Mecodema
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''Mecodema'' is a genus of large flightless
ground beetle Ground beetles are a large, cosmopolitan distribution, cosmopolitan family (biology), family of beetles, the Carabidae, with more than 40,000 species worldwide, around 2,000 of which are found in North America and 2,700 in Europe. As of 2015, it ...
(Carabidae)
endemic Endemism is the state of a species being found only in a single defined geographic location, such as an island, state, nation, country or other defined zone; organisms that are indigenous to a place are not endemic to it if they are also foun ...
to New Zealand. The genus is very diverse in comparison to the other three New Zealand genera ('' Diglymma'', '' Oregus'', '' Orthoglymma'') within the subtribe
Nothobroscina Nothobroscina Roig-Juñent 2000 is one of five subtribes within the ground beetle tribe Broscini. The subtribe consists of ten genera from the Gondwana Gondwana ( ; ) was a large landmass, sometimes referred to as a supercontinent. The re ...
(tribe
Broscini The tribe Broscini is a worldwide group of beetles in the Broscinae subfamily of Carabidae (the ground beetle Ground beetles are a large, cosmopolitan distribution, cosmopolitan family (biology), family of beetles, the Carabidae, with more th ...
). ''Mecodema'' is geographically widespread across both the
North North is one of the four compass points or cardinal directions. It is the opposite of south and is perpendicular to east and west. ''North'' is a noun, adjective, or adverb indicating Direction (geometry), direction or geography. Etymology T ...
and
South Island The South Island ( , 'the waters of Pounamu, Greenstone') is the largest of the three major islands of New Zealand by surface area, the others being the smaller but more populous North Island and Stewart Island. It is bordered to the north by ...
s, as well as numerous offshore islands, including the Three Kings Is., Poor Knights Is., Aotea ( Great Barrier Is.) and Hauturu (Little Barrier Is.), Kapiti Is., Stephens Is., Stewart Is., Chatham Is., Snares Is.


Genus description

Head, frons and vertex often rugose and / or punctate, microsculpture or macrosculpture absent in some species groups (e.g., ''curvidens'' species); eyes prominent, almost hemispherical; a single supraorbital puncture on each side, bearing more than one setae; setose punctures of vertex absent; mandibles large, uniformly narrow to the pointed apex, sharply curved in the apical third, mandibles with a number of dorsal grooves and without a seta in the scrobe; mentum with median process, which is notched to strongly indentate (bifid); usually with 2 setiferous punctures (may be absent) below the process and generally with 4–8 setose (setiferous) punctures along the submentum sclerite; labial palpi with setae on the penultimate segment; terminal segments of all palpi slightly compressed, sub-cylindrical, truncate; ligula chitinous with a median carina, 2 apical setae present; the membranous paraglossae extended well beyond the apex of the ligula; stipes with 2–3 basal setae; antennomeres I–II glabrous, III–V with apical ring, VI–XI setose throughout; antennomeres I–III subcylindrical, IV–VII subglobose and slightly compressed. Prothoracic carina (pronotal lateral margin) distinct with an asymmetrical number of setae along each side, the last never at the posterior lateral sinuation (posterior angle); midline (median groove) and pronotal foveae (laterobasal depressions) always present; pronotal disc often with shallow depressions on each side of apical half. Elytra always striate; fused together along the suture; hind wings absent; a row of setose punctures on the 7th stria and another row (umbilical series) near the lateral carina (elytral margin). Legs slender, the protibia with a pointed apical prolongation on the outer side, greatly distally expanded to distally expanded and shovel-like; male and female tarsi identical, the basal 4 segments of the anterior tarsi are as broad as they are long, usually slightly asymmetrical. Type species: '' Mecodema sculpturatum''
Blanchard Blanchard is a French family name. It is also used as a given name. It derives from the Old French word ''blanchart'' which meant "whitish, bordering upon white". It is also an obsolete term for a white horse. Geographical distribution As of 2014, ...
1853.


Species and subspecies (103)

* ''
Mecodema aberrans ''Mecodema aberrans'' is a medium-sized (14.6–19.5 mm length, 5.3–6.5 mm width) ground beetle endemic to the South Island The South Island ( , 'the waters of Pounamu, Greenstone') is the largest of the three major islands of New Zealan ...
