Morchella Disparilis
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''Morchella'', the true morels, is a
genus Genus (; : genera ) is a taxonomic rank above species and below family (taxonomy), family as used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In bino ...
of edible
sac fungi Ascomycota is a phylum of the kingdom Fungi that, together with the Basidiomycota, forms the subkingdom Dikarya. Its members are commonly known as the sac fungi or ascomycetes. It is the largest phylum of Fungi, with over 64,000 species. The def ...
closely related to anatomically simpler
cup fungi The Pezizaceae (commonly referred to as cup fungi) are a family of fungi in the Ascomycota which produce mushrooms that tend to grow in the shape of a "cup". Spores are formed on the inner surface of the fruit body ( ascoma). The cup shape typi ...
in the order
Pezizales The Pezizales are an order of the subphylum Pezizomycotina within the phylum Ascomycota. The order contains 16 families, 199 genera, and 1683 species. It contains a number of species of economic importance, such as morels, the black and whit ...
( division
Ascomycota Ascomycota is a phylum of the kingdom Fungi that, together with the Basidiomycota, forms the subkingdom Dikarya. Its members are commonly known as the sac fungi or ascomycetes. It is the largest phylum of Fungi, with over 64,000 species. The def ...
). These distinctive
fungi A fungus (: fungi , , , or ; or funguses) is any member of the group of eukaryotic organisms that includes microorganisms such as yeasts and mold (fungus), molds, as well as the more familiar mushrooms. These organisms are classified as one ...
have a
honeycomb A honeycomb is a mass of Triangular prismatic honeycomb#Hexagonal prismatic honeycomb, hexagonal prismatic cells built from beeswax by honey bees in their beehive, nests to contain their brood (eggs, larvae, and pupae) and stores of honey and pol ...
appearance due to the network of ridges with pits composing their
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. Morels are prized by
gourmet Gourmet (, ) is a cultural idea associated with the culinary arts of fine food and drink, or haute cuisine, which is characterized by their high level of refined and elaborate food preparation techniques and displays of balanced meals that have ...
cooks, particularly in Catalan and
French cuisine French cuisine is the cooking traditions and practices of France. In the 14th century, Guillaume Tirel, a Court (royal), court chef known as "Taillevent", wrote ''Le Viandier'', one of the earliest recipe collections of medieval France. In ...
, but can be toxic if consumed raw or undercooked. Due to difficulties in cultivation, commercial harvesting of wild morels has become a multimillion-dollar industry in the temperate
Northern Hemisphere The Northern Hemisphere is the half of Earth that is north of the equator. For other planets in the Solar System, north is defined by humans as being in the same celestial sphere, celestial hemisphere relative to the invariable plane of the Solar ...
, in particular
North America North America is a continent in the Northern Hemisphere, Northern and Western Hemisphere, Western hemispheres. North America is bordered to the north by the Arctic Ocean, to the east by the Atlantic Ocean, to the southeast by South Ameri ...
,
Turkey Turkey, officially the Republic of Türkiye, is a country mainly located in Anatolia in West Asia, with a relatively small part called East Thrace in Southeast Europe. It borders the Black Sea to the north; Georgia (country), Georgia, Armen ...
,
China China, officially the People's Republic of China (PRC), is a country in East Asia. With population of China, a population exceeding 1.4 billion, it is the list of countries by population (United Nations), second-most populous country after ...
, the
Himalayas The Himalayas, or Himalaya ( ), is a mountain range in Asia, separating the plains of the Indian subcontinent from the Tibetan Plateau. The range has some of the Earth's highest peaks, including the highest, Mount Everest. More than list of h ...
,
India India, officially the Republic of India, is a country in South Asia. It is the List of countries and dependencies by area, seventh-largest country by area; the List of countries by population (United Nations), most populous country since ...
, and
Pakistan Pakistan, officially the Islamic Republic of Pakistan, is a country in South Asia. It is the List of countries and dependencies by population, fifth-most populous country, with a population of over 241.5 million, having the Islam by country# ...
where these highly prized fungi are found in abundance. Typified by ''
Morchella esculenta ''Morchella esculenta'' (commonly known as common morel, morel, yellow morel, true morel, morel mushroom, and sponge morel) is a species of fungus in the family Morchellaceae of the Ascomycota. It is one of the most readily recognized of all th ...
'' in 1794, the genus has been the source of considerable
taxonomical In biology, taxonomy () is the scientific study of naming, defining ( circumscribing) and classifying groups of biological organisms based on shared characteristics. Organisms are grouped into taxa (singular: taxon), and these groups are given ...
controversy throughout the years, mostly with regard to the number of species involved, with some
mycologist Mycology is the branch of biology concerned with the study of fungi, including their taxonomy, genetics, biochemical properties, and use by humans. Fungi can be a source of tinder, food, traditional medicine, as well as entheogens, poison, and ...
s recognising as few as three species and others over thirty. Current
molecular phylogenetics Molecular phylogenetics () is the branch of phylogeny that analyzes genetic, hereditary molecular differences, predominantly in DNA sequences, to gain information on an organism's evolutionary relationships. From these analyses, it is possible to ...
suggest there might be over seventy species of ''Morchella'' worldwide, most of them exhibiting high continental
endemism Endemism is the state of a species being found only in a single defined geographic location, such as an island, state, nation, country or other defined zone; organisms that are indigenous to a place are not endemic to it if they are also foun ...
and provincialism. The genus is currently the focus of extensive
phylogenetic In biology, phylogenetics () is the study of the evolutionary history of life using observable characteristics of organisms (or genes), which is known as phylogenetic inference. It infers the relationship among organisms based on empirical dat ...
,
biogeographical Biogeography is the study of the distribution of species and ecosystems in geographic space and through geological time. Organisms and biological communities often vary in a regular fashion along geographic gradients of latitude, elevation, ...
, taxonomical and
nomenclatural Nomenclature (, ) is a system of names or terms, or the rules for forming these terms in a particular field of arts or sciences. (The theoretical field studying nomenclature is sometimes referred to as ''onymology'' or ''taxonymy'' ). The principl ...
studies, and several new species have been described from Australia, Canada, Cyprus, Israel, Spain, and Turkey.


