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''Megophrys'' is a
genus Genus (; : genera ) is a taxonomic rank above species and below family (taxonomy), family as used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In bino ...
of
frog A frog is any member of a diverse and largely semiaquatic group of short-bodied, tailless amphibian vertebrates composing the order (biology), order Anura (coming from the Ancient Greek , literally 'without tail'). Frog species with rough ski ...
s in the
family Family (from ) is a Social group, group of people related either by consanguinity (by recognized birth) or Affinity (law), affinity (by marriage or other relationship). It forms the basis for social order. Ideally, families offer predictabili ...
Megophryidae Megophryidae, commonly known as goose frogs, is a large family of frogs native to the warm southeast of Asia, from the Himalayan foothills eastwards, south to Indonesia and the Greater Sunda Islands in Maritime Southeast Asia, and extending to t ...
. They are
endemic Endemism is the state of a species being found only in a single defined geographic location, such as an island, state, nation, country or other defined zone; organisms that are indigenous to a place are not endemic to it if they are also foun ...
to
Indonesia Indonesia, officially the Republic of Indonesia, is a country in Southeast Asia and Oceania, between the Indian Ocean, Indian and Pacific Ocean, Pacific oceans. Comprising over List of islands of Indonesia, 17,000 islands, including Sumatra, ...
, where they are found on the islands of
Java Java is one of the Greater Sunda Islands in Indonesia. It is bordered by the Indian Ocean to the south and the Java Sea (a part of Pacific Ocean) to the north. With a population of 156.9 million people (including Madura) in mid 2024, proje ...
and
Sumatra Sumatra () is one of the Sunda Islands of western Indonesia. It is the largest island that is fully within Indonesian territory, as well as the list of islands by area, sixth-largest island in the world at 482,286.55 km2 (182,812 mi. ...
. They commonly have elongated upper "eyebrows" and are thus known as Indonesian horned toads. This group was thought to contain many more species and have a much wider distribution prior to recent taxonomic revisions.


Taxonomy

The following species are recognised in the genus ''Megophrys'': * '' Megophrys acehensis'' * '' Megophrys lancip'' * '' Megophrys montana'' * '' Megophrys parallela'' * '' Megophrys selatanensis'' ''Megophrys'' formerly contained over a hundred species, but significant taxonomic revisions have led to the vast majority of these species being moved to other genera, such as '' Xenophrys'', '' Boulenophrys'', '' Atympanophrys'' and ''
Pelobatrachus ''Pelobatrachus'' is a genus of Frog, frogs in the Family (biology), family Megophryidae. It was formerly synonymized with ''Megophrys'' until 2021, when it was revived as a distinct genus. They inhabit Southeast Asia, namely the Malay Peninsula, ...
''. However, there is a divide between studies over this, with most Indian-published studies preferring to classify all these taxa within ''Megophrys'', while Chinese-published studies classify them in their own genera; the
IUCN Red List The International Union for Conservation of Nature (IUCN) Red List of Threatened Species, also known as the IUCN Red List or Red Data Book, founded in 1964, is an inventory of the global conservation status and extinction risk of biological ...
follows the former, while ''Amphibian Species of the World'' and AmphibiaWeb follow the latter.


Endemic ranges

Many ''Megophrys'' species are
endemic Endemism is the state of a species being found only in a single defined geographic location, such as an island, state, nation, country or other defined zone; organisms that are indigenous to a place are not endemic to it if they are also foun ...
to highly restricted geographical areas. *
Southern China Northern China () and Southern China () are two approximate regions that display certain differences in terms of their geography, demographics, economy, and culture. Extent The Qinling–Daba Mountains serve as the transition zone between ...
**
Anhui Anhui is an inland Provinces of China, province located in East China. Its provincial capital and largest city is Hefei. The province is located across the basins of the Yangtze and Huai rivers, bordering Jiangsu and Zhejiang to the east, Jiang ...
*** '' Megophrys huangshanensis'': Huangshan mountains,
Anhui Anhui is an inland Provinces of China, province located in East China. Its provincial capital and largest city is Hefei. The province is located across the basins of the Yangtze and Huai rivers, bordering Jiangsu and Zhejiang to the east, Jiang ...
