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The Mascarene Islands (, ) or Mascarenes or Mascarenhas Archipelago is a group of
island An island or isle is a piece of land, distinct from a continent, completely surrounded by water. There are continental islands, which were formed by being split from a continent by plate tectonics, and oceanic islands, which have never been ...
s in the
Indian Ocean The Indian Ocean is the third-largest of the world's five oceanic divisions, covering or approximately 20% of the water area of Earth#Surface, Earth's surface. It is bounded by Asia to the north, Africa to the west and Australia (continent), ...
east of
Madagascar Madagascar, officially the Republic of Madagascar, is an island country that includes the island of Madagascar and numerous smaller peripheral islands. Lying off the southeastern coast of Africa, it is the world's List of islands by area, f ...
consisting of islands belonging to the
Republic of Mauritius Mauritius, officially the Republic of Mauritius, is an island country in the Indian Ocean, about off the southeastern coast of East Africa, east of Madagascar. It includes the main island (also called Mauritius), as well as Rodrigues, Agalà ...
as well as the French department of
Réunion Réunion (; ; ; known as before 1848) is an island in the Indian Ocean that is an overseas departments and regions of France, overseas department and region of France. Part of the Mascarene Islands, it is located approximately east of the isl ...
. Their name derives from the
Portuguese Portuguese may refer to: * anything of, from, or related to the country and nation of Portugal ** Portuguese cuisine, traditional foods ** Portuguese language, a Romance language *** Portuguese dialects, variants of the Portuguese language ** Portu ...
navigator
Pedro Mascarenhas D. Pedro Mascarenhas (1480 – 16 June 1555) was a Portuguese explorer and colonial administrator. He was the first European to discover the island of Diego Garcia in the Indian Ocean in 1512. He also encountered the Indian Ocean island of Ma ...
, who first visited them in April 1512. The islands share a common
geological Geology (). is a branch of natural science concerned with the Earth and other astronomical objects, the rocks of which they are composed, and the processes by which they change over time. Modern geology significantly overlaps all other Earth s ...
origin beneath the
Mascarene Plateau The Mascarene Plateau is a submarine plateau in the Indian Ocean, north and east of Madagascar. The plateau extends approximately , from Seychelles in the north to Réunion in the south. The plateau covers an area of over of shallow water, wi ...
known as the Mauritia microcontinent which was a
Precambrian The Precambrian ( ; or pre-Cambrian, sometimes abbreviated pC, or Cryptozoic) is the earliest part of Earth's history, set before the current Phanerozoic Eon. The Precambrian is so named because it preceded the Cambrian, the first period of t ...
microcontinent Continental crustal fragments, partly synonymous with microcontinents, are pieces of continents that have broken off from main continental masses to form distinct islands that are often several hundred kilometers from their place of origin. Caus ...
situated between
India India, officially the Republic of India, is a country in South Asia. It is the List of countries and dependencies by area, seventh-largest country by area; the List of countries by population (United Nations), most populous country since ...
and
Madagascar Madagascar, officially the Republic of Madagascar, is an island country that includes the island of Madagascar and numerous smaller peripheral islands. Lying off the southeastern coast of Africa, it is the world's List of islands by area, f ...
until their separation about 70 million years ago. They form a distinct
ecoregion An ecoregion (ecological region) is an ecological and geographic area that exists on multiple different levels, defined by type, quality, and quantity of environmental resources. Ecoregions cover relatively large areas of land or water, and c ...
with unique biodiversity and endemism of flora and fauna.