Description

Morels resemble a
honeycomb A honeycomb is a mass of Triangular prismatic honeycomb#Hexagonal prismatic honeycomb, hexagonal prismatic cells built from beeswax by honey bees in their beehive, nests to contain their brood (eggs, larvae, and pupae) and stores of honey and pol ...
due to the network of ridges with pits composing their
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. Morels have a convoluted head/cap, and are varied in shape and habitat. File:Morchella americana on a mossy riverbank in MI.jpg File:Morchella_americana_closer.JPG File:Morchella esculenta 5.jpg File:Morchella esculenta 6.jpg File:Old holiday shot of Morchella esculenta (GB= Morel mushroom, D= Speise-Morchel, NL= Gewone morielje) - panoramio.jpg File:Morchella esculenta Illinois.jpg


Similar species

When foraging for morels, one must be absolutely sure of identification. There are many look-alikes often referred to as " false morels", most notably ''
Gyromitra ''Gyromitra'' () is a genus of about 18 species of ascomycete fungi. They are a false morel - a frequently toxic mushroom that can be mistaken for edible mushrooms of the genus ''Morchella'' (morels). Taxonomy The name ''Gyromitra'' comes from ...
''. These also include members of the most closely related genus, ''
Verpa ''Verpa'' is a genus of ascomycete fungi related to the morels. Resembling the latter genus in edibility and form, the common name early morels is popular. There are five species in the widespread genus. Taxonomy Analysis of the ribosomal DNA o ...
'', which are highly poisonous. Other mushrooms can also be mistaken for morels, including some species of stinkhorns, or
Phallaceae Phallaceae is a family of fungi, commonly known as stinkhorns, within the order Phallales. Stinkhorns have a worldwide distribution, but are especially prevalent in tropical regions. They are known for their foul-smelling, sticky spore masses ...
, which have a similarly shaped cap but a distinctive foul odor. It is important to take care when harvesting and identifying mushrooms, particularly morels. The key morphological features distinguishing false morels from true morels are as follows: * ''
Gyromitra ''Gyromitra'' () is a genus of about 18 species of ascomycete fungi. They are a false morel - a frequently toxic mushroom that can be mistaken for edible mushrooms of the genus ''Morchella'' (morels). Taxonomy The name ''Gyromitra'' comes from ...
'' species often have a "wrinkled" or "cerebral" (brain-like) appearance to the cap due to multiple wrinkles and folds, rather than the honeycomb appearance of true morels due to ridges and pits. * The caps of morels come in a range of colors including white, grey, black, brown, and yellow. ''Gyromitra esculenta'' has a cap that is usually reddish-brown in colour, but sometimes also chestnut, purplish-brown, or dark brown. * True morels are always hollow when sliced lengthwise, whereas ''Gyromitra'' species are typically chambered in longitudinal sections, while ''
Verpa ''Verpa'' is a genus of ascomycete fungi related to the morels. Resembling the latter genus in edibility and form, the common name early morels is popular. There are five species in the widespread genus. Taxonomy Analysis of the ribosomal DNA o ...
'' species contain a cottony substance inside their stem. The easiest way to distinguish ''Verpa'' species from ''Morchella'' species is to slice them longitudinally ,8 * The caps of ''Verpa'' species ('' V. bohemica'', '' V. conica'' and others) are attached to the stem only at the apex (top of the cap), unlike true morels which have caps that are attached to the stem at, or near the base of the cap.


Taxonomy

The
fruit bodies The sporocarp (also known as fruiting body, fruit body or fruitbody) of fungi is a multicellular structure on which spore-producing structures, such as basidia or asci, are borne. The fruitbody is part of the sexual phase of a fungal life cyc ...
of ''Morchella'' species are highly polymorphic, varying in shape, color, and size. While in many cases they do not exhibit clear-cut distinguishing features microscopically, this has historically contributed to uncertainties in taxonomy. Discriminating between the various taxa described is further hindered by uncertainty over which of these are truly biologically distinct. Remarkably, some authors in the past had suggested that the genus contains as few as 3 to 6 species,Weber, N.S. (1988). In A Morel Hunter's Companion, pp. 111-167. Two Peninsula Press: Lansing. while others recognised as many as 34. Efforts to clarify the situation and re-evaluate old classical names (such as ''
Morchella elata ''Morchella elata'' is a species of fungus in the family Morchellaceae. It is one of many related species commonly known as black morels, and until 2012 the name ''M. elata'' was broadly applied to black morels throughout the globe. Like most ...
'' and others) in accordance to current phylogenetic data have been challenging, due to vague or ambiguous original descriptions and loss of
holotype A holotype (Latin: ''holotypus'') is a single physical example (or illustration) of an organism used when the species (or lower-ranked taxon) was formally described. It is either the single such physical example (or illustration) or one of s ...
material. In 2012, the simultaneous description of several new taxa from Europe by Clowez and North America by Kuo and colleagues resulted in several
synonyms A synonym is a word, morpheme, or phrase that means precisely or nearly the same as another word, morpheme, or phrase in a given language. For example, in the English language, the words ''begin'', ''start'', ''commence'', and ''initiate'' are a ...
further complicating matters, until a transatlantic study by Richard and colleagues resolved many of these issues in 2014. The genus is currently undergoing extensive re-evaluation with regard to the taxonomic status of several species.


