which have never been ...

Kauai Kauai (), anglicized as Kauai ( or ), is one of the main Hawaiian Islands. It has an area of 562.3 square miles (1,456.4 km2), making it the fourth-largest of the islands and the 21st-largest island in the United States. Kauai lies 73 m ...

.David G. Frodin and Rafaël Govaerts. 2003. ''World Checklist and Bibliography of Araliaceae''. Royal Botanic Gardens, Kew. . (See ''External links'' below). It is very rare in the

wild Wild, wild, wilds or wild may refer to: Common meanings * Wilderness, a wild natural environment * Wildlife, an undomesticated organism * Wildness, the quality of being wild or untamed Art, media and entertainment Film and television * ''Wild ...

and some of its original

habitat In ecology, habitat refers to the array of resources, biotic factors that are present in an area, such as to support the survival and reproduction of a particular species. A species' habitat can be seen as the physical manifestation of its ...

has been replaced by

sugar cane Sugarcane or sugar cane is a species of tall, Perennial plant, perennial grass (in the genus ''Saccharum'', tribe Andropogoneae) that is used for sugar Sugar industry, production. The plants are 2–6 m (6–20 ft) tall with stout, jointed, fib ...

plantation Plantations are farms specializing in cash crops, usually mainly planting a single crop, with perhaps ancillary areas for vegetables for eating and so on. Plantations, centered on a plantation house, grow crops including cotton, cannabis, tob ...

s.Earl Edward Sherff. 1956. "Some Recently Collected Dicotyledonous Hawaiian Island and Peruvian Plants". ''American Journal of Botany'' 43(7):475-478. It was thought for some time to be probably

extinct Extinction is the termination of an organism by the death of its Endling, last member. A taxon may become Functional extinction, functionally extinct before the death of its last member if it loses the capacity to Reproduction, reproduce and ...

, but was rediscovered a few years prior to 1967.Benjamin C. Stone. 1967. "A review of the endemic genera of Hawaiian plants" ''Botanical Review'' (Lancaster) 33(3):216-259. Using

cladistic Cladistics ( ; from Ancient Greek 'branch') is an approach to biological classification in which organisms are categorized in groups ("clades") based on hypotheses of most recent common ancestry. The evidence for hypothesized relationships is ...

methods,

phylogenetic In biology, phylogenetics () is the study of the evolutionary history of life using observable characteristics of organisms (or genes), which is known as phylogenetic inference. It infers the relationship among organisms based on empirical dat ...

studies Study or studies may refer to: General * Education **Higher education * Clinical trial * Experiment * Field of study * Observational study * Scientific study * Research * Study skills, abilities and approaches applied to learning Other * Study ...

DNA Deoxyribonucleic acid (; DNA) is a polymer composed of two polynucleotide chains that coil around each other to form a double helix. The polymer carries genetic instructions for the development, functioning, growth and reproduction of al ...

have shown that the closest relative of ''Munroidendron racemosum'' is '' Reynoldsia sandwicensis''.Gregory M. Plunkett and Porter P. Lowry II. 2010. "Paraphyly and polyphyly in ''Polyscias'' sensu lato: molecular evidence and the case for recircumscribing the "pinnate genera" of Araliaceae". ''Plant Diversity and Evolution'' (formerly ''Botanische Jahrbucher'') 128(1-2):23-54. . These two

are now known as ''Polyscias racemosa'' and ''

Polyscias sandwicensis ''Polyscias sandwicensis'' (formerly '' Reynoldsia sandwicensis''), known in Hawaiian as the 'ohe makai or ''Ohe kukuluāeo'', is a species of flowering plant in the family Araliaceae, that is endemic to Hawaii. It is a Hawaiian dry forest tre ...

'', respectively. They are two of the 21 species now placed in ''Polyscias'' subgenus ''Tetraplasandra''.Porter P. Lowry II and Gregory M. Plunkett. 2010. "Recircumscription of ''Polyscias'' (Araliaceae) to include six related genera, with a new infrageneric classification and a synopsis of species". ''

Plant Diversity and Evolution This is a list of academic journals, monographic series Monographic series (alternatively, monographs in series) are scholarly and scientific books released in successive volumes, each of which is structured like a separate book or scholarly mo ...