Geography

The
archipelago An archipelago ( ), sometimes called an island group or island chain, is a chain, cluster, or collection of islands. An archipelago may be in an ocean, a sea, or a smaller body of water. Example archipelagos include the Aegean Islands (the o ...
comprises three large islands,
Mauritius Mauritius, officially the Republic of Mauritius, is an island country in the Indian Ocean, about off the southeastern coast of East Africa, east of Madagascar. It includes the main island (also called Mauritius), as well as Rodrigues, Ag ...
,
Réunion Réunion (; ; ; known as before 1848) is an island in the Indian Ocean that is an overseas departments and regions of France, overseas department and region of France. Part of the Mascarene Islands, it is located approximately east of the isl ...
, and
Rodrigues Rodrigues ( ; Mauritian Creole, Creole: ) is a Autonomous administrative division, autonomous Outer islands of Mauritius, outer island of the Republic of Mauritius in the Indian Ocean, about east of Mauritius. It is part of the Mascarene Isl ...
, plus a number of volcanic remnants in the tropics of the southwestern Indian Ocean, generally between 700 and 1,500 kilometres east of Madagascar. The
terrain Terrain (), alternatively relief or topographical relief, is the dimension and shape of a given surface of land. In physical geography, terrain is the lay of the land. This is usually expressed in terms of the elevation, slope, and orientati ...
includes a variety of
reef A reef is a ridge or shoal of rock, coral, or similar relatively stable material lying beneath the surface of a natural body of water. Many reefs result from natural, abiotic component, abiotic (non-living) processes such as deposition (geol ...
s,
atoll An atoll () is a ring-shaped island, including a coral rim that encircles a lagoon. There may be coral islands or cays on the rim. Atolls are located in warm tropical or subtropical parts of the oceans and seas where corals can develop. Most ...
s, and small islands. They present various
topographical Topography is the study of the forms and features of land surfaces. The topography of an area may refer to the landforms and features themselves, or a description or depiction in maps. Topography is a field of geoscience and planetary scienc ...
and
edaphic Edaphology (from Greek , ''edaphos'' 'ground' + , '' -logia'') is concerned with the influence of soils on living beings, particularly plants. It is one of two main divisions of soil science, the other being pedology. Edaphology includes the st ...
regions. On the largest islands these gave rise to endemism and unusual biodiversity. The climate is oceanic and
tropical The tropics are the regions of Earth surrounding the equator, where the sun may shine directly overhead. This contrasts with the temperate or polar regions of Earth, where the Sun can never be directly overhead. This is because of Earth's ax ...
. Mauritius is located 900 km east of
Madagascar Madagascar, officially the Republic of Madagascar, is an island country that includes the island of Madagascar and numerous smaller peripheral islands. Lying off the southeastern coast of Africa, it is the world's List of islands by area, f ...
. It has an area of 1,865 km2. The highest point is 828 meters above sea level. Mauritius is the most populous of the Mascarene Islands, with a population of 1,252,964. Réunion is located 150 km southwest of Mauritius. It is the largest of the islands, with an area of 2,512 km2.
Piton des Neiges The Piton des Neiges (Snow Peak) is a complex volcano on Réunion, one of the French volcanic islands in the Mascarene Archipelago in the southwestern Indian Ocean. It is located about east of Madagascar. Piton des Neiges is the highest poin ...
(3,069 m), an extinct volcano, is the highest peak on Réunion and in the islands.
Piton de la Fournaise Piton de la Fournaise (; ) is a shield volcano on the eastern side of Réunion island, a French overseas department and region, in the Indian Ocean. It is currently one of the most active volcanoes in the world, along with Kīlauea in the Haw ...
is an active volcano on Réunion which erupts frequently. Rodrigues is located 574 km east of Mauritius. It has an area of 108 km2, and reaches 393 meters elevation.
St. Brandon Saint Brandon (), also known as the Cargados Carajos Shoals, is a southwest Indian Ocean archipelago of sand banks, shoals and islets belonging to the Republic of Mauritius. It lies about northeast of the island of Mauritius. It consists of fiv ...
, also known as the
Cargados Carajos Saint Brandon (), also known as the Cargados Carajos Shoals, is a southwest Indian Ocean archipelago of sand banks, shoals and islets belonging to the Republic of Mauritius. It lies about northeast of the island of Mauritius. It consists of fiv ...
shoals, is a
coral atoll Corals are colonial marine invertebrates within the subphylum Anthozoa of the phylum Cnidaria. They typically form compact colonies of many identical individual polyps. Coral species include the important reef builders that inhabit tropical oce ...
group consisting of a barrier reef, shoals, and low islets. It is the remnant of one -or more- large volcanic islands sitting on the
Mauritia (microcontinent) Mauritia was a Precambrian microcontinent that was situated between India and Madagascar until their separation about 70 million years ago. Being initially attached to the Indian continent, Mauritia separated from it about 60 million years ago ...
which were submerged by rising tides. Today, the thirty or so islands that comprise
St Brandon Saint Brandon (), also known as the Cargados Carajos Shoals, is a southwest Indian Ocean archipelago of sand banks, shoals and islets belonging to the Republic of Mauritius. It lies about northeast of the island of Mauritius. It consists of fiv ...
form part of the Republic of
Mauritius Mauritius, officially the Republic of Mauritius, is an island country in the Indian Ocean, about off the southeastern coast of East Africa, east of Madagascar. It includes the main island (also called Mauritius), as well as Rodrigues, Ag ...
. Around seventeen of the uninhabited islands are administered by the 'Outer Islands Development Corporation' of Mauritius (an entity run by the Prime Minister of Mauritius through the Prime Minister's Office) whilst the remaining Thirteen Islands of St. Brandon are managed by the
Raphael Fishing Company The Raphaël Fishing Company Ltd is a Mauritian fishing company incorporated on 7 July 1927 in Port Louis, Mauritius. It is the second oldest commercial company in Mauritius, after Mauritius Commercial Bank (1828). The company is a fisheries co ...
from three permanently inhabited island bases of Ile Raphael,
L'île du Sud L'île du Sud (South Island, l'île Boisées) is an island located in the St. Brandon archipelago. It is one of the three islands (the others being Île Raphael and L'Île Coco) used as a base of operations for fishing activities by Raphael Fishin ...
(South Island, l'île Boisées) and
L'ÃŽle Coco L'ÃŽle Coco ('Coco Island') is one of the longest islands adjoining the inner lagoon of the St. Brandon archipelago. It is at times inhabited by fishermen as a base for the resident fishing company's fishing activities as well as for fly fishin ...
. The
Raphael Fishing Company The Raphaël Fishing Company Ltd is a Mauritian fishing company incorporated on 7 July 1927 in Port Louis, Mauritius. It is the second oldest commercial company in Mauritius, after Mauritius Commercial Bank (1828). The company is a fisheries co ...
was granted these Thirteen Islands of
St. Brandon Saint Brandon (), also known as the Cargados Carajos Shoals, is a southwest Indian Ocean archipelago of sand banks, shoals and islets belonging to the Republic of Mauritius. It lies about northeast of the island of Mauritius. It consists of fiv ...
under a 1901 Permanent Lease (
999-year lease A 999-year lease, under historic common law, is an essentially permanent lease of property. The lease locations are mainly in Britain, its former colonies, and the Commonwealth. A former colony, the Republic of Mauritius (''The Raphael Fishing Com ...
) which were adjudged as a
Permanent Grant A 999-year lease, under historic common law, is a permanent lease of property. Permanent lease locations are in Britain, its former colonies and the Commonwealth. A former colony, the Republic of Mauritius established a legal precedent through ...
by determination of the
Privy Council (United Kingdom) The Privy Council, formally His Majesty's Most Honourable Privy Council, is a privy council, formal body of advisers to the sovereign of the United Kingdom. Its members, known as privy counsellors, are mainly senior politicians who are curre ...
in 2008. These Thirteen islands of
St. Brandon Saint Brandon (), also known as the Cargados Carajos Shoals, is a southwest Indian Ocean archipelago of sand banks, shoals and islets belonging to the Republic of Mauritius. It lies about northeast of the island of Mauritius. It consists of fiv ...
on Permanent Grant are
L'île du Sud L'île du Sud (South Island, l'île Boisées) is an island located in the St. Brandon archipelago. It is one of the three islands (the others being Île Raphael and L'Île Coco) used as a base of operations for fishing activities by Raphael Fishin ...
(South Island, l'île Boisée), Petit Fou, l'Avocaire, l'île aux Fous,
L'île du Gouvernement l'île du Gouvernement is an island in the St. Brandon archipelago. The island is uninhabited, and mostly functions as a bird and turtle sanctuary. This island was one of those recommended as part of the ''St. Brandon Marine Protected Area'' (M ...
, Petit Mapou, Grand Mapou, La Baleine,
L'ÃŽle Coco L'ÃŽle Coco ('Coco Island') is one of the longest islands adjoining the inner lagoon of the St. Brandon archipelago. It is at times inhabited by fishermen as a base for the resident fishing company's fishing activities as well as for fly fishin ...
,
Île Raphael Île Raphael is an island in the Saint Brandon archipelago, a group of 30 outer islands of Mauritius. The island is named after Veuve Raphaël. Veuve Raphaël's husband was a sea captain and had installations on the corner of rue (route) des ...
, Verronge, l'île aux Bois and Baleines Rocks. There are several submerged banks or shoals: *
Saya de Malha Bank The Saya de Malha Bank (also the Sahia de Malha Bank, modern Portuguese: ''saia de malha'', English: ''mesh skirt'') or Mesh Skirt Bank, is one of the largest submerged ocean banks in the world, a part of the vast undersea Mascarene Plateau. Ge ...
is a large,
submerged bank An ocean bank, sometimes referred to as a fishing bank or simply bank, is a part of the seabed that is shallow compared to its surrounding area, such as a shoal or the top of an seamount, underwater hill. Somewhat like continental margin, conti ...
. Prehistorically it was a group of volcanic islands and was enjoined to the
Great Chagos Bank The Great Chagos Bank, in the Chagos Archipelago, about south of Maldives, is the largest atoll structure in the world, with a total area of . Islands Despite its enormous size, the Great Chagos Bank is largely a submarine structure. Ther ...
until continental drift pushed them apart. *
Soudan Banks The Soudan Banks are a series of underwater high points and reefs off the coast of Africa, known for their good fishing yields as fishing banks. They are administered by Mauritius. The five banks (actually a single feature) lie on the Mascarene Pla ...
are a group of low-lying banks on the
Mascarene Plateau The Mascarene Plateau is a submarine plateau in the Indian Ocean, north and east of Madagascar. The plateau extends approximately , from Seychelles in the north to Réunion in the south. The plateau covers an area of over of shallow water, wi ...
. *
Nazareth Bank Nazareth Bank is a large submerged bank in the Indian Ocean. Geography It lies about 1,040 km east of northern Madagascar and 280 km south of the Saya de Malha Bank. The closest land is Cargados Carajos shoals, a small and remote dependency of ...
is located just north of Cargados Carajos, and prehistorically they were a single geological feature. Today it is a large, shallow fishing bank. * Hawkins Bank is located on the northernmost point of the
Mascarene Plateau The Mascarene Plateau is a submarine plateau in the Indian Ocean, north and east of Madagascar. The plateau extends approximately , from Seychelles in the north to Réunion in the south. The plateau covers an area of over of shallow water, wi ...
.