Early taxonomic history

''Morchella'' Dill. ex Pers. : Fr. was typified by
Christiaan Hendrik Persoon Christiaan Hendrik Persoon (31 December 1761 – 16 November 1836) was a Cape Colony mycologist who is recognized as one of the founders of mycology, mycological Taxonomy (biology), taxonomy. Early life Persoon was born in Cape Colony at ...
in 1794, with ''
Morchella esculenta ''Morchella esculenta'' (commonly known as common morel, morel, yellow morel, true morel, morel mushroom, and sponge morel) is a species of fungus in the family Morchellaceae of the Ascomycota. It is one of the most readily recognized of all th ...
'' designated as the
type species In International_Code_of_Zoological_Nomenclature, zoological nomenclature, a type species (''species typica'') is the species name with which the name of a genus or subgenus is considered to be permanently taxonomically associated, i.e., the spe ...
for the genus. Among early pioneers who took an interest in the genus, were mycologists
Julius Vincenz von Krombholz Julius Vincenz von Krombholz (19 December 1782 – 1 November 1843) was a physician and mycologist born in Oberpolitz (today Horní Police, Czech Republic), northern Bohemia. He studied medicine at the Charles University in Prague, University of P ...
and Émile Boudier, who, in 1834 and 1897 respectively, published several species and varieties, accompanied by meticulously illustrated iconographic plates. The seminal taxon ''
Morchella elata ''Morchella elata'' is a species of fungus in the family Morchellaceae. It is one of many related species commonly known as black morels, and until 2012 the name ''M. elata'' was broadly applied to black morels throughout the globe. Like most ...
'', whose true identity still remains unresolved, was described by
Elias Fries Elias Magnus Fries (15 August 1794 – 8 February 1878) was a Swedish mycologist and botanist. He is sometimes called the "Linnaeus of Mycology". In his works he described and assigned botanical names to hundreds of fungus and lichen sp ...
in 1822, from a fir forest in
Sweden Sweden, formally the Kingdom of Sweden, is a Nordic countries, Nordic country located on the Scandinavian Peninsula in Northern Europe. It borders Norway to the west and north, and Finland to the east. At , Sweden is the largest Nordic count ...
. Other classical, early-proposed names include ''
Morchella deliciosa ''Morchella deliciosa'' is a species of edible fungus in the family Morchellaceae. It was first described scientifically by Elias Magnus Fries Elias Magnus Fries (15 August 1794 – 8 February 1878) was a Swedish mycologist and botanis ...
'', also described by Fries in 1822, ''
Morchella semilibera ''Morchella semilibera'', commonly called the half-free morel, is a species of fungus in the family Morchellaceae. It is native to Eurasia and is edible. Taxonomy DNA analysis has shown that the half-free morels, which appear nearly identical ...
'', the half-free morel, originally described by
de Candolle Augustin Pyramus (or Pyrame) de Candolle (, , ; 4 February 17789 September 1841) was a Swiss people, Swiss botany, botanist. René Louiche Desfontaines launched de Candolle's botanical career by recommending him at a herbarium. Within a couple ...
and sanctioned by Fries in 1822, ''
Morchella vulgaris ''Morchella vulgaris'' is a widespread fungus of the family Morchellaceae (Ascomycota). It was originally described in 1801 as a form of the common yellow morel (''Morchella esculenta'') by mycologist Christiaan Hendrik Persoon, but was later rec ...
'', which was recombined by Samuel Gray as a distinct species in 1821 following a ''forma'' of ''M. esculenta'' previously proposed by
Persoon Persoon is a surname. Notable people with the surname include: * Christiaan Hendrik Persoon (1761–1836), German mycologist * Delfine Persoon (born 1985), Belgian boxer * Sigrid Persoon Sigrid Persoon (born 12 March 1983) is a Belgian former g ...
, and ''
Morchella angusticeps ''Morchella angusticeps'' is a species of fungus in the family Morchellaceae native to eastern North America. Described by Charles Horton Peck Charles Horton Peck (March 30, 1833 – July 11, 1917) was an American mycologist of the 19th and ...
'', a large-spored species described by American mycologist Charles Peck in 1887. ''
Morchella purpurascens ''Morchella purpurascens'', the purple morel, is an ascomycete fungus in the family ''Morchellaceae''. It was first described as a variety of ''Morchella elata'' by Emile Boudier in 1897, based on a plate previously illustrated by Julius Vincen ...
'', the purple morel, was first described by Boudier as a variety of ''M. elata'' in 1897 based on an 1834 plate by Krombholz, and was recombined as a distinct species in 1985 by Emile Jacquetant. ''
Morchella eximia ''Morchella eximia'' is a globally-occurring fungus in the family ''Morchellaceae'' (Ascomycota), first described by Émile Boudier in 1910.Boudier E. (1910). Icones mycologicae ou iconographie des champignons de France principalement discomycè ...
'', a globally-occurring fire-associated species was also described by Boudier in 1910. The old, widely applied name ''
Morchella conica ''Morchella conica'' is an old binomial name previously applied to species of fungi in the family Morchellaceae. It is one of three scientific names that had been commonly used to describe black morels, the others being '' M. angusticeps'' ...
'', featuring in many field guides and literature across several countries, has been shown by Richard and colleagues to be
illegitimate Legitimacy, in traditional Western common law, is the status of a child born to parents who are legally married to each other, and of a child conceived before the parents obtain a legal divorce. Conversely, ''illegitimacy'', also known as ''b ...
.