'' (Putzeys, 1868) * ''
Mecodema allani ''Mecodema allani'' is a ground beetle of the family Carabidae, endemic to the South Island, New Zealand. It is one of two species within the ''laterale'' group, which are large-bodied species with a distinctively broad lateral carina along the ...
'' Fairburn, 1945 * ''
Mecodema alternans ''Mecodema'' is a genus of large flightless ground beetle (Carabidae) endemic to New Zealand. The genus is very diverse in comparison to the other three New Zealand genera ('' Diglymma'', '' Oregus'', '' Orthoglymma'') within the subtribe Nothobr ...
'' Laporte de Castelnau, 1867 ** ''Mecodema alternans hudsoni''
Broun Broun is a surname. It is the Middle English and Scots spelling of Brown. Notable people with the surname include: *Agnes Broun (1732–1820), mother of Scottish poet Robert Burns * Alex Broun (born 1965), Australian playwright and screenwriter *D ...
, 1909 * ''
Mecodema angustulum ''Mecodema'' is a genus of large flightless ground beetle (Carabidae) endemic to New Zealand. The genus is very diverse in comparison to the other three New Zealand genera ('' Diglymma'', '' Oregus'', '' Orthoglymma'') within the subtribe Nothobr ...
''
Broun Broun is a surname. It is the Middle English and Scots spelling of Brown. Notable people with the surname include: *Agnes Broun (1732–1820), mother of Scottish poet Robert Burns * Alex Broun (born 1965), Australian playwright and screenwriter *D ...
, 1914 * ''
Mecodema antarcticum ''Mecodema antarcticum'' is a carnivorous carabid beetle that burrows in sand above the high tide mark on New Zealand sandy beaches. First described by Francis de Laporte de Castelnau in 1867 as ''Brullea antarctica'', it has since been reassig ...
'' (Laporte de Castelnau, 1867) * ''
Mecodema aoteanoho ''Mecodema aoteanoho'' is a medium-sized (19–24 mm length, 5.5–7 mm width) ground beetle, the only such beetle endemic to Great Barrier Island (Aotea), Hauraki Gulf, Auckland, New Zealand. ''Mecodema aoteanoho'' is a sister species to '' Me ...
'' Seldon & Leschen, 2011 * ''
Mecodema argentum ''Mecodema argentum'' is a large-bodied (29–32 mm length, 8.5–10 mm width) ground beetle found in the northern region of the Coromandel Range, New Zealand and relatively abundant in the hills above Coromandel town. ''Mecodema argen ...
'' Seldon & Buckley, 2019 * ''
Mecodema atrox ''Mecodema atrox'' is a medium-sized (17–24 mm length, 6–8 mm width) ground beetle species that is closely related to '' Mecodema curvidens'' (a very widespread species throughout the Bay of Plenty and Central Plateau). ''Mecodema ...
'' Britton, 1949 * '' Mecodema atuanui'' Seldon & Buckley, 2019 * '' Mecodema brittoni'' Townsend, 1965 * '' Mecodema bullatum'' Lewis, 1902 * '' Mecodema chaiup'' Seldon, 2015 * '' Mecodema chiltoni''
Broun Broun is a surname. It is the Middle English and Scots spelling of Brown. Notable people with the surname include: *Agnes Broun (1732–1820), mother of Scottish poet Robert Burns * Alex Broun (born 1965), Australian playwright and screenwriter *D ...
, 1917 * '' Mecodema constrictum'' Broun, 1881 * '' Mecodema costellum''
Broun Broun is a surname. It is the Middle English and Scots spelling of Brown. Notable people with the surname include: *Agnes Broun (1732–1820), mother of Scottish poet Robert Burns * Alex Broun (born 1965), Australian playwright and screenwriter *D ...
, 1903 ** ''Mecodema costellum gordonense''
Broun Broun is a surname. It is the Middle English and Scots spelling of Brown. Notable people with the surname include: *Agnes Broun (1732–1820), mother of Scottish poet Robert Burns * Alex Broun (born 1965), Australian playwright and screenwriter *D ...