Geology

The islands are
volcanic A volcano is commonly defined as a vent or fissure in the crust of a planetary-mass object, such as Earth, that allows hot lava, volcanic ash, and gases to escape from a magma chamber below the surface. On Earth, volcanoes are most often fo ...
in origin;
Saya de Malha The Saya de Malha Bank (also the Sahia de Malha Bank, modern Portuguese: ''saia de malha'', English: ''mesh skirt'') or Mesh Skirt Bank, is one of the largest submerged ocean banks in the world, a part of the vast undersea Mascarene Plateau. Ge ...
(35 mya) was the first of the Mascarene islands to rise out of the Indian Ocean due to the
Réunion hotspot The Réunion hotspot is a volcanic hotspot which currently lies under the island of Réunion in the Indian Ocean. The Chagos-Laccadive Ridge and the southern part of the Mascarene Plateau are volcanic traces of the Réunion hotspot. The hotspo ...
, followed by
Nazareth Bank Nazareth Bank is a large submerged bank in the Indian Ocean. Geography It lies about 1,040 km east of northern Madagascar and 280 km south of the Saya de Malha Bank. The closest land is Cargados Carajos shoals, a small and remote dependency of ...
(approximately 2 mya later), Soudan Bank and
Cargados Carajos Saint Brandon (), also known as the Cargados Carajos Shoals, is a southwest Indian Ocean archipelago of sand banks, shoals and islets belonging to the Republic of Mauritius. It lies about northeast of the island of Mauritius. It consists of fiv ...
. The youngest islands to form were Mauritius (7–10 mya), the oldest of the existing islands, created along with the undersea Rodrigues ridge. The islands of Rodrigues and Réunion were created in the last two million years. Réunion is the largest of the islands (2,500 km2), followed by Mauritius (1,900 km2) and Rodrigues (110 km2). Eventually, Saya de Malha, Nazareth and Soudan were completely submerged, Cargados Carajos remaining as a coral atoll. The Réunion hotspot was beginning to cool and Rodrigues came out as a small island. Réunion is home to the highest peaks in the Mascarenes, the
shield volcano A shield volcano is a type of volcano named for its low profile, resembling a shield lying on the ground. It is formed by the eruption of highly fluid (low viscosity) lava, which travels farther and forms thinner flows than the more viscous lava ...
es
Piton des Neiges The Piton des Neiges (Snow Peak) is a complex volcano on Réunion, one of the French volcanic islands in the Mascarene Archipelago in the southwestern Indian Ocean. It is located about east of Madagascar. Piton des Neiges is the highest poin ...
(3,069 m) and
Piton de la Fournaise Piton de la Fournaise (; ) is a shield volcano on the eastern side of Réunion island, a French overseas department and region, in the Indian Ocean. It is currently one of the most active volcanoes in the world, along with Kīlauea in the Haw ...
(2,525 m). Piton de la Fournaise, on the southeastern corner of Réunion, is one of the most active volcanoes in the world, erupting last in December 2021.
Piton de la Petite Rivière Noire Piton de la Petite Rivière Noire (''Little Black River Peak'') is the highest mountain on the island of Mauritius, in the Indian Ocean The Indian Ocean is the third-largest of the world's five oceanic divisions, covering or approximately 2 ...
(828 m) is the highest peak on Mauritius, and the gentle hills of Rodrigues rise to only 390 m. The
Mascarene Plateau The Mascarene Plateau is a submarine plateau in the Indian Ocean, north and east of Madagascar. The plateau extends approximately , from Seychelles in the north to Réunion in the south. The plateau covers an area of over of shallow water, wi ...
is an undersea plateau that extends approximately 2000 km, from the
Seychelles Seychelles (, ; ), officially the Republic of Seychelles (; Seychellois Creole: ), is an island country and archipelagic state consisting of 155 islands (as per the Constitution) in the Indian Ocean. Its capital and largest city, Victoria, ...
to Réunion. The plateau covers an area of over 115,000 km2 of shallow water, with depths ranging from 8 to 150 meters, plunging to 4000 m to the
abyssal plain An abyssal plain is an underwater plain on the deep ocean floor, usually found at depths between . Lying generally between the foot of a continental rise and a mid-ocean ridge, abyssal plains cover more than 50% of the Earth's surface. They ...
at its edges. The southern part of the plateau, including the
Saya de Malha Bank The Saya de Malha Bank (also the Sahia de Malha Bank, modern Portuguese: ''saia de malha'', English: ''mesh skirt'') or Mesh Skirt Bank, is one of the largest submerged ocean banks in the world, a part of the vast undersea Mascarene Plateau. Ge ...
,
Nazareth Bank Nazareth Bank is a large submerged bank in the Indian Ocean. Geography It lies about 1,040 km east of northern Madagascar and 280 km south of the Saya de Malha Bank. The closest land is Cargados Carajos shoals, a small and remote dependency of ...
,
Soudan Banks The Soudan Banks are a series of underwater high points and reefs off the coast of Africa, known for their good fishing yields as fishing banks. They are administered by Mauritius. The five banks (actually a single feature) lie on the Mascarene Pla ...
and
Cargados Carajos Saint Brandon (), also known as the Cargados Carajos Shoals, is a southwest Indian Ocean archipelago of sand banks, shoals and islets belonging to the Republic of Mauritius. It lies about northeast of the island of Mauritius. It consists of fiv ...
Shoals (Saint Brandon) (then one large island), was formed by the Réunion hotspot. These were once volcanic islands, much like Mauritius and Réunion, which have now sunk or eroded to below sea level or, in the case of the Cargados Carajos, to low coral islands. The Saya de Malha Bank formed 35 million years ago, and the Nazareth Bank and the Cargados Carajos shoals after that.
Limestone Limestone is a type of carbonate rock, carbonate sedimentary rock which is the main source of the material Lime (material), lime. It is composed mostly of the minerals calcite and aragonite, which are different Polymorphism (materials science) ...
banks found on the plateau are the remnants of
coral reef A coral reef is an underwater ecosystem characterized by reef-building corals. Reefs are formed of colonies of coral polyps held together by calcium carbonate. Most coral reefs are built from stony corals, whose polyps cluster in group ...
s, indicating that the plateau was a succession of islands. Some of the banks may have been islands as recently as 18,000–6,000 years ago, when sea levels were as much as 130 meters lower during the most recent
ice age An ice age is a long period of reduction in the temperature of Earth's surface and atmosphere, resulting in the presence or expansion of continental and polar ice sheets and alpine glaciers. Earth's climate alternates between ice ages, and g ...
.


History

The early colonial history of the islands, like that of the Caribbean, featured a succession of takeovers between rival powers: the Portuguese, Dutch, French and British all ruled some or all of the islands. Around 1507, the explorer
Diogo Fernandes Pereira Diogo Fernandes Pereira, sometimes called simply Diogo Fernandes, was a Portuguese 16th-century navigator, originally from Setúbal, Portugal. Diogo Fernandes was the first known European captain to visit the island of Socotra in 1503 and the di ...
discovered the island group. The area remained under nominal Portuguese rule until
Étienne de Flacourt Étienne de Flacourt (1607–1660) was a French governor of Madagascar, born in Orléans in 1607. He was named governor of Madagascar by the French East India Company in 1648. Flacourt restored order among the French soldiers, who had mutinied ...
arrived with a French naval squadron and took possession in 1649. From 4 June 1735 to 23 March 1746, a single French Mascarene Islands chartered colony under one ''
gouverneur général Governor-general (plural governors-general), or governor general (plural governors general), is the title of an official, most prominently associated with the British Empire. In the context of the governors-general and former British colonies, ...
'' (governor general) contained Isle de France (now
Mauritius Mauritius, officially the Republic of Mauritius, is an island country in the Indian Ocean, about off the southeastern coast of East Africa, east of Madagascar. It includes the main island (also called Mauritius), as well as Rodrigues, Ag ...
),
Île Bourbon Ile or ILE may refer to: Ile * Ile, a Puerto Rican singer * Ile District (disambiguation), multiple places * Ilé-Ifẹ̀, an ancient Yoruba city in south-western Nigeria * Interlingue (ISO 639:ile), a planned language * Isoleucine, an amino aci ...
(Réunion) and Séchelles (Seychelles). On 14 July 1767 this became a French crown colony, still under one governor general. From 3 February 1803 until 2 September 1810 the French colony of ''Indes-Orientales'', under a '' capitaine général'' (captain-general), included Réunion and (nominally) the Seychelles.


Mauritius

It is postulated from navigational charts, such as the Cantino planisphere of 1502, that Arab sailors first discovered Mauritius around 975, calling it Dina Arobi (''abandoned island''). The earliest confirmed discovery on record was in 1507 by Portuguese sailors. The Dutch took physical possession in 1598, establishing a succession of short-lived settlements over a period of about 120 years, before abandoning their efforts in 1710. France took control in 1715, renaming it ''Isle de France''. In 1810, the United Kingdom invaded the island to protect its East India ships coming around the Cape of Good Hope from French pirates. Four years later, at the Treaty of Paris, France ceded Mauritius and its dependencies to the United Kingdom. Under
British Mauritius Mauritius was a Crown colony off the southeast coast of Africa. Formerly part of the French colonial empire, British rule in Mauritius was established ''de facto'' with the invasion of Isle de France in November 1810, and ''de jure'' by the sub ...
(1810-1968), the island successfully developed as a sugar cane-based plantation economy and colony until independence in 1968.


Rodrigues

Rodrigues was first discovered by the
Arabs Arabs (,  , ; , , ) are an ethnic group mainly inhabiting the Arab world in West Asia and North Africa. A significant Arab diaspora is present in various parts of the world. Arabs have been in the Fertile Crescent for thousands of yea ...
but named after Portuguese navigator
Diogo Rodrigues Dom Diogo de Azevedo Roiz (né Rodrigues; – 21 April 1577) was a Portuguese explorer of the Indian Ocean who sailed as an ordinary helmsmanAuguste Toussaint, ''History of the Indian Ocean'' (Chicago: University Press, 1966), pp. 109 un ...
. It was under Dutch control in 1601 and settled by the French in 1691. Britain took possession of Rodrigues in 1809. When Mauritius gained independence in 1968, Rodrigues was forcefully joined to it. Rodrigues remains an autonomous region of Mauritius.


Réunion

Réunion was discovered first by the Arabs then by the Portuguese, who named it Santa Apolónia. It was then occupied by the French as part of Mauritius. It was first inhabited by French mutineers who arrived on the island between 1646 and 1669. It was given its current name in 1793. From 1810 to 1815 it was held by the British, before being returned to France. Réunion became an
overseas department The overseas departments and regions of France (, ; DROM) are the five departments and regions of the French Republic which are located outside European France (also known as " metropolitan France"). These overseas entities have exactly the s ...
of France in 1946.