Classification

About 80 species of ''Morchella'' were described until the turn of the 21st century (per the ''
Index Fungorum ''Index Fungorum'' is an international project to index all formal names (scientific names) in the fungus kingdom. As of 2015, the project is based at the Royal Botanic Gardens, Kew, one of three partners along with Landcare Research and th ...
''), a number of which were later shown to be illegitimate or synonyms. As molecular tools became widely available in the new millennium, a revived interest in the genus commenced and several new species were proposed. In 2008 Kuo described ''
Morchella tomentosa ''Morchella tomentosa'', commonly called the gray, fuzzy foot, or black foot morel, is a species of fungus in the family Morchellaceae. ''M. tomentosa'' is a fire-associated species described from western North America, formally described as ...
'' from burned coniferous forests in western North America. In 2010 IÅŸiloÄŸlu and colleagues described '' Morchella anatolica'', a basal species from
Turkey Turkey, officially the Republic of Türkiye, is a country mainly located in Anatolia in West Asia, with a relatively small part called East Thrace in Southeast Europe. It borders the Black Sea to the north; Georgia (country), Georgia, Armen ...
later shown to be sister to ''
Morchella rufobrunnea ''Morchella rufobrunnea'', commonly known as the blushing morel, is a species of ascomycete fungus in the family Morchellaceae. A choice edible species, the fungus was described as new to science in 1998 by mycologists Gastón Guzmán and Fide ...
''. A study by Clowez described over 20 new species in 2012, while later in the same year, another study by Kuo and colleagues described 19 species from
North America North America is a continent in the Northern Hemisphere, Northern and Western Hemisphere, Western hemispheres. North America is bordered to the north by the Arctic Ocean, to the east by the Atlantic Ocean, to the southeast by South Ameri ...
. However, several of these newly proposed names later turned out to be synonyms. An extensive taxonomical and nomenclatural revision of the genus provided by Richard and colleagues in 2014, applied names to 30 of the genealogical lineages recognized so far and clarified several synonymities. Also in 2014, Elliott and colleagues described ''
Morchella australiana ''Morchella australiana'' is a species of ascomycete fungus in the family Morchellaceae. Described as new to science in 2014, it is found in New South Wales, South Australia, Tasmania and Victoria, Australia. The type locality was in the temper ...
'' from
sclerophyll Sclerophyll is a type of vegetation that is adapted to long periods of dryness and heat. The plants feature hard leaves, short Internode (botany), internodes (the distance between leaves along the stem) and leaf orientation which is parallel or ...
forests in
Australia Australia, officially the Commonwealth of Australia, is a country comprising mainland Australia, the mainland of the Australia (continent), Australian continent, the island of Tasmania and list of islands of Australia, numerous smaller isl ...
, while Clowez and colleagues described '' Morchella fluvialis'' from riparian forests in
Spain Spain, or the Kingdom of Spain, is a country in Southern Europe, Southern and Western Europe with territories in North Africa. Featuring the Punta de Tarifa, southernmost point of continental Europe, it is the largest country in Southern Eur ...
. In 2015, Loizides and colleagues clarified the taxonomy of ''
Morchella tridentina ''Morchella tridentina'' is a cosmopolitan species of ascomycete fungus in the family Morchellaceae. Commonly referred to as the mountain blond or western blond morel in North America, it produces conical, grey to buff ascocarp, fruit bodies tha ...
'', a cosmopolitan species described under many names, and recombined ''
Morchella kakiicolor ''Morchella kakiicolor'' is a species of fungus in the family ''Morchellaceae'' (Ascomycota). It was originally proposed as a form of ''Morchella quercus-ilicis'' in a 2012 study by Philippe Clowez, but was later re-combined as an autonomous spec ...
'' as a distinct species. Later in the same year, Clowez and colleagues described '' Morchella palazonii'' from Spain, while Voitk and colleagues described ''
Morchella laurentiana ''Morchella laurentiana'' is a species of fungus in the family Morchellaceae described as new to science in 2016. It is known only from the Saint Lawrence River basin in the Canadian province of Newfoundland and Labrador. It is in the ''Morchell ...
'' from
Canada Canada is a country in North America. Its Provinces and territories of Canada, ten provinces and three territories extend from the Atlantic Ocean to the Pacific Ocean and northward into the Arctic Ocean, making it the world's List of coun ...
and '' Morchella eohespera'', a cosmopolitan species present in several continents. In an extensive phylogenetic and morphological study from
Cyprus Cyprus (), officially the Republic of Cyprus, is an island country in the eastern Mediterranean Sea. Situated in West Asia, its cultural identity and geopolitical orientation are overwhelmingly Southeast European. Cyprus is the List of isl ...
in 2016, Loizides and colleagues added two more
Mediterranean The Mediterranean Sea ( ) is a sea connected to the Atlantic Ocean, surrounded by the Mediterranean basin and almost completely enclosed by land: on the east by the Levant in West Asia, on the north by Anatolia in West Asia and Southern ...
species, '' Morchella arbutiphila'' and '' Morchella disparilis'', and resurrected ''
Morchella dunensis ''Morchella dunensis'', the morel of the dunes, is a species of fungus in the family ''Morchellaceae'' (Ascomycota). It was first described as a form of ''Morchella esculenta'' by Castañera and colleagues in 1996, but was later recombined as a d ...
'' as an autonomous species. In the same year, Taşkın and colleagues described four of the previously unnamed phylospecies from Turkey: ''
Morchella conifericola ''Morchella'', the true morels, is a genus of edible sac fungi closely related to anatomically simpler cup fungi in the order Pezizales ( division Ascomycota). These distinctive fungi have a honeycomb appearance due to the network of ridges wi ...
'', ''
Morchella feekensis ''Morchella'', the true morels, is a genus of edible sac fungi closely related to anatomically simpler cup fungi in the order Pezizales ( division Ascomycota). These distinctive fungi have a honeycomb appearance due to the network of ridges wi ...
'', ''
Morchella magnispora ''Morchella'', the true morels, is a genus of edible sac fungi closely related to anatomically simpler cup fungi in the order Pezizales ( division Ascomycota). These distinctive fungi have a honeycomb appearance due to the network of ridges wi ...
'' and ''
Morchella mediteterraneensis ''Morchella'', the true morels, is a genus of edible sac fungi closely related to anatomically simpler cup fungi in the order Pezizales ( division Ascomycota). These distinctive fungi have a honeycomb appearance due to the network of ridges w ...
''.


Section ''Rufobrunnea''

*'' Morchella anatolica'' **synonym: ''
Morchella lanceolata ''Morchella anatolica'' is a rare species of ascomycete fungus in the family Morchellaceae. It was described as new to science in 2010 from southwest Anatolia Anatolia (), also known as Asia Minor, is a peninsula in West Asia that makes up ...
'' *''
Morchella rufobrunnea ''Morchella rufobrunnea'', commonly known as the blushing morel, is a species of ascomycete fungus in the family Morchellaceae. A choice edible species, the fungus was described as new to science in 1998 by mycologists Gastón Guzmán and Fide ...
''


Section ''Morchella''

*'' Morchella americana'' **synonyms: ''
Morchella californica ''Morchella americana'' (also called the yellow morel) is a North American species of fungus in the family Morchellaceae. Taxonomy The species was described as new to science in 2012. In 2014, Richard et al. clarified its taxonomic status, retai ...
'', ''
Morchella claviformis ''Morchella americana'' (also called the yellow morel) is a North American species of fungus in the family Morchellaceae. Taxonomy The species was described as new to science in 2012. In 2014, Richard et al. clarified its taxonomic status, retai ...
'', '' Morchella esculentoides'', '' Morchella populina'' *'' Morchella castaneae'' **synonyms: '' Morchella brunneorosea'', ''
Morchella brunneorosea var. sordida ''Morchella'', the true morels, is a genus of edible sac fungi closely related to anatomically simpler cup fungi in the order Pezizales ( division Ascomycota). These distinctive fungi have a honeycomb appearance due to the network of ridges wi ...
'' *'' Morchella diminutiva'' *''
Morchella dunensis ''Morchella dunensis'', the morel of the dunes, is a species of fungus in the family ''Morchellaceae'' (Ascomycota). It was first described as a form of ''Morchella esculenta'' by Castañera and colleagues in 1996, but was later recombined as a d ...
'' **synonyms: ''
Morchella esculenta f. dunensis ''Morchella dunensis'', the morel of the dunes, is a species of fungus in the family ''Morchellaceae'' (Ascomycota). It was first described as a Form (botany), form of ''Morchella esculenta'' by Castañera and colleagues in 1996, but was later re ...
'', ''
Morchella andalusiae ''Morchella'', the true morels, is a genus of edible sac fungi closely related to anatomically simpler cup fungi in the order Pezizales ( division Ascomycota). These distinctive fungi have a honeycomb appearance due to the network of ridges wi ...
'' *''
Morchella esculenta ''Morchella esculenta'' (commonly known as common morel, morel, yellow morel, true morel, morel mushroom, and sponge morel) is a species of fungus in the family Morchellaceae of the Ascomycota. It is one of the most readily recognized of all th ...
'' **synonyms: ''
Morchella pseudoumbrina ''Morchella esculenta'' (commonly known as common morel, morel, yellow morel, true morel, morel mushroom, and sponge morel) is a species of fungus in the family Morchellaceae of the Ascomycota. It is one of the most readily recognized of all the ...
'', ''
Morchella pseudoviridis ''Morchella esculenta'' (commonly known as common morel, morel, yellow morel, true morel, morel mushroom, and sponge morel) is a species of fungus in the family Morchellaceae of the Ascomycota. It is one of the most readily recognized of all the ...
'' *'' Morchella fluvialis'' *'' Morchella galilaea'' *'' Morchella palazonii'' *''
Morchella prava ''Morchella prava'' is a species of fungus in the family Morchellaceae The Morchellaceae are a family of ascomycete fungi in the order Pezizales. According to a standard reference work, the family has contained at least 49 species distributed ...
'' *''
Morchella sceptriformis ''Morchella sceptriformis'' is a species of fungus in the family Morchellaceae (Ascomycota). It was described as new to science in a 2012 study by Clowez, and corresponds to phylogenetic lineage ''Mes''-3. ''Morchella virginiana'', described lat ...
'' **synonym: ''
Morchella virginiana ''Morchella sceptriformis'' is a species of fungus in the family Morchellaceae (Ascomycota). It was described as new to science in a 2012 study by Clowez, and corresponds to phylogenetic lineage ''Mes''-3. ''Morchella virginiana'', described lat ...
'' *'' Morchella steppicola'' *'' Morchella ulmaria'' **synonym: ''Morchella cryptica'' *''
Morchella vulgaris ''Morchella vulgaris'' is a widespread fungus of the family Morchellaceae (Ascomycota). It was originally described in 1801 as a form of the common yellow morel (''Morchella esculenta'') by mycologist Christiaan Hendrik Persoon, but was later rec ...
'' **synonyms: ''Morchella acerina'', ''Morchella anthracina'', ''Morchella lepida'', ''Morchella robiniae'', ''Morchella spongiola''