**
Guangdong ) means "wide" or "vast", and has been associated with the region since the creation of Guang Prefecture in AD 226. The name "''Guang''" ultimately came from Guangxin ( zh, labels=no, first=t, t= , s=广信), an outpost established in Han dynasty ...
*** '' Megophrys acuta'': Heishiding, Fengkai County,
Guangdong ) means "wide" or "vast", and has been associated with the region since the creation of Guang Prefecture in AD 226. The name "''Guang''" ultimately came from Guangxin ( zh, labels=no, first=t, t= , s=广信), an outpost established in Han dynasty ...
*** '' Megophrys obesa'': Heishiding, Fengkai County,
Guangdong ) means "wide" or "vast", and has been associated with the region since the creation of Guang Prefecture in AD 226. The name "''Guang''" ultimately came from Guangxin ( zh, labels=no, first=t, t= , s=广信), an outpost established in Han dynasty ...
**
Jiangxi ; Gan: ) , translit_lang1_type2 = , translit_lang1_info2 = , translit_lang1_type3 = , translit_lang1_info3 = , image_map = Jiangxi in China (+all claims hatched).svg , mapsize = 275px , map_caption = Location ...
*** '' Megophrys cheni'': Luoxiao Mountains *** '' Megophrys lini'': Luoxiao Mountains *** '' Megophrys jinggangensis'': Luoxiao Mountains **
Hunan Hunan is an inland Provinces of China, province in Central China. Located in the middle reaches of the Yangtze watershed, it borders the Administrative divisions of China, province-level divisions of Hubei to the north, Jiangxi to the east, Gu ...
*** '' Megophrys sangzhiensis'':
Sangzhi County Sangzhi () is a county in Hunan Province, China, it is under administration of the prefecture-level city of Zhangjiajie. Located on the northern margin of Hunan, Sangzhi County is bordered to the east by Cili County, to the south by Yongding a ...
,
Hunan Hunan is an inland Provinces of China, province in Central China. Located in the middle reaches of the Yangtze watershed, it borders the Administrative divisions of China, province-level divisions of Hubei to the north, Jiangxi to the east, Gu ...
*** '' Megophrys caudoprocta'': Tianping Mountain,
Sangzhi County Sangzhi () is a county in Hunan Province, China, it is under administration of the prefecture-level city of Zhangjiajie. Located on the northern margin of Hunan, Sangzhi County is bordered to the east by Cili County, to the south by Yongding a ...
,
Hunan Hunan is an inland Provinces of China, province in Central China. Located in the middle reaches of the Yangtze watershed, it borders the Administrative divisions of China, province-level divisions of Hubei to the north, Jiangxi to the east, Gu ...
*** '' Megophrys tuberogranulatus'': Tianzishan Nature Reserve,
Sangzhi County Sangzhi () is a county in Hunan Province, China, it is under administration of the prefecture-level city of Zhangjiajie. Located on the northern margin of Hunan, Sangzhi County is bordered to the east by Cili County, to the south by Yongding a ...
,
Hunan Hunan is an inland Provinces of China, province in Central China. Located in the middle reaches of the Yangtze watershed, it borders the Administrative divisions of China, province-level divisions of Hubei to the north, Jiangxi to the east, Gu ...
*** '' Megophrys mangshanensis'': Yizhang County,
Hunan Hunan is an inland Provinces of China, province in Central China. Located in the middle reaches of the Yangtze watershed, it borders the Administrative divisions of China, province-level divisions of Hubei to the north, Jiangxi to the east, Gu ...
**
Chongqing ChongqingPostal Romanization, Previously romanized as Chungking ();. is a direct-administered municipality in Southwestern China. Chongqing is one of the four direct-administered municipalities under the State Council of the People's Republi ...
*** '' Megophrys baolongensis'': Baolong (抱龙镇),
Wushan County, Chongqing Wushan County () is a County (People's Republic of China), county located in Chongqing municipality. It occupies roughly and has a population of about 600,000. The county seat is located at the western entrance to the Wu Gorge in the Three Gorge ...
*** '' Megophrys wushanensis'':
Wu Mountains Wuxia Gorge (), sometimes called Great Gorge (), is the second gorge of the Three Gorges system on the Yangtze River, People's Republic of China. Formed by the Wu River, it stretches from Wushan to Guandukou, and is located downstream of ...