Climate

The climate of the islands is tropical. At low elevations summer temperatures (December to April) average 30 Â°C. Winter temperatures (May to November) are generally cooler, around 25 Â°C. Temperatures are cooler in the mountains, averaging 18 Â°C. During the winter short-lived snow can fall on Réunion's high peaks. Southeasterly
trade winds The trade winds or easterlies are permanent east-to-west prevailing winds that flow in the Earth's equatorial region. The trade winds blow mainly from the northeast in the Northern Hemisphere and from the southeast in the Southern Hemisphere ...
blow throughout the year. Rainfall is generally higher on the windward sides of the islands. On Mauritius average annual rainfall in the lowlands varies from 1905 mm on the windward southeast coast to 890 mm on the leeward side. Rainfall in the mountains is higher, varying from 2540 mm to 4445 mm annually.
Tropical cyclone A tropical cyclone is a rapidly rotating storm system with a low-pressure area, a closed low-level atmospheric circulation, strong winds, and a spiral arrangement of thunderstorms that produce heavy rain and squalls. Depending on its locat ...
s occur occasionally, bringing heavy rain and high winds, and causing erosion and landslides.


Flora

The habitats of the Macarenes vary with the islands' size, topography, age, and proximity to Madagascar, the nearest major land mass. As is common among remote islands, the Mascarene fauna and flora display a high degree of
endemism Endemism is the state of a species being found only in a single defined geographic location, such as an island, state, nation, country or other defined zone; organisms that are indigenous to a place are not endemic to it if they are also foun ...
; they include over a thousand species of which several hundred are endemic. The Mascarene islands form a distinct
ecoregion An ecoregion (ecological region) is an ecological and geographic area that exists on multiple different levels, defined by type, quality, and quantity of environmental resources. Ecoregions cover relatively large areas of land or water, and c ...
known as the Mascarene forests. Until Europeans first settled the islands in the sixteenth century the Mascarenes are not known to have harbored any human population, and the islands' wildlife was still flourishing during the early days of settlement. Today much of the natural vegetation is gone, and there are many
introduced species An introduced species, alien species, exotic species, adventive species, immigrant species, foreign species, non-indigenous species, or non-native species is a species living outside its native distributional range, but which has arrived ther ...
on the islands. In prehistoric times the islands were covered in a diverse range of
tropical moist broadleaf forest Tropical and subtropical moist broadleaf forests (TSMF), also known as tropical moist forest, is a subtropical and tropical forest habitat type defined by the World Wide Fund for Nature (WWF). Description TSMF is generally found in large ...
s. The plant communities are not homogeneous, and comprise at least five fairly distinct vegetation zones that reflect variations in altitude and in moisture regime. These include dry lowland forests, semi-dry sclerophyllous forests, lowland rainforests, montane cloud forests, and high-elevation heathlands on Réunion.Thébaud, Christophe & Warren, Ben & Cheke, Anthony & Strasberg, Dominique. (2009). Mascarene Islands, Biology