Section ''Distantes''

*''
Morchella angusticeps ''Morchella angusticeps'' is a species of fungus in the family Morchellaceae native to eastern North America. Described by Charles Horton Peck Charles Horton Peck (March 30, 1833 – July 11, 1917) was an American mycologist of the 19th and ...
'' *'' Morchella arbutiphila'' *''
Morchella australiana ''Morchella australiana'' is a species of ascomycete fungus in the family Morchellaceae. Described as new to science in 2014, it is found in New South Wales, South Australia, Tasmania and Victoria, Australia. The type locality was in the temper ...
'' *''Morchella brunnea'' *''
Morchella conifericola ''Morchella'', the true morels, is a genus of edible sac fungi closely related to anatomically simpler cup fungi in the order Pezizales ( division Ascomycota). These distinctive fungi have a honeycomb appearance due to the network of ridges wi ...
'' *''
Morchella deliciosa ''Morchella deliciosa'' is a species of edible fungus in the family Morchellaceae. It was first described scientifically by Elias Magnus Fries Elias Magnus Fries (15 August 1794 – 8 February 1878) was a Swedish mycologist and botanis ...
'' **synonym: ''
Morchella conica ''Morchella conica'' is an old binomial name previously applied to species of fungi in the family Morchellaceae. It is one of three scientific names that had been commonly used to describe black morels, the others being '' M. angusticeps'' ...
'' *'' Morchella disparilis'' *''Morchella dunalii'' **synonym: ''Morchella fallax'' *''
Morchella elata ''Morchella elata'' is a species of fungus in the family Morchellaceae. It is one of many related species commonly known as black morels, and until 2012 the name ''M. elata'' was broadly applied to black morels throughout the globe. Like most ...
'' *'' Morchella eohespera'' *''
Morchella eximia ''Morchella eximia'' is a globally-occurring fungus in the family ''Morchellaceae'' (Ascomycota), first described by Émile Boudier in 1910.Boudier E. (1910). Icones mycologicae ou iconographie des champignons de France principalement discomycè ...
'' **synonyms: ''Morchella anthracophila'', ''Morchella carbonaria'', ''Morchella septimelata'' *''Morchella eximioides'' *''Morchella exuberans'' **synonym: ''Morchella capitata'' *''
Morchella feekensis ''Morchella'', the true morels, is a genus of edible sac fungi closely related to anatomically simpler cup fungi in the order Pezizales ( division Ascomycota). These distinctive fungi have a honeycomb appearance due to the network of ridges wi ...
'' *''Morchella iberica'' *''Morchella importuna'' *''
Morchella kakiicolor ''Morchella kakiicolor'' is a species of fungus in the family ''Morchellaceae'' (Ascomycota). It was originally proposed as a form of ''Morchella quercus-ilicis'' in a 2012 study by Philippe Clowez, but was later re-combined as an autonomous spec ...
'' **synonym: ''Morchella quercus-ilicis f. kakiicolor'' *''
Morchella laurentiana ''Morchella laurentiana'' is a species of fungus in the family Morchellaceae described as new to science in 2016. It is known only from the Saint Lawrence River basin in the Canadian province of Newfoundland and Labrador. It is in the ''Morchell ...
'' *''
Morchella magnispora ''Morchella'', the true morels, is a genus of edible sac fungi closely related to anatomically simpler cup fungi in the order Pezizales ( division Ascomycota). These distinctive fungi have a honeycomb appearance due to the network of ridges wi ...
'' *''Morchella mediterraneensis'' *''Morchella populiphila'' *''Morchella pulchella'' *''Morchella punctipes'' *''
Morchella purpurascens ''Morchella purpurascens'', the purple morel, is an ascomycete fungus in the family ''Morchellaceae''. It was first described as a variety of ''Morchella elata'' by Emile Boudier in 1897, based on a plate previously illustrated by Julius Vincen ...
'' **synonyms: ''Morchella elata var. purpurascens'', ''
Morchella conica ''Morchella conica'' is an old binomial name previously applied to species of fungi in the family Morchellaceae. It is one of three scientific names that had been commonly used to describe black morels, the others being '' M. angusticeps'' ...
'', ''Morchella conica var. purpurascens'', ''Morchella conica var. crassa'' *''
Morchella semilibera ''Morchella semilibera'', commonly called the half-free morel, is a species of fungus in the family Morchellaceae. It is native to Eurasia and is edible. Taxonomy DNA analysis has shown that the half-free morels, which appear nearly identical ...
'' **synonyms: ''Morchella gigas'', ''Morchella gigas var. tintinnabulum'', ''Morchella hybrida'', ''Morchella undosa'', ''Morchella varisiensis'', ''Morchella esculenta var. crassipes'', ''Phallus gigas'', ''Eromitra gigas'', ''Phallus undosus'', ''Phallus crassipes'', ''Mitrophora hybrida'', ''Mitrophora hybrida var. crassipes'', ''Ptychoverpa gigas'', ''Helvella hybrida'' *''Morchella septentrionalis'' *''Morchella sextelata'' *''Morchella snyderi'' *''
Morchella tomentosa ''Morchella tomentosa'', commonly called the gray, fuzzy foot, or black foot morel, is a species of fungus in the family Morchellaceae. ''M. tomentosa'' is a fire-associated species described from western North America, formally described as ...
'' *''
Morchella tridentina ''Morchella tridentina'' is a cosmopolitan species of ascomycete fungus in the family Morchellaceae. Commonly referred to as the mountain blond or western blond morel in North America, it produces conical, grey to buff ascocarp, fruit bodies tha ...
'' **synonyms: ''Morchella quercus-ilicis'', ''Morchella frustrata'', ''Morchella elatoides'', ''Morchella elatoides var. elagans'', ''Morchella conica var. pseudoeximia''