'' (formerly ''Botanische Jahrbucher'') 128(1-2):55-84. . (See ''External links'' below). ''Polyscias racemosa'' is known in cultivation in Hawaii.Clyde T. Imada, George W. Staples, and Derral R. Herbst. undated. Annotated Checklist of Cultivated Plants of Hawai‘i. (See ''External links'' below). Cultivation procedures for ''Polyscias racemosa'' have been studied.Kerin E. Lilleeng-Rosenberger. 2005. ''Growing Hawaii's Native Plants''. Mutual Publishing.

Description

''Polyscias racemosa'' is a small

tree In botany, a tree is a perennial plant with an elongated stem, or trunk, usually supporting branches and leaves. In some usages, the definition of a tree may be narrower, e.g., including only woody plants with secondary growth, only ...

growing to tall, with a straight

trunk Trunk may refer to: Biology * Trunk (anatomy), synonym for torso * Trunk (botany), a tree's central superstructure, and the stem of woody plants * Trunk of corpus callosum, in neuroanatomy * Elephant trunk, the proboscis of an elephant Comput ...

, spreading

branch A branch, also called a ramus in botany, is a stem that grows off from another stem, or when structures like veins in leaves are divided into smaller veins. History and etymology In Old English, there are numerous words for branch, includ ...

es, and smooth, grey

bark Bark may refer to: Common meanings * Bark (botany), an outer layer of a woody plant such as a tree or stick * Bark (sound), a vocalization of some animals (which is commonly the dog) Arts and entertainment * ''Bark'' (Jefferson Airplane album), ...

.Warren L. Wagner, Derral R. Herbst, and Sy H. Sohmer. ''Manual of the Flowering Plants of Hawaii'', Revised Edition, 1999. Bishop Museum Press: Hololulu Like many

members Member may refer to: * Military jury, referred to as "Members" in military jargon * Element (mathematics), an object that belongs to a mathematical set * In object-oriented programming, a member of a class ** Field (computer science), entries in ...

of ''Polyscias'', it is sparingly branched and thick- stemmed, with large

imparipinnate Pinnation (also called pennation) is the arrangement of feather-like or multi-divided features arising from both sides of a common axis. Pinnation occurs in biological morphology, in crystals, such as some forms of ice or metal crystals, and in ...

leaves, but not as extreme in these characteristics as is '' Polyscias nodosa''. Its

leaves A leaf (: leaves) is a principal appendage of the stem of a vascular plant, usually borne laterally above ground and specialized for photosynthesis. Leaves are collectively called foliage, as in "autumn foliage", while the leaves, stem, ...

are

pinnate Pinnation (also called pennation) is the arrangement of feather-like or multi-divided features arising from both sides of a common axis. Pinnation occurs in biological morphology, in crystals, such as some forms of ice or metal crystals, and ...

, long, with oval leaflets, each of which is over long. These trees are

dry season The dry season is a yearly period of low rainfall, especially in the tropics. The weather in the tropics is dominated by the tropical rain belt, which moves from the northern to the southern tropics and back over the course of the year. The t ...

deciduous In the fields of horticulture and botany, the term deciduous () means "falling off at maturity" and "tending to fall off", in reference to trees and shrubs that seasonally shed Leaf, leaves, usually in the autumn; to the shedding of petals, aft ...

, dropping most of their leaves during their

summer Summer or summertime is the hottest and brightest of the four temperate seasons, occurring after spring and before autumn. At or centred on the summer solstice, daylight hours are the longest and darkness hours are the shortest, with day ...

blooming Bloom or blooming may refer to: Science and technology Biology * Bloom, one or more flowers on a flowering plant * Algal bloom, a rapid increase or accumulation in the population of algae in an aquatic system * Jellyfish bloom, a collective n ...

season A season is a division of the year based on changes in weather, ecology, and the number of daylight hours in a given region. On Earth, seasons are the result of the axial parallelism of Earth's axial tilt, tilted orbit around the Sun. In temperat ...