/ref> Coastal habitats include beach vegetation, coastal
wetland A wetland is a distinct semi-aquatic ecosystem whose groundcovers are flooded or saturated in water, either permanently, for years or decades, or only seasonally. Flooding results in oxygen-poor ( anoxic) processes taking place, especially ...
s, and
swamp forest Freshwater swamp forests, or flooded forests, are forests which are inundated with freshwater, either permanently or seasonally. They normally occur along the lower reaches of rivers and around freshwater lakes. Freshwater swamp forests are foun ...
s, grading into lowland rain forest on the windward sides of the islands and lowland dry forests to leeward. The dry lowland forests are found on the leeward sides of the islands, from sea level to 200 metres in areas with less than 1000 mm of average annual rainfall. Palms are the dominant trees, including species of ''
Latania ''Latania'', commonly known as latan palm or latania palm, is a genus of flowering plant in the Arecaceae, palm tree family (biology), family, native to the Mascarene Islands in the western Indian Ocean. Species The genus contains three species ...
'' and '' Dictyosperma album'', along with the palm-like screw-pines (''
Pandanus ''Pandanus'' is a genus of monocots with about 578 accepted species. They are palm-like, dioecious trees and shrubs native to the Old World tropics and subtropics. Common names include pandan, screw palm and screw pine. The genus is classified ...
'' spp.). Semi-dry
sclerophyllous Sclerophyll is a type of vegetation that is adapted to long periods of dryness and heat. The plants feature hard leaves, short internodes (the distance between leaves along the stem) and leaf orientation which is parallel or oblique to direct ...
forests A forest is an ecosystem characterized by a dense community of trees. Hundreds of definitions of forest are used throughout the world, incorporating factors such as tree density, tree height, land use, legal standing, and ecological functio ...
are found between the coast and 360 metres elevation on Mauritius and Rodrigues, and from 200 to 750 metres elevation on the western slopes of Réunion, where only small patches of such forest now remain. Average annual rainfall in the semi-dry forests ranges from 1000 to 1500 mm. Tree genera and families well-represented in the semi-dry forests include
ebony Ebony is a dense black/brown hardwood, coming from several species in the genus '' Diospyros'', which also includes the persimmon tree. A few ''Diospyros'' species, such as macassar and mun ebony, are dense enough to sink in water. Ebony is fin ...
species in the genus ''
Diospyros ''Diospyros'' is a genus of over 700 species of deciduous and evergreen trees and shrubs. The majority are native to the tropics, with only a few species extending into temperate regions. Individual species valued for their hard, heavy, dark timb ...
'' (
Ebenaceae The Ebenaceae are a family of flowering plants belonging to order Ericales. The family includes ebony and persimmon among about 768 species of trees and shrubs. It is distributed across the tropical and warmer temperate regions of the world. It i ...
), '' Pleurostylia'' (
Celastraceae The Celastraceae (staff-vine or bittersweet) are a family of 98 genera and 1,350 species of herbs, vines, shrubs and small trees, belonging to the order Celastrales. The great majority of the genera are tropical, with only ''Celastrus'' (the staf ...
), ''
Foetidia ''Foetidia'' is a genus of flowering plants first described as a genus in 1788. Most authors place the genus in the family Lecythidaceae, the sole genus in the subfamily Foetidioideae, but some prefer to treat it as a distinct family, the Foeti ...
'' (
Lecythidaceae The Lecythidaceae ( ) comprise a family of about 20 genera and 250–300 species of woody plants native to tropical South America, Africa (including Madagascar), Asia and Australia. Well known members of the family include the cannonball tree ...
), ''Olea europaea'' subsp. ''cuspidata'' (
Oleaceae Oleaceae, also known as the olive family or sometimes the lilac family, is a taxonomic family of flowering shrubs, trees, and a few lianas in the order Lamiales. It presently comprises 28 genera, one of which is recently extinct.Peter S. Gree ...
), '' Cossinia pinnata'' (
Sapindaceae The Sapindaceae are a family (biology), family of flowering plants in the order Sapindales known as the soapberry family. It contains 138 genera and 1,858 accepted species. Examples include Aesculus, horse chestnut, maples, ackee and lychee. The ...
) ''
Dombeya ''Dombeya'' is a flowering plant genus. Traditionally included in the family Sterculiaceae, it is included in the expanded Malvaceae in the APG and most subsequent systematics. These plants are known by a number of vernacular names which sometim ...
'' (
Malvaceae Malvaceae (), or the mallows, is a family of flowering plants estimated to contain 244 genera with 4225 known species. Well-known members of economic importance include Theobroma cacao, cacao, Cola (plant), cola, cotton, okra, Hibiscus sabdariffa, ...
), '' Terminalia bentzoe'' (
Combretaceae The Combretaceae, often called the white mangrove family, are a family of flowering plant Flowering plants are plants that bear flowers and fruits, and form the clade Angiospermae (). The term angiosperm is derived from the Ancient Greek, G ...
), and ''
Sideroxylon boutonianum ''Sideroxylon'' is a genus of trees in the family Sapotaceae described as a genus by Linnaeus in 1753. They are collectively known as bully trees. The generic name is derived from the Greek words σιδηÏος (''sideros''), meaning "iron", and ...
, Sideroxylon borbonicum'', and various species of ''
Mimusops ''Mimusops'' is a genus of plants in the family Sapotaceae, described as a genus by Carl Linnaeus, Linnaeus in 1753. ''Mimusops'' is native to tropical and subtropical regions of Asia, Africa, Australia, and various oceanic islands.Allaby, Micha ...
'' in family
Sapotaceae 240px, '' Madhuca longifolia'' var. ''latifolia'' in Narsapur, Medak district, India The Sapotaceae are a family of flowering plants belonging to the order (biology)">order Ericales">family (biology)">family of flowering plants belonging to th ...
. Shrubs in the semi-dry forests include several spectacular endemic species of ''
Hibiscus ''Hibiscus'' is a genus of flowering plants in the Malva, mallow family, Malvaceae. The genus is quite large, comprising List of Hibiscus species, several hundred species that are Native plant, native to warm temperate, Subtropics, subtropical ...
'' (
Malvaceae Malvaceae (), or the mallows, is a family of flowering plants estimated to contain 244 genera with 4225 known species. Well-known members of economic importance include Theobroma cacao, cacao, Cola (plant), cola, cotton, okra, Hibiscus sabdariffa, ...
), ''
Zanthoxylum ''Zanthoxylum'' is a genus of about 250 species of deciduous and evergreen trees, shrubs and climbers in the Family (biology), family Rutaceae that are native to warm temperate and Subtropics, subtropical areas worldwide. It is the type gen ...
'' (
Rutaceae The Rutaceae () is a family (biology), family, commonly known as the rueRUTACEAE
in Bo ...
), ''
Obetia ficifolia ''Obetia'' is a genus of dioecious plants in the family Urticaceae, with stinging hairs. The genus contains the following species: *''Obetia carruthersiana'' (Hiern) Rendle *''Obetia ficifolia'' Gaudich. *''Obetia madagascariensis'' (Juss. ''ex'' ...
'' (
Urticaceae The Urticaceae are a family, the nettle family, of flowering plants. The family name comes from the genus ''Urtica''. The Urticaceae include a number of well-known and useful plants, including nettles in the genus ''Urtica'', ramie (''Boehmeria ...
), and ''
Scolopia heterophylla ''Scolopia'' is an Old World genus of plants in the family Salicaceae. Species include: * ''Scolopia braunii'' (Klotzsch) & Sleumer - an Australian rainforest tree *''Scolopia buxifolia'' Gagnep. *''Scolopia chinensis'' (Lour.) Clos *''Scolopi ...
'' (
Flacourtiaceae The Flacourtiaceae is a defunct family of flowering plants whose former members have been scattered to various families, mostly to the Achariaceae and Salicaceae. It was so vaguely defined that hardly anything seemed out of place there and it bec ...
). The lowland rain forests are characterized by dense
evergreen In botany, an evergreen is a plant which has Leaf, foliage that remains green and functional throughout the year. This contrasts with deciduous plants, which lose their foliage completely during the winter or dry season. Consisting of many diffe ...
forests, composed of diverse species creating a canopy of 30 meters or more in height. They are found on the Mauritius' eastern lowlands from the coast to 800–900 metres elevation, and above 360 m on the western slopes. On Réunion lowland rain forests are found between 750 and 1100 m. They occur in areas with average annual rainfall of 1500 to 6000 mm. Characteristic trees include species of ''
Mimusops ''Mimusops'' is a genus of plants in the family Sapotaceae, described as a genus by Carl Linnaeus, Linnaeus in 1753. ''Mimusops'' is native to tropical and subtropical regions of Asia, Africa, Australia, and various oceanic islands.Allaby, Micha ...
'' and '' Labourdonnaisia'' (
Sapotaceae 240px, '' Madhuca longifolia'' var. ''latifolia'' in Narsapur, Medak district, India The Sapotaceae are a family of flowering plants belonging to the order (biology)">order Ericales">family (biology)">family of flowering plants belonging to th ...
), '' Hernandia mascarenensis'' (
Hernandiaceae The Hernandiaceae are a family of flowering plants (angiosperms) in the order Laurales. Consisting of five genera with about 58 known species, they are distributed over the world's tropical areas, some of them widely distributed in coastal areas ...
), ''
Calophyllum ''Calophyllum'' is a genus of tropical flowering plants in the family Calophyllaceae. They are mainly distributed in Asia, with some species in Africa, the Americas, Australasia, and the Pacific Islands. History Members of the genus ''Calophyll ...
'' (
Clusiaceae The Clusiaceae or Guttiferae Juss. (1789) (''nom. alt. et cons.'' = alternative and valid name) are a family (biology), family of plants including 13 genera and ca 750 species. Several former members of Clusiacae are now placed in Calophyllaceae ...
), and species of ''
Syzygium ''Syzygium'' () is a genus of flowering plants that belongs to the myrtle family, Myrtaceae. The genus comprises about 1200 species, and has a native range that extends from Africa and Madagascar through southern Asia east through the Pacific. I ...
'', ''
Eugenia ''Eugenia'' is a genus of flowering plants in the myrtle family Myrtaceae. It has a worldwide, although highly uneven, distribution in tropical and subtropical regions. The bulk of the approximately 1,100 species occur in the New World tropics, ...
'', ''
Sideroxylon ''Sideroxylon'' is a genus of trees in the family (biology), family Sapotaceae described as a genus by Carl Linnaeus, Linnaeus in 1753. They are collectively known as bully trees. The generic name is derived from the Greek language, Greek words ...
'', and '' Monimiastrum'' (
Myrtaceae Myrtaceae (), the myrtle family, is a family of dicotyledonous plants placed within the order Myrtales. Myrtle, pÅhutukawa, bay rum tree, clove, guava, acca (feijoa), allspice, and eucalyptus are some notable members of this group. All ...
). Characteristic shrubs include species of ''
Gaertnera ''Gaertnera'' is a genus of flowering plants in the family Rubiaceae. There are at least 85 species distributed across the Old World tropics from Africa to Asia.
'', '' Chassalia'', ''
Bertiera ''Bertiera'' is a genus of flowering plants in the family Rubiaceae. It comprises 57 species with most known from tropical Africa, five known from various :Indian Ocean islands and five found in the tropics of the Americas. Taxonomy The genus ...
'', and ''
Coffea ''Coffea'' is a genus of flowering plants in the family Rubiaceae. ''Coffea'' species are shrubs or small trees native to tropical and southern Africa and tropical Asia. The seeds of some species, called coffee beans, are used to flavor variou ...
'' in family
Rubiaceae Rubiaceae () is a family (biology), family of flowering plants, commonly known as the coffee, madder, or bedstraw family. It consists of terrestrial trees, shrubs, lianas, or herbs that are recognizable by simple, opposite leaves with Petiole ( ...
. Other plants include