Unresolved classification

*''Morchella anteridiformis'' *''Morchella apicata'' *''Morchella bicostata'' *''Morchella conicopapyracea'' *''Morchella crassipes'' *''Morchella deqinensis'' *''Morchella distans'' *''Morchella guatemalensis'' *''Morchella herediana'' *''Morchella hetieri'' *''Morchella hortensis'' *''Morchella hotsonii'' *''Morchella hungarica'' *''Morchella inamoena'' *''Morchella intermedia'' *''Morchella meiliensis'' *''Morchella miyabeana'' *''Morchella neuwirthii'' *''Morchella norvegiensis'' *''Morchella patagonica'' *''Morchella patula'' *''Morchella pragensis'' *''Morchella procera'' *''Morchella pseudovulgaris'' *''Morchella rielana'' *''Morchella rigida'' *''Morchella rigidoides'' *''Morchella smithiana'' *''Morchella sulcata'' *''Morchella tasmanica'' *''Morchella tatari'' *''Morchella tibetica'' *''Morchella umbrina'' *''Morchella umbrinovelutipes'' *''Morchella vaporaria''


Phylogeny

Early
phylogenetic In biology, phylogenetics () is the study of the evolutionary history of life using observable characteristics of organisms (or genes), which is known as phylogenetic inference. It infers the relationship among organisms based on empirical dat ...
analyses supported the hypothesis that the genus comprises only a few species with considerable phenotype, phenotypic variation. Subsequent multigenic DNA studies, however, have revealed more than a dozen genealogically distinct species in
North America North America is a continent in the Northern Hemisphere, Northern and Western Hemisphere, Western hemispheres. North America is bordered to the north by the Arctic Ocean, to the east by the Atlantic Ocean, to the southeast by South Ameri ...
and at least as many in Europe. DNA studies revealed three discrete clades, or genetic groups, consisting of the "white morels" (''
Morchella rufobrunnea ''Morchella rufobrunnea'', commonly known as the blushing morel, is a species of ascomycete fungus in the family Morchellaceae. A choice edible species, the fungus was described as new to science in 1998 by mycologists Gastón Guzmán and Fide ...
'' and ''Morchella anatolica, M. anatolica''), the "yellow morels" (''Morchella esculenta, M. esculenta'' and others), and the "black morels" (''Morchella elata, M. elata'' and others). The fire-associated species ''
Morchella tomentosa ''Morchella tomentosa'', commonly called the gray, fuzzy foot, or black foot morel, is a species of fungus in the family Morchellaceae. ''M. tomentosa'' is a fire-associated species described from western North America, formally described as ...
'', commonly known as the "gray morel", is distinct for its tomentose, fine hairs on the cap ridges and sclerotium, sclerotia-like underground structures, and may also deserve its own clade based on DNA evidence. Within the yellow and black clades, there are dozens of distinct species, many endemism, endemic to individual continents or regions. This species-rich view is supported by studies in Western Europe, Turkey,
Cyprus Cyprus (), officially the Republic of Cyprus, is an island country in the eastern Mediterranean Sea. Situated in West Asia, its cultural identity and geopolitical orientation are overwhelmingly Southeast European. Cyprus is the List of isl ...
, Israel,
China China, officially the People's Republic of China (PRC), is a country in East Asia. With population of China, a population exceeding 1.4 billion, it is the list of countries by population (United Nations), second-most populous country after ...
, Patagonia, and the
Himalayas The Himalayas, or Himalaya ( ), is a mountain range in Asia, separating the plains of the Indian subcontinent from the Tibetan Plateau. The range has some of the Earth's highest peaks, including the highest, Mount Everest. More than list of h ...
. Early ancestral reconstruction tests by O'Donnell and collaborators postulated a Western United States, western North American origin of morels and the genus was estimated to have diverged from its closest genealogical relatives ''
Verpa ''Verpa'' is a genus of ascomycete fungi related to the morels. Resembling the latter genus in edibility and form, the common name early morels is popular. There are five species in the widespread genus. Taxonomy Analysis of the ribosomal DNA o ...
'' and ''Disciotis'' in the early Cretaceous, approximately 129 million years ago (Mya). This date was later revised by Du and collaborators, placing the divergence of the genus in the late Jurassic, approximately 154 Mya. However, neither of these reconstructions had included '' Morchella anatolica'' in the analyses, whose phylogenetic placement remained at the time unresolved. Following genetic testing of isotype collection of ''M. anatolica'' by TaÅŸkın and colleagues, this species was shown to nest in the ancestral /Rufobrunnea clade, together with the transcontinental ''M. rufobrunnea''. This cast doubts over the accuracy of the original reconstructions, since both species of the ancestral /Rufobrunnea clade are present in the Mediterranean, while ''M. anatolica'' is altogether absent from North America. Updated ancestral area reconstructions by Loizides and colleagues using an expanded 79-species data set, have in 2021 refuted the previous hypothesis and designated the Mediterranean basin as the most probable place of origin of morels.


Distribution and habitat

Morels can be found in the temperate
Northern Hemisphere The Northern Hemisphere is the half of Earth that is north of the equator. For other planets in the Solar System, north is defined by humans as being in the same celestial sphere, celestial hemisphere relative to the invariable plane of the Solar ...
, in particular North America, Turkey, China, the Himalayas,
India India, officially the Republic of India, is a country in South Asia. It is the List of countries and dependencies by area, seventh-largest country by area; the List of countries by population (United Nations), most populous country since ...
, and
Pakistan Pakistan, officially the Islamic Republic of Pakistan, is a country in South Asia. It is the List of countries and dependencies by population, fifth-most populous country, with a population of over 241.5 million, having the Islam by country# ...
. Yellow morels (''
Morchella esculenta ''Morchella esculenta'' (commonly known as common morel, morel, yellow morel, true morel, morel mushroom, and sponge morel) is a species of fungus in the family Morchellaceae of the Ascomycota. It is one of the most readily recognized of all th ...
'' and related species) are more commonly found under deciduous trees rather than conifers, while black morels (''Morchella elata, M. elata'' and related species) are mostly found in coniferous forests, disturbed ground and recently burned areas. '' Morchella galilaea'', and occasionally ''M. rufobrunnea'', appear to fruit in the autumn or winter months rather than spring, which is the typical fruiting season for morels. In the American Pacific Northwest, they can be found from April to August. Efforts to Fungiculture, cultivate morels at a large scale have rarely been successful and the commercial morel industry relies on the harvest of wild mushrooms.