, 1917 ** ''Mecodema costellum lewisi''
Broun Broun is a surname. It is the Middle English and Scots spelling of Brown. Notable people with the surname include: *Agnes Broun (1732–1820), mother of Scottish poet Robert Burns * Alex Broun (born 1965), Australian playwright and screenwriter *D ...
, 1908 ** ''Mecodema costellum obesum'' Townsend, 1965 * '' Mecodema costipenne''
Broun Broun is a surname. It is the Middle English and Scots spelling of Brown. Notable people with the surname include: *Agnes Broun (1732–1820), mother of Scottish poet Robert Burns * Alex Broun (born 1965), Australian playwright and screenwriter *D ...
, 1914 * '' Mecodema crenaticolle'' Redtenbacher, 1868 * '' Mecodema crenicolle'' Castelnau, 1867 * '' Mecodema curvidens'' (
Broun Broun is a surname. It is the Middle English and Scots spelling of Brown. Notable people with the surname include: *Agnes Broun (1732–1820), mother of Scottish poet Robert Burns * Alex Broun (born 1965), Australian playwright and screenwriter *D ...
, 1915) * '' Mecodema ducale'' Sharp, 1886 * '' Mecodema dunense'' Townsend, 1965 * '' Mecodema dunnorum'' Seldon & Buckley, 2019 * '' Mecodema dux'' Britton, 1949 * '' Mecodema elongatum'' Castelnau, 1867 * '' Mecodema femorale''
Broun Broun is a surname. It is the Middle English and Scots spelling of Brown. Notable people with the surname include: *Agnes Broun (1732–1820), mother of Scottish poet Robert Burns * Alex Broun (born 1965), Australian playwright and screenwriter *D ...
, 1921 * '' Mecodema florae'' Britton, 1949 * '' Mecodema fulgidum''
Broun Broun is a surname. It is the Middle English and Scots spelling of Brown. Notable people with the surname include: *Agnes Broun (1732–1820), mother of Scottish poet Robert Burns * Alex Broun (born 1965), Australian playwright and screenwriter *D ...
, 1881 * '' Mecodema genesispotini'' Seldon & Buckley, 2019 * '' Mecodema godzilla'' Seldon & Buckley, 2019 * '' Mecodema gourlayi'' Britton, 1949 * ''
Mecodema haakuturi ''Mecodema'' is a genus of large flightless ground beetle (Carabidae) endemic to New Zealand. The genus is very diverse in comparison to the other three New Zealand genera ('' Diglymma'', '' Oregus'', '' Orthoglymma'') within the subtribe Nothobr ...
'' Seldon & Pou, 2024 * '' Mecodema haunoho'' Seldon & Leschen, 2011 * '' Mecodema hector'' Britton, 1949 * '' Mecodema howitti'' Castelnau, 1867 * '' Mecodema huttense''
Broun Broun is a surname. It is the Middle English and Scots spelling of Brown. Notable people with the surname include: *Agnes Broun (1732–1820), mother of Scottish poet Robert Burns * Alex Broun (born 1965), Australian playwright and screenwriter *D ...
, 1915 * '' Mecodema impressum'' Castelnau, 1867 * '' Mecodema infimate'' Lewis, 1902 * '' Mecodema integratum'' Townsend, 1965 * '' Mecodema jacinda'' Seldon & Buckley, 2019 * '' Mecodema kipjac'' Seldon & Buckley, 2019 * '' Mecodema kokoroiho'' Seldon & Buckley, 2019 * '' Mecodema kokoromatua'' Seldon, Leschen & Liebherr, 2012 * '' Mecodema laeviceps''
Broun Broun is a surname. It is the Middle English and Scots spelling of Brown. Notable people with the surname include: *Agnes Broun (1732–1820), mother of Scottish poet Robert Burns * Alex Broun (born 1965), Australian playwright and screenwriter *D ...
, 1904 * '' Mecodema laterale''
Broun Broun is a surname. It is the Middle English and Scots spelling of Brown. Notable people with the surname include: *Agnes Broun (1732–1820), mother of Scottish poet Robert Burns * Alex Broun (born 1965), Australian playwright and screenwriter *D ...