bamboo Bamboos are a diverse group of mostly evergreen perennial plant, perennial flowering plants making up the subfamily (biology), subfamily Bambusoideae of the grass family Poaceae. Giant bamboos are the largest members of the grass family, in th ...
s like ''
Nastus borbonicus ''Nastus'' is a genus of slender, erect, scrambling or climbing bamboos in the grass family. It is native to Southeast Asia, Papuasia, and certain islands in the Indian Ocean (Madagascar and Réunion). ;Species ;formerly included see ''Arundin ...
'', numerous species of orchids (e.g., ''
Angraecum ''Angraecum'', also known as comet orchid, is a genus of the family Orchidaceae native to tropical and South Africa, as well as Sri Lanka. It contains 223 species. Etymology Despite the genus's distribution being largely confined to Africa and i ...
'', ''
Bulbophyllum ''Bulbophyllum'' is a genus of mostly Epiphyte, epiphytic and Lithophyte, lithophytic orchids in the Family (biology), family Orchidaceae. It is the largest genus in the orchid family and one of the List of the largest genera of flowering plants, ...
'') and ferns e.g., ''
Asplenium ''Asplenium'' is a genus of about 700 species of ferns, often treated as the only genus in the family (biology), family Aspleniaceae, though other authors consider ''Hymenasplenium'' separate, based on molecular phylogenetic analysis of DNA seque ...
'', ''
Hymenophyllum ''Hymenophyllum'' is a genus of ferns in the family Hymenophyllaceae. Its name means "membranous leaf", referring to the very thin translucent tissue of the fronds, which gives rise to the common name filmy fern for this and other thin-leaved fer ...
'', ''
Trichomanes ''Trichomanes'' is a genus of ferns in the family Hymenophyllaceae, termed bristle ferns. The circumscription of the genus is disputed. All ferns in the genus are filmy ferns, with leaf tissue typically 2 cells thick. This thinness generally nec ...
'', '' Elaphoglossum'', and ''
Marattia ''Marattia'' is a small genus of basal, large, fleshy eusporangiate ferns. It is the type genus of the family Marattiaceae, order Marattiales and class Marattiopsida. Formerly considered to be a much larger genus, genetic analysis has shown t ...
fraxinea''. The
cloud forest A cloud forest, also called a water forest, primas forest, or tropical montane cloud forest, is a generally tropical or subtropical, evergreen, Montane forest, montane, Tropical and subtropical moist broadleaf forests, moist forest characteri ...
s are evergreen rain forests found on mountain slopes with high rainfall. They occur on Réunion between 800 and 1900 metres elevation on eastern slopes with an average annual rainfall of 2000–10,000 mm, and between 1100 and 2000 metres on western slopes with an average annual rainfall of 2000–3000 mm. By 2005, relatively intact cloud forests still covered approximately 44,000 ha on Réunion. On Mauritius they are restricted to a small area of Montagne Cocotte in the island's southwest, above 750 metres elevation and with average annual rainfall of 4500–5500 mm. Trees form a dense canopy 6 to 10 metres high. Typical canopy trees include species of ''
Dombeya ''Dombeya'' is a flowering plant genus. Traditionally included in the family Sterculiaceae, it is included in the expanded Malvaceae in the APG and most subsequent systematics. These plants are known by a number of vernacular names which sometim ...
'' on Réunion, and species of ''
Monimia ''Monimia'' is a genus of trees of the family Monimiaceae, within the major group of ''Angiosperms'' (flowering plants). It is endemic to the Mascarene Islands The Mascarene Islands (, ) or Mascarenes or Mascarenhas Archipelago is a group of ...
'' and ''Tambourissa'' (
Monimiaceae The Monimiaceae is a family (biology), family of flowering plants in the magnoliid Order (biology), order Laurales.Peter F. Stevens (2001 onwards). "Monimiaceae" At: Angiosperm Phylogeny Website. At: Botanical Databases At: Missouri Botanical Gar ...
) on both islands. There is an understory of small trees and shrubs including species of ''
Psiadia ''Psiadia'', commonly known as daisy trees, is a genus of mostly woody Asian and African plants in the tribe Astereae within the family Asteraceae. The genus is distributed throughout the Western Indian Ocean, with species reported from continent ...
'' (
Asteraceae Asteraceae () is a large family (biology), family of flowering plants that consists of over 32,000 known species in over 1,900 genera within the Order (biology), order Asterales. The number of species in Asteraceae is rivaled only by the Orchi ...
) and ''
Melicope ''Melicope'' is a genus of about 240 species of shrubs and trees in the family Rutaceae, occurring from the Hawaiian Islands across the Pacific Ocean to tropical Asia, Australia and New Zealand. Plants in the genus ''Melicope'' have simple or ...
'' (
Rutaceae The Rutaceae () is a family (biology), family, commonly known as the rueRUTACEAE
in Bo ...
). The forests are rich in
epiphyte An epiphyte is a plant or plant-like organism that grows on the surface of another plant and derives its moisture and nutrients from the air, rain, water (in marine environments) or from debris accumulating around it. The plants on which epiphyt ...
s (orchids, ferns, mosses, lichens), emergent
tree fern Tree ferns are arborescent (tree-like) ferns that grow with a trunk (botany), trunk elevating the fronds above ground level, making them trees. Many extant tree ferns are members of the order Cyatheales, to which belong the families Cyatheaceae ( ...
s (''
Cyathea ''Cyathea'' is a genus of tree ferns, the type genus of the fern order Cyatheales. The genus name ''Cyathea'' is derived from the Greek ''kyatheion'', meaning "little cup", and refers to the cup-shaped sori on the underside of the fronds. ...
'' species), and, originally the palms ''
Acanthophoenix rubra ''Acanthophoenix rubra'', the barbel palm, is a critically endangered Arecaceae, palm endemic to Mauritius, Rodrigues, and La Reunion that is prized for its edible palm hearts. Naming and description It is also known as the red- or yellow- Bar ...
'', but poaching has wiped out palms in many areas of Réunion. Three monodominant plant communities are also found in the cloud forests – forests of ''
Acacia heterophylla ''Acacia heterophylla'', the highland tamarind,Forestry Abstracts, University of Oxford Commonwealth Forestry Bureau, p.327, 1939 is a tree (or shrub in its higher places) Endemism, endemic to Réunion, Réunion island where it is commonly named ...
'' (Fabaceae) as the canopy tree, which are similar to the ''
Acacia koa ''Acacia koa'', commonly known as koa, is a species of Flowering plant, flowering tree in the Family (biology), family Fabaceae. It is Endemism, endemic to the Hawaiian Islands, where it is the second most common tree. The highest populations ar ...
'' forests in
Hawaii Hawaii ( ; ) is an island U.S. state, state of the United States, in the Pacific Ocean about southwest of the U.S. mainland. One of the two Non-contiguous United States, non-contiguous U.S. states (along with Alaska), it is the only sta ...
, thickets dominated by '' Erica reunionensis'' (
Ericaceae The Ericaceae () are a Family (biology), family of flowering plants, commonly known as the heath or heather family, found most commonly in acidic and infertile growing conditions. The family is large, with about 4,250 known species spread acros ...
), and hyperhumid forests of screw-pine (''
Pandanus montanus ''Pandanus montanus'' is a species of monocots in the genus ''Pandanus'', that is found in wet forest at an elevation of 400–1700 meters. Description It is a small, slender tree of 3–5 m in height. Its fruit-head is initially held up erect ...
''). Subalpine shrub occurs on the volcanoes on Réunion, above the treeline at 1800 to 2000 metres elevation. Average annual rainfall is from 2000 to 6000 mm, and frosts occur regularly in winter. Characteristic shrubs include ''Erica reunionensis'' and '' Erica galioides'' (Ericaceae), ''
Stoebe passerinoides ''Stoebe'' is a genus of plants in the tribe Gnaphalieae within the family Asteraceae. The genus is confined to the Cape Provinces of South Africa. ; Species ; formerly included A few dozen species now regarded as members of other genera: '' Di ...
'' and species of ''
Hubertia ''Hubertia'' is a genus of flowering plants in the daisy family Asteraceae () is a large family of flowering plants that consists of over 32,000 known species in over 1,900 genera within the order Asterales. The number of species in Astera ...
'' and ''
Psiadia ''Psiadia'', commonly known as daisy trees, is a genus of mostly woody Asian and African plants in the tribe Astereae within the family Asteraceae. The genus is distributed throughout the Western Indian Ocean, with species reported from continent ...
'' (Asteraceae), and '' Phylica nitida'' (
Rhamnaceae The Rhamnaceae are a large Family (biology), family of flowering plants, mostly trees, shrubs, and some vines, commonly called the buckthorn family. Rhamnaceae is included in the order Rosales. The family contains about 55 genera and 950 specie ...
). Herbaceous endemic species include '' Heterochaenia rivalsii'' (Campanulaceae), ''Eriotrix commersonii'' (Asteraceae), and ''
Cynoglossum borbonicum ''Cynoglossum'' is a genus of small-flowered plants in the Family (biology), family Boraginaceae (borage family). It includes 81 species native to Eurasia, Africa, New Guinea, and Australia. ''Cynoglossum officinale'', the common hound's-tongue, ...
'' (
Boraginaceae Boraginaceae, the Borago, borage or forget-me-not family, includes about 2,000 species of shrubs, trees, and herbs in 146 to 154 genus, genera with a worldwide distribution. The APG IV system from 2016 classifies the Boraginaceae as single famil ...
). Recent volcanic deposits on Réunion's volcanic summits are covered by sparse grasslands rich in endemic grasses, including ''
Festuca borbonica ''Festuca'' (fescue) is a genus of flowering plants belonging to the grass family Poaceae (subfamily Pooideae). They are evergreen or herbaceous perennial tufted grasses with a height range of and a cosmopolitan distribution, occurring on every ...
'', ''
Agrostis salaziensis ''Agrostis'' (bent or bentgrass) is a large and very nearly cosmopolitan genus of plants in the grass family, found in nearly all the countries in the world. It has been bred as a GMO creeping bent grass. Species , Plants of the World Online a ...
'', and ''
Cenchrus afer ''Cenchrus'' is a widespread genus of plants in the grass family, native to tropical and warm temperate regions of the world. Its species are native to many countries in Asia, Africa, Australia, the Americas, and various oceanic islands. Commo ...
''. along with orchids like '' Disa borbonica''. Ericoid thickets and thickets of the small tree '' Sophora denudata'' (
Fabaceae Fabaceae () or Leguminosae,International Code of Nomen ...
) are found on weathered volcanic substrates. Most of the indigenous Mascarene
flora Flora (: floras or florae) is all the plant life present in a particular region or time, generally the naturally occurring (indigenous (ecology), indigenous) native plant, native plants. The corresponding term for animals is ''fauna'', and for f ...
and fauna are thought to have descended originally from Madagascan and
Africa Africa is the world's second-largest and second-most populous continent after Asia. At about 30.3 million km2 (11.7 million square miles) including adjacent islands, it covers 20% of Earth's land area and 6% of its total surfac ...
n ancestors. The four largest families present in the Mascarene Islands are
Myrtaceae Myrtaceae (), the myrtle family, is a family of dicotyledonous plants placed within the order Myrtales. Myrtle, pÅhutukawa, bay rum tree, clove, guava, acca (feijoa), allspice, and eucalyptus are some notable members of this group. All ...
,
Rubiaceae Rubiaceae () is a family (biology), family of flowering plants, commonly known as the coffee, madder, or bedstraw family. It consists of terrestrial trees, shrubs, lianas, or herbs that are recognizable by simple, opposite leaves with Petiole ( ...
,
Orchidaceae Orchids are plants that belong to the family (biology), family Orchidaceae (), a diverse and widespread group of flowering plants with blooms that are often colourful and fragrant. Orchids are cosmopolitan distribution, cosmopolitan plants that ...
, and
Euphorbiaceae Euphorbiaceae (), the spurge family, is a large family of flowering plants. In English, they are also commonly called euphorbias, which is also the name of Euphorbia, the type genus of the family. Most spurges, such as ''Euphorbia paralias'', ar ...