. Its small, pale

yellow Yellow is the color between green and orange on the spectrum of light. It is evoked by light with a dominant wavelength of roughly 575585 nm. It is a primary color in subtractive color systems, used in painting or color printing. In t ...

flower Flowers, also known as blooms and blossoms, are the reproductive structures of flowering plants ( angiosperms). Typically, they are structured in four circular levels, called whorls, around the end of a stalk. These whorls include: calyx, m ...

s hang in long, rope-like strands. The

inflorescence In botany, an inflorescence is a group or cluster of flowers arranged on a plant's Plant stem, stem that is composed of a main branch or a system of branches. An inflorescence is categorized on the basis of the arrangement of flowers on a mai ...

racemose A raceme () or racemoid is an unbranched, indeterminate type of inflorescence bearing flowers having short floral stalks along the shoots that bear the flowers. The oldest flowers grow close to the base and new flowers are produced as the shoo ...

in form, with up to 250 flowers.Peter S. Green (author) and Mary Grierson (illustrator). 1996. ''A Hawaiian Florilegium: Botanical Portraits from Paradise''. University of Hawaii Press: Honolulu, Hawaii. . File:Polyscias racemosa (5454956091).jpg, Flowers File:Polyscias racemosa (5455568842).jpg, Inflorescence File:Starr-090421-6250-Polyscias racemosa-leaves-Keikilani Rd Pukalani-Maui (24952473915).jpg, Leaves File:Starr 061108-9821 Munroidendron racemosum.jpg, New Leaves

Habitat and range

''Polyscias racemosa'' occurs in coastal mesic and mixed mesic forests at

elevation The elevation of a geographic location (geography), ''location'' is its height above or below a fixed reference point, most commonly a reference geoid, a mathematical model of the Earth's sea level as an equipotential gravitational equipotenti ...

s of , where it grows on exposed cliffs and ridges. Associated

plant Plants are the eukaryotes that form the Kingdom (biology), kingdom Plantae; they are predominantly Photosynthesis, photosynthetic. This means that they obtain their energy from sunlight, using chloroplasts derived from endosymbiosis with c ...

species include papala kepau (''

Pisonia umbellifera ''Ceodes umbellifera'', Synonym (taxonomy), synonym ''Pisonia umbellifera'', commonly known as the birdlime tree or bird catcher tree, is a species of plant in the Nyctaginaceae family. The evergreen shrub has soft wood, small pink or yellow flowe ...

''), āwikiwiki (''

Canavalia ''Canavalia'' is a genus of plants in the legume family (Fabaceae) that comprises approximately 62 species of tropical vines. Members of the genus are commonly known as jack-beans. It has a pantropical distribution. The species of ''Canavalia'' ...

galeata Galeata () is a ''comune'' (municipality) in the Province of Forlì-Cesena in the Italian region Emilia-Romagna, located about southeast of Bologna and about southwest of Forlì. Galeata borders the following municipalities: Civitella di Romag ...

''), ilima (''

Sida fallax ''Sida fallax'', known as yellow ilima, golden mallow,, or Ilima is a species of Herbaceous plant, herbaceous flowering plant in the ''Hibiscus'' family, Malvaceae, indigenous to the Hawaiian Archipelago and other Pacific Islands. Plants may be e ...

''), ōlulu (''

Brighamia insignis ''Brighamia insignis'', commonly known as ''ʻŌlulu'' or ''Alula'' in Hawaiian, or colloquially as the vulcan palm or cabbage on a stick, is a species of Hawaiian lobelioid in the bellflower family, Campanulaceae. It is native to the islands o ...

''), alahee (''

Psydrax odorata ''Psydrax odorata'', known as ''alahee'' in Hawaiian, is a species of flowering shrub or small tree in the coffee family, Rubiaceae. It is native to the Pacific Islands, New Guinea and Australia. Description The species range from in height, h ...