, 1917 * '' Mecodema litoreum''
Broun Broun is a surname. It is the Middle English and Scots spelling of Brown. Notable people with the surname include: *Agnes Broun (1732–1820), mother of Scottish poet Robert Burns * Alex Broun (born 1965), Australian playwright and screenwriter *D ...
, 1886 * '' Mecodema longicolle''
Broun Broun is a surname. It is the Middle English and Scots spelling of Brown. Notable people with the surname include: *Agnes Broun (1732–1820), mother of Scottish poet Robert Burns * Alex Broun (born 1965), Australian playwright and screenwriter *D ...
, 1923 * '' Mecodema lucidum'' Castelnau, 1867 * '' Mecodema manaia'' Seldon & Leschen, 2011 * '' Mecodema metallicum'' Sharp, 1886 * '' Mecodema minax'' Britton, 1949 * '' Mecodema mohi'' Seldon & Buckley, 2019 * '' Mecodema moniliferum'' (Bates, 1867) * '' Mecodema morio'' ( Castelnau, 1867) * '' Mecodema ngaiatonga'' Seldon & Buckley, 2019 * '' Mecodema ngaitahuhu'' Seldon & Buckley, 2019 * '' Mecodema nitidum''
Broun Broun is a surname. It is the Middle English and Scots spelling of Brown. Notable people with the surname include: *Agnes Broun (1732–1820), mother of Scottish poet Robert Burns * Alex Broun (born 1965), Australian playwright and screenwriter *D ...
, 1903 * '' Mecodema oblongum'' (
Broun Broun is a surname. It is the Middle English and Scots spelling of Brown. Notable people with the surname include: *Agnes Broun (1732–1820), mother of Scottish poet Robert Burns * Alex Broun (born 1965), Australian playwright and screenwriter *D ...
, 1882) * '' Mecodema oconnori''
Broun Broun is a surname. It is the Middle English and Scots spelling of Brown. Notable people with the surname include: *Agnes Broun (1732–1820), mother of Scottish poet Robert Burns * Alex Broun (born 1965), Australian playwright and screenwriter *D ...
, 1912 * '' Mecodema oregoides'' (
Broun Broun is a surname. It is the Middle English and Scots spelling of Brown. Notable people with the surname include: *Agnes Broun (1732–1820), mother of Scottish poet Robert Burns * Alex Broun (born 1965), Australian playwright and screenwriter *D ...
, 1894) * '' Mecodema papake'' Seldon & Buckley, 2019 * '' Mecodema parataiko'' Seldon & Leschen, 2011 * '' Mecodema pavidum'' Townsend, 1965 * '' Mecodema perexiguus'' Seldon & Buckley, 2019 * '' Mecodema persculptum'' Broun, 1915 * '' Mecodema pluto'' Britton, 1949 * '' Mecodema politanum''
Broun Broun is a surname. It is the Middle English and Scots spelling of Brown. Notable people with the surname include: *Agnes Broun (1732–1820), mother of Scottish poet Robert Burns * Alex Broun (born 1965), Australian playwright and screenwriter *D ...
, 1917 * '' Mecodema ponaiti'' Seldon & Leschen, 2011 * '' Mecodema proximum'' Britton, 1949 * '' Mecodema puiakium'' Johns & Ewers in Johns, 2007 * '' Mecodema pulchellum'' Townsend, 1965 * '' Mecodema punctatum'' (Laporte de Castelnau, 1867) * '' Mecodema punctellum''
Broun Broun is a surname. It is the Middle English and Scots spelling of Brown. Notable people with the surname include: *Agnes Broun (1732–1820), mother of Scottish poet Robert Burns * Alex Broun (born 1965), Australian playwright and screenwriter *D ...
, 1921 * '' Mecodema puncticolle''
Broun Broun is a surname. It is the Middle English and Scots spelling of Brown. Notable people with the surname include: *Agnes Broun (1732–1820), mother of Scottish poet Robert Burns * Alex Broun (born 1965), Australian playwright and screenwriter *D ...
, 1915 Mecodema sculpturatum'' Blanchard, 1843">='' Mecodema sculpturatum'' Blanchard, 1843* '' Mecodema quoinense''
Broun Broun is a surname. It is the Middle English and Scots spelling of Brown. Notable people with the surname include: *Agnes Broun (1732–1820), mother of Scottish poet Robert Burns * Alex Broun (born 1965), Australian playwright and screenwriter *D ...