, which comprise between 193 and 223 species each, for a total of 831 species, or 26.9% of the flowering plant flora. Another seven families contain 80 or more species each:
Poaceae Poaceae ( ), also called Gramineae ( ), is a large and nearly ubiquitous family of monocotyledonous flowering plants commonly known as grasses. It includes the cereal grasses, bamboos, the grasses of natural grassland and species cultivate ...
,
Apocynaceae Apocynaceae (, from '' Apocynum'', Greek for "dog-away") is a family of flowering plants that includes trees, shrubs, herbs, stem succulents, and vines, commonly known as the dogbane family, because some taxa were used as dog poison. Notable mem ...
,
Cyperaceae The Cyperaceae () are a family of graminoid (grass-like), monocotyledonous flowering plants known as wikt:sedge, sedges. The family (biology), family is large; botanists have species description, described some 5,500 known species in about 90 ...
,
Cunoniaceae Cunoniaceae is a family of 27 genera and about 335 species of woody plants in the order Oxalidales, mostly found in the tropical and wet temperate regions of the Southern Hemisphere. The greatest diversity of genera are in Australia and Tasmania ...
,
Rutaceae The Rutaceae () is a family (biology), family, commonly known as the rueRUTACEAE
in Bo ...
,
Araliaceae The Araliaceae are a family of flowering plants composed of about 43 genera and around 1500 species consisting of primarily woody plants and some herbaceous plants commonly called the ginseng family. The morphology of Araliaceae varies widely ...
, and
Sapotaceae 240px, '' Madhuca longifolia'' var. ''latifolia'' in Narsapur, Medak district, India The Sapotaceae are a family of flowering plants belonging to the order (biology)">order Ericales">family (biology)">family of flowering plants belonging to th ...
, representing an additional ca. 660 species in all, 21.3% of the angiosperm flora. Ferns are prominent components of the biotas of the islands, especially in the tropical forest. Most ferns disperse easily via ornithochory of their spores, allowing fairly frequent colonization from Madagascar and exchange among the Mascarene islands. The Mascarenes are home to many endemic species of
Dombeyoideae Dombeyoideae is a widely distributed subfamily of the Malvaceae, as proposed by the APG. Most of the plants placed here were once assembled with more or less related genera in the paraphyletic Sterculiaceae; a lesser number were placed in the Ti ...
, the
monotypic In biology, a monotypic taxon is a taxonomic group (taxon) that contains only one immediately subordinate taxon. A monotypic species is one that does not include subspecies or smaller, infraspecific taxa. In the case of genera, the term "unisp ...
genus '' Psiloxylon'' ('' Psiloxylon mauritianum''), and members of the family
Monimiaceae The Monimiaceae is a family (biology), family of flowering plants in the magnoliid Order (biology), order Laurales.Peter F. Stevens (2001 onwards). "Monimiaceae" At: Angiosperm Phylogeny Website. At: Botanical Databases At: Missouri Botanical Gar ...
(the genera ''
Monimia ''Monimia'' is a genus of trees of the family Monimiaceae, within the major group of ''Angiosperms'' (flowering plants). It is endemic to the Mascarene Islands The Mascarene Islands (, ) or Mascarenes or Mascarenhas Archipelago is a group of ...
'' and ''
Trochetia ''Trochetia'' is a subgenus of flowering plants, now placed in the genus '' Ruizia'' and the family Malvaceae (formerly in the Sterculiaceae, but genera now usually subsumed in the Dombeyoideae and other subfamilies). Species are endemic to the M ...
'', and species of ''
Tambourissa ''Tambourissa'' is a genus of plant in family Monimiaceae. Its range includes Madagascar, the Comoro Islands, Réunion, and Mauritius. It contains the following accepted species, according to ThePlantList.org: * '' Tambourissa alaticarpa'' Lorenc ...
'') and
Escalloniaceae Escalloniaceae is a family of flowering plants consisting of about 130 species in eight genera. In the APG II system it is one of eight families in the euasterids II clade (campanulids) that are unplaced as to order. More recent research has pro ...
. Indigenous trees include species of the genera ''
Ocotea ''Ocotea'' is a genus of flowering plants belonging to the family Lauraceae. Many are evergreen trees with lauroid leaves. There are over 520 species currently accepted within the genus, distributed mostly in tropical and subtropical areas of ...
,
Erythrina ''Erythrina'' is a genus of plants in the pea family, Fabaceae. It contains about 130 species, which are distributed in tropical and subtropical regions worldwide. They are trees, with the larger species growing up to in height. These species ...
,
Sideroxylon ''Sideroxylon'' is a genus of trees in the family (biology), family Sapotaceae described as a genus by Carl Linnaeus, Linnaeus in 1753. They are collectively known as bully trees. The generic name is derived from the Greek language, Greek words ...
'', and ''
Foetidia ''Foetidia'' is a genus of flowering plants first described as a genus in 1788. Most authors place the genus in the family Lecythidaceae, the sole genus in the subfamily Foetidioideae, but some prefer to treat it as a distinct family, the Foeti ...
''. The palm genera ''
Latania ''Latania'', commonly known as latan palm or latania palm, is a genus of flowering plant in the Arecaceae, palm tree family (biology), family, native to the Mascarene Islands in the western Indian Ocean. Species The genus contains three species ...
'', ''
Hyophorbe ''Hyophorbe'' is a genus of five known species of flowering plants in the family Arecaceae, native to the Mascarene Islands in the Indian Ocean. All five species can attain heights of over 6 meters, and two of the species develop swollen trunks t ...
'', ''
Acanthophoenix ''Acanthophoenix'' is a genus of flowering plants in the Arecaceae, palm family from the Mascarene Islands in the Indian Ocean, where they are commonly called palmiste rouge. A genus long in flux, three species are currently recognized, though u ...
'', ''
Dictyosperma ''Dictyosperma'' is a monotypic genus of flowering plant in the palm family found in the Mascarene Islands in the Indian Ocean (Mauritius, Réunion and Rodrigues). The sole species, ''Dictyosperma album'', is widely cultivated in the tropics but ...
'', and ''
Tectiphiala ''Tectiphiala ferox'', or ''palmiste bouglé'', is a species of flowering plant in the family Arecaceae. It is endemic to Mauritius. It is threatened by habitat loss. It is the sole species in the genus ''Tectiphiala''. ''Tectiphiala ferox'' was ...
'' are all endemic to the Mascarene Islands. Other endemic genera include ''
Berenice Berenice (, ''Bereníkē'') is the Ancient Macedonian form of the Attic Greek name ''Pherenikē'', which means "bearer of victory" . Berenika, priestess of Demeter in Lete ca. 350 BC, is the oldest epigraphical evidence. The Latin variant Veron ...
, Heterochaenia, Nesocodon, Ruizia'', and ''
Astiria ''Ruizia'' is a genus of flowering plants belonging to the family Malvaceae. It contains 13 species native mostly to the Mascarene Islands (Mauritius and Réunion), and one species native to Madagascar. The genus was described in 1786 by Antonio ...
''. There are 24 species of trees and shrubs in the Mascarenes from subfamily
Dombeyoideae Dombeyoideae is a widely distributed subfamily of the Malvaceae, as proposed by the APG. Most of the plants placed here were once assembled with more or less related genera in the paraphyletic Sterculiaceae; a lesser number were placed in the Ti ...
, 23 of which are endemic to the islands. The Macarenes' endemic species are polyphyletic and split into nine clades. ''
Trochetia ''Trochetia'' is a subgenus of flowering plants, now placed in the genus '' Ruizia'' and the family Malvaceae (formerly in the Sterculiaceae, but genera now usually subsumed in the Dombeyoideae and other subfamilies). Species are endemic to the M ...
'' appears monophyletic and more closely related to ''
Eriolaena ''Eriolaena'' is a genus of flowering plants. Traditionally included in the family (biology), family Sterculiaceae, it is included now in the recently expanded Malvaceae.Hinsley, S. R. (2007)Synonymy of Malvaceae The genus is distributed in Asia ...
'' and '' Helmiopsis'' than to ''
Dombeya ''Dombeya'' is a flowering plant genus. Traditionally included in the family Sterculiaceae, it is included in the expanded Malvaceae in the APG and most subsequent systematics. These plants are known by a number of vernacular names which sometim ...
''. All ''Dombeya'' taxa are included in a clade together with '' Ruizia'' and ''
Astiria ''Ruizia'' is a genus of flowering plants belonging to the family Malvaceae. It contains 13 species native mostly to the Mascarene Islands (Mauritius and Réunion), and one species native to Madagascar. The genus was described in 1786 by Antonio ...
'', this means that ''Dombeya'' is paraphyletic. In terms of breeding systems the Malagasy Dombeyoideae are
hermaphroditic A hermaphrodite () is a sexually reproducing organism that produces both male and female gametes. Animal species in which individuals are either male or female are gonochoric, which is the opposite of hermaphroditic. The individuals of many ...
, whereas those of the Mascarenes are considered
dioecious Dioecy ( ; ; adj. dioecious, ) is a characteristic of certain species that have distinct unisexual individuals, each producing either male or female gametes, either directly (in animals) or indirectly (in seed plants). Dioecious reproduction is ...
. The
polyphyly A polyphyletic group is an assemblage that includes organisms with mixed evolutionary origin but does not include their most recent common ancestor. The term is often applied to groups that share similar features known as homoplasies, which ar ...
of the Mascarene Dombeyoideae suggests that dioecy has been acquired several times. At least five colonization events from Madagascar to the Mascarene archipelago occurred. The evolutionary history of two lineages of Mascarene Domeyoideae seems to be related to adaption to xeric habitats. The flora of the islands co-evolved with the island's unique fauna over millions of years. The human-caused extinction of several of the Mascarenes' land animals, and the introduction of exotic animals to the islands by humans, has disrupted the reproduction and thriving of various island plants. For example, the Tambalacoque tree (''Sideroxylon grandiflorum''), often called the dodo tree, is not producing young trees and is threatened with extinction. The extinct
dodo The dodo (''Raphus cucullatus'') is an extinction, extinct flightless bird that was endemism, endemic to the island of Mauritius, which is east of Madagascar in the Indian Ocean. The dodo's closest relative was the also-extinct and flightles ...
, ''
Cylindraspis ''Cylindraspis'' is a genus of recently extinct giant tortoises. All of its species lived in the Mascarene Islands (Mauritius, Rodrigues and Réunion) in the Indian Ocean and all are now extinct due to hunting and introduction of non-native pred ...
''
tortoises Tortoises ( ) are reptiles of the family Testudinidae of the order Testudines (Latin for "tortoise"). Like other turtles, tortoises have a turtle shell, shell to protect from predation and other threats. The shell in tortoises is generally hard ...
, and
broad-billed parrot The broad-billed parrot or raven parrot (''Lophopsittacus mauritianus'') is a large extinct parrot in the family (biology), family Psittaculidae. It was endemism in birds, endemic to the Mascarene island of Mauritius. The species was first refe ...
s, along with
fruit bat Megabats constitute the family Pteropodidae of the order Chiroptera. They are also called fruit bats, Old World fruit bats, or—especially the genera ''Acerodon'' and ''Pteropus''—flying foxes. They are the only member of the superfamil ...
s, played an important role in dispersing the seeds. Passing through the animals' digestive tract aided in the seeds' germination. However the trees' decline may be mostly due to introduced
crab-eating macaque The crab-eating macaque (''Macaca fascicularis''), also known as the long-tailed macaque or cynomolgus macaque, is a cercopithecine primate native to Southeast Asia. As a synanthropic species, the crab-eating macaque thrives near human settlem ...
s (''Macaca fascicularis'') destroying unripened seeds.