,
Chongqing ChongqingPostal Romanization, Previously romanized as Chungking ();. is a direct-administered municipality in Southwestern China. Chongqing is one of the four direct-administered municipalities under the State Council of the People's Republi ...
**
Yunnan Yunnan; is an inland Provinces of China, province in Southwestern China. The province spans approximately and has a population of 47.2 million (as of 2020). The capital of the province is Kunming. The province borders the Chinese provinces ...
*** '' Megophrys binchuanensis'': NW
Yunnan Yunnan; is an inland Provinces of China, province in Southwestern China. The province spans approximately and has a population of 47.2 million (as of 2020). The capital of the province is Kunming. The province borders the Chinese provinces ...
*** '' Megophrys daweimontis'': Mount Dawei, Pingbian Miao Autonomous County,
Yunnan Yunnan; is an inland Provinces of China, province in Southwestern China. The province spans approximately and has a population of 47.2 million (as of 2020). The capital of the province is Kunming. The province borders the Chinese provinces ...
*** '' Megophrys gigantica'': Jingdong Yi Autonomous County and
Yongde County Yongde County () is a county in the west of Yunnan province, China. It is under the administration of the prefecture-level city A prefecture-level city () or prefectural city is an administrative division of the China, People's Republic of ...
, southwestern
Yunnan Yunnan; is an inland Provinces of China, province in Southwestern China. The province spans approximately and has a population of 47.2 million (as of 2020). The capital of the province is Kunming. The province borders the Chinese provinces ...
**
Sichuan Sichuan is a province in Southwestern China, occupying the Sichuan Basin and Tibetan Plateau—between the Jinsha River to the west, the Daba Mountains to the north, and the Yunnan–Guizhou Plateau to the south. Its capital city is Cheng ...
*** '' Megophrys binlingensis'': Binling,
Hongya County Hongya () is a county of Sichuan Province, China. It is under the administration of Meishan city. Administrative divisions Hongya County comprises 12 towns: *Zhige 止戈镇 *Hongchuan 洪川镇 *Yuping 余坪镇 *Caoyutan 槽渔滩镇 *Zhon ...
,
Sichuan Sichuan is a province in Southwestern China, occupying the Sichuan Basin and Tibetan Plateau—between the Jinsha River to the west, the Daba Mountains to the north, and the Yunnan–Guizhou Plateau to the south. Its capital city is Cheng ...
*** '' Megophrys wawuensis'': Mount Wawu,
Hongya County Hongya () is a county of Sichuan Province, China. It is under the administration of Meishan city. Administrative divisions Hongya County comprises 12 towns: *Zhige 止戈镇 *Hongchuan 洪川镇 *Yuping 余坪镇 *Caoyutan 槽渔滩镇 *Zhon ...
,
Sichuan Sichuan is a province in Southwestern China, occupying the Sichuan Basin and Tibetan Plateau—between the Jinsha River to the west, the Daba Mountains to the north, and the Yunnan–Guizhou Plateau to the south. Its capital city is Cheng ...
*** '' Megophrys nankiangensis'': northern
Sichuan Sichuan is a province in Southwestern China, occupying the Sichuan Basin and Tibetan Plateau—between the Jinsha River to the west, the Daba Mountains to the north, and the Yunnan–Guizhou Plateau to the south. Its capital city is Cheng ...
and southern
Gansu Gansu is a provinces of China, province in Northwestern China. Its capital and largest city is Lanzhou, in the southeastern part of the province. The seventh-largest administrative district by area at , Gansu lies between the Tibetan Plateau, Ti ...
**
Guizhou ) , image_skyline = , image_caption = , image_map = Guizhou in China (+all claims hatched).svg , mapsize = 275px , map_alt = Map showing the location of Guizhou Province , map_caption = Map s ...
*** '' Megophrys shuichengensis'': Fenghuang Village, Shuicheng County,
Guizhou ) , image_skyline = , image_caption = , image_map = Guizhou in China (+all claims hatched).svg , mapsize = 275px , map_alt = Map showing the location of Guizhou Province , map_caption = Map s ...