''), kōpiko (''

Psychotria ''Psychotria'' is a large genus of flowering plants in the coffee family Rubiaceae, with over 1,600 species. The genus has a pantropical distribution and members of the genus are small understorey trees in tropical forests. Some species are endan ...

'' spp.), olopua (''

Nestegis sandwicensis ''Notelaea sandwicensis'', commonly known as Hawai'i olive or ''olopua'', is a species of flowering tree in the olive family, Oleaceae, that is endemic to Hawaii. It is found on all major islands at elevations of in coastal mesic and mixed mes ...

''), ahakea ('' Bobea timonioides''), hala pepe ('' Pleomele aurea''), and ālaa (''

Planchonella sandwicensis ''Planchonella sandwicensis'' is a species of flowering tree in the sapodilla family, Sapotaceae, that is endemic to the main islands of Hawaii. Names for this species in the Hawaiian language include ''Ālaa'', ''Āulu'' and ''Ēlaa''. Descrip ...

''). It occurs naturally in only three locations on Kauai: Nounou Mountain, the cliffs of the Nā Pali Coast, and Haupu Ridge near Nāwiliwili Bay.

History

''Polyscias racemosa'' first entered the

botanical literature Botany, also called plant science, is the branch of natural science and biology studying plants, especially their anatomy, taxonomy, and ecology. A botanist or plant scientist is a scientist who specialises in this field. "Plant" and "botany ...

in 1917, when it was described and named as ''Tetraplasandra racemosa'' by Charles Noyes Forbes.Charles Noyes Forbes. 1917. "New Hawaiian Plants.-VI." ''Occasional Papers of the Bernice Pauahi Bishop Museum'' 6(4):51. (See ''External links'' below). Earl Edward Sherff felt that this species was uniquely distinct from the rest of ''Tetraplasandra'', so he erected a new genus for it, ''Munroidendron'', in 1952.''Munroidendron'' in

International Plant Names Index The International Plant Names Index (IPNI) describes itself as "a database of the names and associated basic bibliographical details of seed plants, ferns and lycophytes." Coverage of plant names is best at the rank of species and genus. It inclu ...

. (see ''External links'' below). The genus was named for

George Campbell Munro George Campbell Munro (10 May 1866 – 4 December 1963) was a New Zealand born pioneer of Hawaiian botany and ornithology. He settled on a ranch in Lanai and wrote one of the first books on the birds of Hawaii, many species of which are now extinct ...

(1866-1963), described by Umberto Quattrocchi as "a pioneer in Hawaiian

ornithology Ornithology, from Ancient Greek ὄρνις (''órnis''), meaning "bird", and -logy from λόγος (''lógos''), meaning "study", is a branch of zoology dedicated to the study of birds. Several aspects of ornithology differ from related discip ...

botany Botany, also called plant science, is the branch of natural science and biology studying plants, especially Plant anatomy, their anatomy, Plant taxonomy, taxonomy, and Plant ecology, ecology. A botanist or plant scientist is a scientist who s ...

, and

horticulture Horticulture (from ) is the art and science of growing fruits, vegetables, flowers, trees, shrubs and ornamental plants. Horticulture is commonly associated with the more professional and technical aspects of plant cultivation on a smaller and mo ...

;

plant collector A botanical specimen, also called a plant specimen, is a biological specimen of a plant (or part of a plant) used for scientific purposes. Preserved collections of algae, fungi, slime molds, and other organisms traditionally studied by botanists a ...

in the Hawaiian Islands".Umberto Quattrocchi. 2000. ''CRC World Dictionary of Plant Names'', volume III. CRC Press: Baton Rouge, New York, London, Washington DC. (vol. III). (see ''External links'' below). ''Dendron'' is a

Greek Greek may refer to: Anything of, from, or related to Greece, a country in Southern Europe: *Greeks, an ethnic group *Greek language, a branch of the Indo-European language family **Proto-Greek language, the assumed last common ancestor of all kno ...