Transcontinental species

Although many species within ''Morchella'' exhibit continental
endemism Endemism is the state of a species being found only in a single defined geographic location, such as an island, state, nation, country or other defined zone; organisms that are indigenous to a place are not endemic to it if they are also foun ...
and provincialism, several species have been phylogenetically shown to be present in more than one continent. So far, the list of transcontinental species includes ''Morchella americana, M. americana'', ''Morchella eohespera, M. eohespera'', ''Morchella eximia, M. eximia'', ''Morchella exuberans, M. exuberans'', ''Morchella galilaea, M. galilaea'', ''Morchella importuna, M. importuna'', ''Morchella populiphila, M. populiphila'', ''Morchella pulchella, M. pulchella'', ''Morchella rufobrunnea, M. rufobrunnea'', ''Morchella semilibera, M. semilibera'', ''Morchella sextelata, M. sextelata'', ''Morchella steppicola, M. steppicola'', and ''Morchella tridentina, M. tridentina''. The reasons behind the widespread, cosmopolitan distribution of these species, are still puzzling. Some authors have hypothesized that such transcontinental occurrences are the result of accidental anthropogenic introduced species, introductions, but this view has been disputed by others, who suggested an old and natural distribution, at least for some of these species which appear to be linked to indigenous flora. Long-distance spore dispersal has also been suggested as a possible biological dispersal, dispersal mechanism for some species, especially those belonging to fire-adapted Lineage (evolution), lineages. It has been suggested that the widespread but disjunct distribution of some morel species, especially early diverging lineages like ''Morchella rufobrunnea, M. rufobrunnea'' and ''Morchella tridentina, M. tridentina'', may be the result of climatic refugia from the Quaternary glaciation.


Ecology

The ecology of ''Morchella'' species is not well understood. Many species appear to form symbiotic relationship, symbiotic or endophytic relationships with trees, while others appear to act as saprotrophs. Tree species associated with ''Morchella'' vary greatly depending on the individual species, continent, or region. Trees commonly associated with morels in Europe and across the
Mediterranean The Mediterranean Sea ( ) is a sea connected to the Atlantic Ocean, surrounded by the Mediterranean basin and almost completely enclosed by land: on the east by the Levant in West Asia, on the north by Anatolia in West Asia and Southern ...
include ''Abies'' (fir), ''Pinus'' (pine), ''Populus'' (poplar), ''Ulmus'' (elm), ''Quercus'' (oak), ''Arbutus'' (strawberry trees), ''Castanea (plant), Castanea'' (chestnut), ''Alnus'' (alder), ''Olea'' (olive trees), ''Malus'' (apple trees), and ''Fraxinus'' (ash). In western North America morels are often found in coniferous forests, including species of ''Pinus'' (pine), ''Abies'' (fir), ''Larix'' (larch), and ''Pseudotsuga'' (Douglas-fir), as well as in ''Populus'' (cottonwood) riparian zone, riparian forests. Deciduous trees commonly associated with morels in the northern hemisphere include ''Fraxinus'' (ash), ''Platanus'' (sycamore), ''Liriodendron'' (tulip tree), dead and dying elms, Populus sect. Aigeiros, cottonwoods, and old apple trees (remnants of orchards). Due to their springtime phenology (March–May), morels are hardly ever found in the vicinity of common poisonous mushrooms such as the death cap (''Amanita phalloides''), the sulphur tuft (''Hypholoma fasciculare''), or the fly agaric (''Amanita muscaria''). They can, however, occur alongside false morels (''
Gyromitra ''Gyromitra'' () is a genus of about 18 species of ascomycete fungi. They are a false morel - a frequently toxic mushroom that can be mistaken for edible mushrooms of the genus ''Morchella'' (morels). Taxonomy The name ''Gyromitra'' comes from ...
'' and ''
Verpa ''Verpa'' is a genus of ascomycete fungi related to the morels. Resembling the latter genus in edibility and form, the common name early morels is popular. There are five species in the widespread genus. Taxonomy Analysis of the ribosomal DNA o ...
'' species) and elfin saddles (''Helvella'' species), which also appear in spring.


Association with wildfire

Certain ''Morchella'' species (''Morchella eximia, M. eximia'', ''Morchella importuna, M. importuna'', ''Morchella tomentosa, M. tomentosa'' and others) exhibit a Pyrophyte, pyrophilic behaviour and may grow abundantly in forests which have been recently burned by a forest fire, fire. Moderate-intensity fires are reported to produce higher abundances of morels than low- or high-intensity fires. This is caused by the soil becoming more alkaline as the result of wood ash combining with water and being absorbed into the soil which triggers the morels to fruit. Alkali soil, Alkaline soil conditions which trigger fruiting have been observed and exploited with small-scale commercial cultivation of morels.Archived a
Ghostarchive
and th
Wayback Machine
Where fire suppression is practiced, morels often grow in small numbers in the same spot, year after year. If these areas are overrun by wildfire they often produce a bumper crop of black morels the following spring. Commercial pickers and buyers in North America target recently burned areas for this reason. These spots may be closely guarded by mushroom pickers, as morels are widely regarded as a delicacy and often a cash crop.


Cultivation

Due to the mushroom's prized fruit bodies, several attempts have been made to grow the fungus in culture. In 1901, Repin reported successfully obtaining fruit bodies in a cave in which cultures had been established in flower pots nine years previously in 1892. More recently, small-scale commercial growers have had success growing morels by using partially shaded rows of mulched wood. The rows of mulch piles are inoculated with morel mushroom spores in a solution of water and molasses which are poured over the piles of mulch and then they are allowed to grow undisturbed for several weeks. A solution of wood ashes mixed in water and diluted is subsequently poured over the rows of wood mulch which triggers fruiting of the morels. Morels are known to appear after fires and the alkalinity produced by wood ash mixed with water initiate fruit body formation for most species of morels. In 2021 it was announced that indoor cultivation of black morels had been successfully achieved after decades of research and experimentation with methods by The Danish Morel Project. The project has been able to cultivate 20 lbs of morels per square yard or around 10 kg per square metre with cost estimates expected to be similar to producing white button mushrooms (''Agaricus bisporus''). Previous attempts at cultivation had managed to produce Sclerotium, sclerotia but encountered issues in getting them to reliably fruit. One of the breakthroughs with this project was growing them in a climate controlled environment in conjunction with grass which is involved in stimulating fruiting in the morel mycelium. Cultivation in this manner has been noted to produce superior morels for culinary uses since they can be assured to be insect, slug and dirt free and therefore do not need to be washed and cleaned like foraged morels. Since washing morels can negatively impact the texture, reliable cultivation may result in more versatility with this ingredient in the kitchen as well as making the delicacy more affordable and accessible.


