''Megalocottus'' is a small

genus Genus (; : genera ) is a taxonomic rank above species and below family (taxonomy), family as used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In bino ...

of marine

ray-finned fish Actinopterygii (; ), members of which are known as ray-finned fish or actinopterygians, is a class of bony fish that comprise over 50% of living vertebrate species. They are so called because of their lightly built fins made of webbings of sk ...

es belonging to the

family Family (from ) is a Social group, group of people related either by consanguinity (by recognized birth) or Affinity (law), affinity (by marriage or other relationship). It forms the basis for social order. Ideally, families offer predictabili ...

Cottidae The Cottidae are a family of fish in the superfamily Cottoidea, the sculpins. It is the largest sculpin family, with about 275 species in 70 genera.Kane, E. A. and T. E. Higham. (2012)Life in the flow lane: differences in pectoral fin morphology ...

, the typical sculpins. These fishes are found in the western

Pacific Ocean The Pacific Ocean is the largest and deepest of Earth's five Borders of the oceans, oceanic divisions. It extends from the Arctic Ocean in the north to the Southern Ocean, or, depending on the definition, to Antarctica in the south, and is ...

Taxonomy

''Megalocottus'' was first proposed as a

monospecific genus In biology, a monotypic taxon is a taxonomic group (taxon) that contains only one immediately subordinate taxon. A monotypic species is one that does not include subspecies or smaller, infraspecific taxa. In the case of genera, the term "unispe ...

in 1861 by the American

biologist A biologist is a scientist who conducts research in biology. Biologists are interested in studying life on Earth, whether it is an individual Cell (biology), cell, a multicellular organism, or a Community (ecology), community of Biological inter ...

Theodore Gill Theodore Nicholas Gill (March 21, 1837 – September 25, 1914) was an American ichthyologist, mammalogist, malacologist, and librarian. Career Born and educated in New York City under private tutors, Gill early showed interest in natural hist ...

with ''Cottus paltycephalus'' which had been described in 1814 by

Peter Simon Pallas Peter Simon Pallas Fellow of the Royal Society, FRS FRSE (22 September 1741 – 8 September 1811) was a Prussia, Prussian zoologist, botanist, Ethnography, ethnographer, Exploration, explorer, Geography, geographer, Geology, geologist, Natura ...

from

Kamchatka The Kamchatka Peninsula (, ) is a peninsula in the Russian Far East, with an area of about . The Pacific Ocean and the Sea of Okhotsk make up the peninsula's eastern and western coastlines, respectively. Immediately offshore along the Pacific ...

and the

Sea of Okhotsk The Sea of Okhotsk; Historically also known as , or as ; ) is a marginal sea of the western Pacific Ocean. It is located between Russia's Kamchatka Peninsula on the east, the Kuril Islands on the southeast, Japan's island of Hokkaido on the sou ...

as its only species. The 5th edition of ''

Fishes of the World ''Fishes of the World'' is a standard reference for the systematics of fishes. It was first written in 1976 by the American ichthyologist Joseph S. Nelson (1937–2011). Now in its fifth edition (2016), the work is a comprehensive overview of t ...

'' classifies this genus in the

subfamily In biological classification, a subfamily (Latin: ', plural ') is an auxiliary (intermediate) taxonomic rank, next below family but more inclusive than genus. Standard nomenclature rules end botanical subfamily names with "-oideae", and zo ...

Cottinae Cottinae is a subfamily of ray-finned fishes belonging to the Family (biology), family Cottidae, the typical sculpins. The subfamily has species throughout the northern hemisphere in both marine and freshwater habitats. Genera The following gener ...

of the family Cottidae but other authorities classify it in the subfamily Myoxocephalinae of the family

Psychrolutidae The fish family Psychrolutidae (commonly known as blobfishes, flathead sculpins, or tadpole sculpins) contains over 35 recognized species in 8 genera. This family consists of bottom-dwelling marine sculpins shaped like tadpoles, with large heads ...

, although others place the subfamily Myoxocephalinae within the Cottidae.

Etymology

''Megalocottus'' prefixes ''megalo'' meaning "great" or large" with Cottus, presumed to be a reference to the large size of the

type species In International_Code_of_Zoological_Nomenclature, zoological nomenclature, a type species (''species typica'') is the species name with which the name of a genus or subgenus is considered to be permanently taxonomically associated, i.e., the spe ...

Species

''Megalocottus'' currently contains two recognized: * '' Megalocottus platycephalus'' (Pallas, 1814) (Belligerent sculpin) * '' Megalocottus taeniopterus'' Kner, 1868 (Southern flathead sculpin) ''

Fishbase FishBase is a global species database of fish species (specifically finfish). It is the largest and most extensively accessed online database on adult finfish on the web.

'' retains these two species as valid, other authorities treat the Southern flathead sculpin as a subspecies of the belligerent sculpin while others treat it as a ''M. taeniopterus'' as a

junior synonym In taxonomy, the scientific classification of living organisms, a synonym is an alternative scientific name for the accepted scientific name of a taxon. The botanical and zoological codes of nomenclature treat the concept of synonymy differently. ...

of ''M. platycephalus''. This taxon is a

sister taxon In phylogenetics, a sister group or sister taxon, also called an adelphotaxon, comprises the closest relative(s) of another given unit in an evolutionary tree. Definition The expression is most easily illustrated by a cladogram: Taxon A and ...

to the genus ''

Myoxocephalus ''Myoxocephalus'' is a genus of marine ray-finned fishes belonging to the family Cottidae, the typical sculpins. They are found in the northern Pacific, Arctic and Atlantic Oceans, with a few species in lakes. Taxonomy ''Myoxocephalus'' was firs ...

''. In 2020 workers undertook a comparison of the

Mitochondrial DNA Mitochondrial DNA (mtDNA and mDNA) is the DNA located in the mitochondrion, mitochondria organelles in a eukaryotic cell that converts chemical energy from food into adenosine triphosphate (ATP). Mitochondrial DNA is a small portion of the D ...

of the

fourhorn sculpin The fourhorn sculpin (''Myoxocephalus quadricornis'') is a species of ray-finned fish belonging to the family Cottidae, the typical sculpins. This species has a Holarctic distribution and can be found in marine, brackish and fresh waters. Taxono ...

(''Myoxocephalus quadricornis'') and the belligerent sculpin and found that the fourhorn sculpin was more closely related to the belligerent sculpin than it was to the other species in the genus ''Myoxocephalus'', they proposed that the fourhorn sculpin be reclassified as a member of the genus ''Megalocottus''.

References

{{Taxonbar, from=Q19921645 Cottinae Taxa named by Theodore Gill Ray-finned fish genera