word for "tree". Munro was apparently the first collector to see his

eponym An eponym is a noun after which or for which someone or something is, or is believed to be, named. Adjectives derived from the word ''eponym'' include ''eponymous'' and ''eponymic''. Eponyms are commonly used for time periods, places, innovati ...

ous genus, ''Munroidendron''.Earl Edward Sherff. 1952. "''Munroidendron'', a new genus of Araliaceous trees from the island of Kauai". ''Botanical Leaflets'' 7(section V):21-24. published by the author. Sherff separated ''Munroidendron'' from ''Tetraplasandra'' on the basis of five characters: the absence of

umbel UMBEL (Upper Mapping and Binding Exchange Layer) is a logically organized knowledge graph of 34,000 concepts and entity types that can be used in information science for relating information from disparate sources to one another. It was retired ...

lules, the

arrangement In music, an arrangement is a musical adaptation of an existing composition. Differences from the original composition may include reharmonization, melodic paraphrasing, orchestration, or formal development. Arranging differs from orchestr ...

of the

s in a

raceme A raceme () or racemoid is an unbranched, indeterminate growth, indeterminate type of inflorescence bearing flowers having short floral stalks along the shoots that bear the flowers. The oldest flowers grow close to the base and new flowers are ...

, the sunken,

diamond Diamond is a Allotropes of carbon, solid form of the element carbon with its atoms arranged in a crystal structure called diamond cubic. Diamond is tasteless, odourless, strong, brittle solid, colourless in pure form, a poor conductor of e ...

-shaped

pedicel Pedicle or pedicel may refer to: Human anatomy *Pedicle of vertebral arch, the segment between the transverse process and the vertebral body, and is often used as a radiographic marker and entry point in vertebroplasty and kyphoplasty procedures ...

scars, the long,

persistence Persistence or Persist may refer to: Math and computers * Image persistence, in LCD monitors * Persistence (computer science), the characteristic of data that outlives the execution of the program that created it * Persistence of a number, a ma ...

of the subtending floral

bract In botany, a bract is a modified or specialized leaf, associated with a reproductive structure such as a flower, inflorescence axis or cone scale. Bracts are usually different from foliage leaves in size, color, shape or texture. They also lo ...

s, and the

insertion Insertion may refer to: *Insertion (anatomy), the point of a tendon or ligament onto the skeleton or other part of the body *Insertion (genetics), the addition of DNA into a genetic sequence *Insertion, several meanings in medicine, see ICD-10-PCS ...

of the

stamen The stamen (: stamina or stamens) is a part consisting of the male reproductive organs of a flower. Collectively, the stamens form the androecium., p. 10 Morphology and terminology A stamen typically consists of a stalk called the filament ...

s in only one

whorl A whorl ( or ) is an individual circle, oval, volution or equivalent in a whorled pattern, which consists of a spiral or multiple concentric objects (including circles, ovals and arcs). In nature File:Photograph and axial plane floral diagra ...

, even when numerous. It has been shown that, in spite of its appearance, the inflorescence is not truly a raceme because it is determinate. Sherff divided the species now known as ''Polyscias racemosa'' into three

varieties Variety may refer to: Arts and entertainment Entertainment formats * Variety (radio) * Variety show, in theater and television Films * ''Variety'' (1925 film), a German silent film directed by Ewald Andre Dupont * ''Variety'' (1935 film), ...

: var. racemosa, var. forbesii, and var. macdanielsii. These have been described as "not sufficiently distinct to be retained". The establishment of ''Munroidendron'' was contentious from the beginning. William R. Philipson said that ''Munroidendron'' "comprises a single species with such a distinct inflorescence and corolla that it can well claim generic

status Status (Latin plural: ''statūs''), is a state, condition, or situation, and may refer to: * Status (law) ** Legal status, in law ** Political status, in international law ** Small entity status, in patent law ** Status conference ** Status c ...