Toxicity

The consumption of ''Morchella'' species can have adverse effects. In 2023, a Montana sushi restaurant serving them was linked to 51 people who experienced gastrointestinal illness, with two reported deaths and three other hospitalizations. The consumption of raw morels in particular is advised against. An unknown toxin can be neutralized via cooking. Additionally, cooked morels can reportedly cause upset stomach, upset stomachs when consumed with alcohol. When eating this fungus for the first time, it is advised to consume a small amount to minimize any allergic reaction. As with all fungi, morels for consumption must be clean and free of decay. Morels growing in old apple orchards previously treated with the deprecated insecticide lead arsenate may bioaccumulation, accumulate levels of toxic lead and arsenic that are unsuitable for human consumption.


Uses

Morels, "almost universally associated with spring," can be found in many habitats. Morel may be more likely to fruit during a period of increasing heat following a chilly period, a preference which is credited for their abundance in areas with cold winters. Black morels (''
Morchella elata ''Morchella elata'' is a species of fungus in the family Morchellaceae. It is one of many related species commonly known as black morels, and until 2012 the name ''M. elata'' was broadly applied to black morels throughout the globe. Like most ...
'') are often found on land that has been disturbed by logging burning.


Nutrition

Raw morel mushrooms are 90% water, 5% carbohydrates, 3% protein (nutrient), protein, and 1% fat. A 100 gram reference amount supplies 31 calories, and is a rich source of iron (94% of the Daily Value, DV), manganese, phosphorus, zinc, and vitamin D (34% DV, if having been exposed to sunlight or artificial ultraviolet light). Raw morels contain moderate levels of several B vitamins (table).


Gastronomical value and culinary uses

They have been called "prized delicacies...they are so esteemed in Europe that people used to set fire to their own forests in hopes of eliciting a bountiful morel crop the next spring!" Morels are a feature of many cuisines, including French cuisine#Provence-Alpes-Côte d'Azur, Provençal. Their flavor is prized by chefs worldwide, with recipes and preparation methods designed to highlight and preserve it. As with most Edible mushroom, edible fungi, they are best when collected or bought fresh. They are sometimes added to meat and poultry dishes and soups, and can be used as pasta fillings. As morels are known to contain thermolabile toxins, they must always be cooked before eating. Morels can be preserved in several ways: They can be flash freezing, 'flash frozen' by simply running under cold water or putting them in a bucket to soak for a few minutes, then spread on a Sheet pan, baking tray and placed into a freezer. After freezing, they keep very well with the frozen glaze for a long time in airtight containers. However, when thawed they can sometimes turn slightly mushy, so they are best frozen after steaming or frying. Due to their natural porosity, morels may contain trace amounts of soil which cannot be easily washed out. Any visible soil should be removed with a brush, after cutting the body in half lengthwise, if needed. Mushroom hunters sometimes recommend soaking morels in a bowl of salt water briefly prior to cooking, although many chefs would disagree. Food drying, Drying is a popular and effective method for long-term storage, and morels are widely available commercially in this form. Any Larva, insect larvae which might be present in the fruit bodies usually drop out during the drying process.Wild About Mushrooms: Morels
Mssf.org. Retrieved on 2012-04-17.
Dried morels can then be reconstituted by soaking for 10–20 minutes in warm water or milk, and the soaking liquid can be used as stock. The flavor of morels is not just appreciated by humans; in Yellowstone National Park, black morels are also known to be consumed by grizzly bears (''Ursus arctos horribilis'').


In popular culture

Morel hunting is a common springtime activity. Mushroom collectors may carry a mesh collecting bag, so the spores can scatter as one carries the harvest. Every spring, hundreds of morel enthusiasts gather in Boyne City, Michigan for the National Morel Mushroom Festival, a century-old event. As one observer stated, "if there is a modern, North American reenactment of Geoffrey Chaucer's ''Canterbury Tales'' this is it." Other festivals and hunting competitions in
North America North America is a continent in the Northern Hemisphere, Northern and Western Hemisphere, Western hemispheres. North America is bordered to the north by the Arctic Ocean, to the east by the Atlantic Ocean, to the southeast by South Ameri ...
include the Illinois State Morel Mushroom Hunting Championship, the Ottawa Midwest Morel Fest and the Mesick Michigan Mushroom Festival. In Tyler Childers 2019 song "All Your'n", he mentions "Fried morels and fine hotels" in the first line of the 2nd verse. In the Farm life sim, farming sim video game ''Stardew Valley'', morels are a consumable item that can be found in the Secret Woods during spring. If the player chooses to allow mushrooms to grow in the Farm Cave, morels have an 8.1% chance of spawning each day regardless of season.


Vernacular names

''Morchella'' species have been called by many local names; some of the more colorful include ''dryland fish'', because when sliced lengthwise then breaded and fried, their outline resembles the shape of a fish; ''hickory chickens'', as they are known in many parts of Kentucky; and ''merkels'' or ''miracles'', based on folklore, of how a mountain family was saved from starvation by eating morels. In parts of West Virginia, they are known as ''molly moochers'', ''muggins'', or ''muggles''. Due to the partial structural and textural similarity to some species of ''Porifera'' (sponges), other common names for any true morel are ''sponge mushroom'' and ''waffle mushroom''. In the Appalachian woodlands, morels have also been called ''haystacks'', or ''snakeheads''. The Finnish language, Finnish vernacular name ''huhtasieni'', refers to ''huhta'', area cleared for agriculture by the slash and burn method. The scientific name of the genus ''Morchella'' itself, is thought to have derived from ''morchel'', an old German language, German word close to "Möhre", carrot or beet, due to similarity in shape. File:Morilles.jpg, left, Yellow morels in France File:Morchella elata group 38336.jpg, Black morels in British Columbia, Canada File:Morel in Ben Shemen Israel.jpg, Morel in a house garden in Ben Shemen, Israel


See also

*


References


External links


2000 Pezizales
on Ascomycete.org


"A Beginner's Guide to Hunting Morel Mushrooms"
from ''Field and Stream''
Colorado Morels
{{Authority control Morchella, Edible fungi, Morel Pezizales genera