*
Tibet Tibet (; ''Böd''; ), or Greater Tibet, is a region in the western part of East Asia, covering much of the Tibetan Plateau and spanning about . It is the homeland of the Tibetan people. Also resident on the plateau are other ethnic groups s ...
** '' Megophrys medogensis'': Mêdog County,
Tibet Tibet (; ''Böd''; ), or Greater Tibet, is a region in the western part of East Asia, covering much of the Tibetan Plateau and spanning about . It is the homeland of the Tibetan people. Also resident on the plateau are other ethnic groups s ...
** '' Megophrys zhangi'': Zhangmu, Nyalam County,
Tibet Tibet (; ''Böd''; ), or Greater Tibet, is a region in the western part of East Asia, covering much of the Tibetan Plateau and spanning about . It is the homeland of the Tibetan people. Also resident on the plateau are other ethnic groups s ...
*
Northeast India Northeast India, officially the North Eastern Region (NER), is the easternmost region of India representing both a geographic and political Administrative divisions of India, administrative division of the country. It comprises eight States and ...
** '' Megophrys megacephala'': East Khasi Hills District,
Meghalaya Meghalaya (; "the abode of clouds") is a states and union territories of India, state in northeast India. Its capital is Shillong. Meghalaya was formed on 21 January 1972 by carving out two districts from the Assam: the United Khasi Hills an ...
** '' Megophrys oropedion'': East Khasi Hills District,
Meghalaya Meghalaya (; "the abode of clouds") is a states and union territories of India, state in northeast India. Its capital is Shillong. Meghalaya was formed on 21 January 1972 by carving out two districts from the Assam: the United Khasi Hills an ...
** '' Megophrys vegrandis'':
West Kameng District West Kameng (pronounced ) is a district of Arunachal Pradesh in northeastern India. It accounts for 8.86% of the total area of the state. The name is derived from the Kameng river, a tributary of the Brahmaputra, that flows through the distri ...
,
Arunachal Pradesh Arunachal Pradesh (; ) is a States and union territories of India, state in northeast India. It was formed from the North-East Frontier Agency (NEFA) region, and India declared it as a state on 20 February 1987. Itanagar is its capital and la ...
** '' Megophrys ancrae'':
Changlang District Changlang district (Pron:/tʃæŋˈlæŋ/) is located in the Indian state of Arunachal Pradesh, located south of Lohit District, Lohit district and north of Tirap District, Tirap district. Naga people reside here . As of 2011 it is the second ...
,
Arunachal Pradesh Arunachal Pradesh (; ) is a States and union territories of India, state in northeast India. It was formed from the North-East Frontier Agency (NEFA) region, and India declared it as a state on 20 February 1987. Itanagar is its capital and la ...
** '' Megophrys serchhipii'': Serchhip,
Mizoram Mizoram is a states and union territories of India, state in northeastern India, with Aizawl as its Capital city, capital and largest city. It shares 722-kilometres (449 miles) of international borders with Bangladesh to the west, and Myanmar t ...
** '' Megophrys zunhebotoensis'': Nguti (Sukhalu), Zunheboto District,
Nagaland Nagaland () is a States and union territories of India, state in the northeast India, north-eastern region of India. It is bordered by the Indian states of Arunachal Pradesh to the north, Assam to the west, Manipur to the south, and the Naga Sel ...
*
Cardamom Mountains The Cardamom Mountains (, ; , ), or the Krâvanh Mountains, is a mountain range in the southwest part of Cambodia and Eastern Thailand. The majority of the range is within Cambodia. The silhouette of the Cardamom Mountains appears in the Sea ...
,
Cambodia Cambodia, officially the Kingdom of Cambodia, is a country in Southeast Asia on the Mainland Southeast Asia, Indochinese Peninsula. It is bordered by Thailand to the northwest, Laos to the north, and Vietnam to the east, and has a coastline ...
** '' Megophrys auralensis'': Phnom Aural, Kampong Speu Province ** '' Megophrys damrei'': Dâmrei Mountains, Kampot Province *
Vietnam Vietnam, officially the Socialist Republic of Vietnam (SRV), is a country at the eastern edge of mainland Southeast Asia, with an area of about and a population of over 100 million, making it the world's List of countries and depende ...