.William R. Philipson. 1970. "A redefinition of ''Gastonia'' and related genera (Araliaceae)". ''Blumea'' 18(2):497-505. In 1971, a

pollen Pollen is a powdery substance produced by most types of flowers of seed plants for the purpose of sexual reproduction. It consists of pollen grains (highly reduced Gametophyte#Heterospory, microgametophytes), which produce male gametes (sperm ...

study indicated that ''Munroidendron'' might be embedded in ''Tetraplasandra''.Charles C. Tseng. 1971. "Light and Scanning Electron Microscopic Studies on Pollen of ''Tetraplasandra'' (Araliaceae) and Relatives". ''American Journal of Botany'' 58(6):505-516. This result was not supported by

molecular phylogenetic Molecular phylogenetics () is the branch of phylogeny that analyzes genetic, hereditary molecular differences, predominantly in DNA sequences, to gain information on an organism's evolutionary relationships. From these analyses, it is possible to ...

based on

DNA sequence A nucleic acid sequence is a succession of bases within the nucleotides forming alleles within a DNA (using GACT) or RNA (GACU) molecule. This succession is denoted by a series of a set of five different letters that indicate the order of the nu ...

s of

nuclear Nuclear may refer to: Physics Relating to the nucleus of the atom: *Nuclear engineering *Nuclear physics *Nuclear power *Nuclear reactor *Nuclear weapon *Nuclear medicine *Radiation therapy *Nuclear warfare Mathematics * Nuclear space *Nuclear ...

and

chloroplast A chloroplast () is a type of membrane-bound organelle, organelle known as a plastid that conducts photosynthesis mostly in plant cell, plant and algae, algal cells. Chloroplasts have a high concentration of chlorophyll pigments which captur ...

DNA regions. These studies show that ''

'' (formerly '' Reynoldsia sandwicensis'' is not most closely related to other species of '' Reynoldsia'', but is

sister A sister is a woman or a girl who shares parents or a parent with another individual; a female sibling. The male counterpart is a brother. Although the term typically refers to a familial relationship, it is sometimes used endearingly to ref ...

to ''Polyscias racemosa'', (formerly ''Munroidendron'').Annemarie Costello and Timothy J. Motley. 2007. "Phylogenetics of the Tetraplasandra Group (Araliaceae) Inferred from ITS, 5S-NTS, and Morphology". ''Systematic Botany'' 32(2):464-477. This pair is then sister to a

monophyletic In biological cladistics for the classification of organisms, monophyly is the condition of a taxonomic grouping being a clade – that is, a grouping of organisms which meets these criteria: # the grouping contains its own most recent co ...

'' Tetraplasandra'' in the

sense A sense is a biological system used by an organism for sensation, the process of gathering information about the surroundings through the detection of Stimulus (physiology), stimuli. Although, in some cultures, five human senses were traditio ...

of Philipson (1970). This pair of species and the nine species formerly in ''Tetraplasandra'' form a

clade In biology, a clade (), also known as a Monophyly, monophyletic group or natural group, is a group of organisms that is composed of a common ancestor and all of its descendants. Clades are the fundamental unit of cladistics, a modern approach t ...

and comprise all of the Hawaiian species of ''Polyscias''. The 11 species of this "Hawaiian clade" and 10 species from

Malesia Malesia is a biogeographical region straddling the Equator and the boundaries of the Indomalayan and Australasian realms. It is a phytogeographical floristic region in the Paleotropical kingdom. It was first recognized as a distinct region ...

Melanesia Melanesia (, ) is a subregion of Oceania in the southwestern Pacific Ocean. It extends from New Guinea in the west to the Fiji Islands in the east, and includes the Arafura Sea. The region includes the four independent countries of Fiji, Vanu ...

, and southern

Polynesia Polynesia ( , ) is a subregion of Oceania, made up of more than 1,000 islands scattered over the central and southern Pacific Ocean. The indigenous people who inhabit the islands of Polynesia are called Polynesians. They have many things in ...

constitute ''Polyscias'' subgenus Tetraplasandra.

Description

Habitat and range

History

References

External links