, 1912 * '' Mecodema rectolineatum'' Laporte de Castelnau, 1867 * '' Mecodema regulus'' Britton, 1964 * ''
Mecodema rex ''Mecodema'' is a genus of large flightless ground beetle (Carabidae) Endemism, endemic to New Zealand. The genus is very diverse in comparison to the other three New Zealand genera (''Diglymma'', ''Oregus'', ''Orthoglymma'') within the subtribe ...
'' Britton, 1949 * '' Mecodema rugiceps'' Sharp, 1886 ** ''Mecodema rugiceps anomalum'' Townsend, 1965 * '' Mecodema rusticulus'' Seldon & Buckley, 2019 * '' Mecodema scitulum'' Broun, 1894 * '' Mecodema sculpturatum'' Blanchard, 1843 * '' Mecodema simplex'' Castelnau, 1867 * '' Mecodema spiniferum''
Broun Broun is a surname. It is the Middle English and Scots spelling of Brown. Notable people with the surname include: *Agnes Broun (1732–1820), mother of Scottish poet Robert Burns * Alex Broun (born 1965), Australian playwright and screenwriter *D ...
, 1880 * '' Mecodema striatum''
Broun Broun is a surname. It is the Middle English and Scots spelling of Brown. Notable people with the surname include: *Agnes Broun (1732–1820), mother of Scottish poet Robert Burns * Alex Broun (born 1965), Australian playwright and screenwriter *D ...
, 1904 * '' Mecodema strictum'' Britton, 1949 * '' Mecodema temata'' Seldon & Buckley, 2019 * '' Mecodema tenaki'' Seldon & Leschen, 2011 * '' Mecodema teparawhau'' Seldon & Buckley, 2019 * '' Mecodema teroroa'' Seldon & Buckley, 2019 * '' Mecodema tewhara'' Seldon & Buckley, 2019 * '' Mecodema tibiale'' (Laporte de Castelnau, 1867) * '' Mecodema tuhoe'' Seldon & Buckley, 2019 * '' Mecodema undecimus'' Seldon & Buckley, 2019 * '' Mecodema validum''
Broun Broun is a surname. It is the Middle English and Scots spelling of Brown. Notable people with the surname include: *Agnes Broun (1732–1820), mother of Scottish poet Robert Burns * Alex Broun (born 1965), Australian playwright and screenwriter *D ...
, 1923 * '' Mecodema wharekahika'' Seldon & Buckley, 2019 * '' Mecodema xylanthrax'' Seldon & Buckley, 2019 * '' Mecodema yconomus'' Seldon & Buckley, 2019 * '' Mecodema zonula'' Seldon & Buckley, 2019


References


Further reading

* Seldon, D. S. (2015). A unique species of ''Mecodema'' (Carabidae: Broscini) from the Hawke's Bay region, New Zealand, with implications for North–South Island zoogeography. ''New Zealand Entomologist'', ''38''(1), 28–35. * Seldon, D. S., & Leschen, R. A. (2011). Revision of the ''Mecodema curvidens'' species group (Coleoptera: Carabidae: Broscini). ''Zootaxa'', ''2829''(1), 1-45. * Seldon, D. S., Leschen, R. A. B., & Liebherr, J. K. (2012). A new species of ''Mecodema'' (Carabidae: Broscini) from Northland, New Zealand, with notes on a newly observed structure within the female genitalia. ''New Zealand Entomologist'', ''35''(1), 39–50. * Townsend, J. I. (1965). Notes on the genus ''Mecodema'' (Coleoptera: Carabidae) with descriptions of new species from the South Island of New Zealand. ''New Zealand Journal of Science'', ''8''(3), 301–318. * Townsend, J. I. (1971). Entomology of the Aucklands and other islands south of New Zealand: Coleoptera: Carabidae: Broscini. ''Pacific Insects Monograph'', ''27'', 173–184.
NZ Entomologist



Landcare Research
{{Authority control Carabidae genera Endemic beetles of New Zealand Taxa named by Émile Blanchard