** '' Megophrys latidactyla'': Pù Mát National Park,
Vietnam Vietnam, officially the Socialist Republic of Vietnam (SRV), is a country at the eastern edge of mainland Southeast Asia, with an area of about and a population of over 100 million, making it the world's List of countries and depende ...
*
Thailand Thailand, officially the Kingdom of Thailand and historically known as Siam (the official name until 1939), is a country in Southeast Asia on the Mainland Southeast Asia, Indochinese Peninsula. With a population of almost 66 million, it spa ...
** '' Megophrys lekaguli'':
Chanthaburi Chanthaburi (, ) is a town ('' thesaban mueang'') in the east of Thailand, on the banks of the Chanthaburi River. It is the capital of the Chanthaburi Province and the Mueang Chanthaburi District. The town covers the two ''tambons'' Talat an ...
and Sa Kaeo Provinces, eastern
Thailand Thailand, officially the Kingdom of Thailand and historically known as Siam (the official name until 1939), is a country in Southeast Asia on the Mainland Southeast Asia, Indochinese Peninsula. With a population of almost 66 million, it spa ...
** '' Megophrys takensis'': Tak Province,
Thailand Thailand, officially the Kingdom of Thailand and historically known as Siam (the official name until 1939), is a country in Southeast Asia on the Mainland Southeast Asia, Indochinese Peninsula. With a population of almost 66 million, it spa ...
*
Borneo Borneo () is the List of islands by area, third-largest island in the world, with an area of , and population of 23,053,723 (2020 national censuses). Situated at the geographic centre of Maritime Southeast Asia, it is one of the Greater Sunda ...
** '' Megophrys baluensis'':
Sabah Sabah () is a States and federal territories of Malaysia, state of Malaysia located in northern Borneo, in the region of East Malaysia. Sabah has land borders with the Malaysian state of Sarawak to the southwest and Indonesia's North Kalima ...
** '' Megophrys dringi'': Mount Mulu,
Sarawak Sarawak ( , ) is a States and federal territories of Malaysia, state of Malaysia. It is the largest among the 13 states, with an area almost equal to that of Peninsular Malaysia. Sarawak is located in East Malaysia in northwest Borneo, and is ...
*
Sumatra Sumatra () is one of the Sunda Islands of western Indonesia. It is the largest island that is fully within Indonesian territory, as well as the list of islands by area, sixth-largest island in the world at 482,286.55 km2 (182,812 mi. ...
** '' Megophrys parallela'':
West Sumatra West Sumatra () is a Provinces of Indonesia, province of Indonesia. It is on the west coast of the island of Sumatra and includes the Mentawai Islands off that coast. West Sumatra borders the Indian Ocean to the west, as well as the provinces of ...
*
Philippines The Philippines, officially the Republic of the Philippines, is an Archipelagic state, archipelagic country in Southeast Asia. Located in the western Pacific Ocean, it consists of List of islands of the Philippines, 7,641 islands, with a tot ...
** '' Megophrys ligayae'': Balabac and
Palawan Palawan (, ), officially the Province of Palawan (; ), is an archipelagic province of the Philippines that is located in the region of Mimaropa. It is the largest province in the country in terms of total area of . The capital and largest c ...
** '' Megophrys stejnegeri'':
Mindanao Mindanao ( ) is the List of islands of the Philippines, second-largest island in the Philippines, after Luzon, and List of islands by population, seventh-most populous island in the world. Located in the southern region of the archipelago, the ...
*
Indonesia Indonesia, officially the Republic of Indonesia, is a country in Southeast Asia and Oceania, between the Indian Ocean, Indian and Pacific Ocean, Pacific oceans. Comprising over List of islands of Indonesia, 17,000 islands, including Sumatra, ...
** '' Megophrys montana'':
Java Java is one of the Greater Sunda Islands in Indonesia. It is bordered by the Indian Ocean to the south and the Java Sea (a part of Pacific Ocean) to the north. With a population of 156.9 million people (including Madura) in mid 2024, proje ...


References


External links


Sound recordings of ''Megophrys'' at BioAcoustica
{{Authority control Megophryidae Amphibians of Asia Amphibian genera Taxa named by Heinrich Kuhl