Fauna

Much of the Mascarenes' native fauna has become endangered or extinct since the human settlement of the islands in the 17th century. Settlers cleared most of the forests for agriculture and grazing, and introduced many exotic species, including
pig The pig (''Sus domesticus''), also called swine (: swine) or hog, is an omnivorous, domesticated, even-toed, hoofed mammal. It is named the domestic pig when distinguishing it from other members of the genus '' Sus''. Some authorities cons ...
s,
rat Rats are various medium-sized, long-tailed rodents. Species of rats are found throughout the order Rodentia, but stereotypical rats are found in the genus ''Rattus''. Other rat genera include '' Neotoma'' (pack rats), '' Bandicota'' (bandicoo ...
s,
cat The cat (''Felis catus''), also referred to as the domestic cat or house cat, is a small domesticated carnivorous mammal. It is the only domesticated species of the family Felidae. Advances in archaeology and genetics have shown that the ...
s,
monkey Monkey is a common name that may refer to most mammals of the infraorder Simiiformes, also known as simians. Traditionally, all animals in the group now known as simians are counted as monkeys except the apes. Thus monkeys, in that sense, co ...
s, and
mongoose A mongoose is a small terrestrial carnivorous mammal belonging to the family Herpestidae. This family has two subfamilies, the Herpestinae and the Mungotinae. The Herpestinae comprises 23 living species that are native to southern Europe, A ...
s.


Birds

Many of the Mascarene birds evolved into flightless forms, the most famous of which was the
extinct Extinction is the termination of an organism by the death of its Endling, last member. A taxon may become Functional extinction, functionally extinct before the death of its last member if it loses the capacity to Reproduction, reproduce and ...
flightless
pigeon Columbidae is a bird family consisting of doves and pigeons. It is the only family in the order Columbiformes. These are stout-bodied birds with small heads, relatively short necks and slender bills that in some species feature fleshy ceres. ...
, the
dodo The dodo (''Raphus cucullatus'') is an extinction, extinct flightless bird that was endemism, endemic to the island of Mauritius, which is east of Madagascar in the Indian Ocean. The dodo's closest relative was the also-extinct and flightles ...
of Mauritius. Thirteen additional species of birds became extinct, including the
Réunion flightless ibis Réunion (; ; ; known as before 1848) is an island in the Indian Ocean that is an overseas departments and regions of France, overseas department and region of France. Part of the Mascarene Islands, it is located approximately east of the isl ...
,
broad-billed parrot The broad-billed parrot or raven parrot (''Lophopsittacus mauritianus'') is a large extinct parrot in the family (biology), family Psittaculidae. It was endemism in birds, endemic to the Mascarene island of Mauritius. The species was first refe ...
,
red rail The red rail (''Aphanapteryx bonasia'') is an extinct species of rail that was endemic to the Mascarene island of Mauritius, east of Madagascar in the Indian Ocean. It had a close relative on Rodrigues island, the likewise extinct Rodrigues r ...
, Rodrigues rail, and
Rodrigues solitaire The Rodrigues solitaire (''Pezophaps solitaria'') is an extinct flightless bird that was endemism, endemic to the island of Rodrigues, east of Madagascar in the Indian Ocean. Genetically within the family of Columbidae, pigeons and doves, it wa ...
. Contemporary sources state that the dodo used
gizzard stones A gastrolith, also called a stomach stone or gizzard stone, is a rock held inside a gastrointestinal tract. Gastroliths in some species are retained in the muscular gizzard and used to grind food in animals lacking suitable grinding teeth. In oth ...
and could swallow ''Ocotea'' fruits. Seed distribution of at least some ''Ocotea'' species is performed by
frugivorous A frugivore ( ) is an animal that thrives mostly on raw fruits or succulent fruit-like produce of plants such as roots, shoots, nuts and seeds. Approximately 20% of mammalian herbivores eat fruit. Frugivores are highly dependent on the abundance ...
birds and there exist a few reports of "solitaires" from the Mascarenes without mention of which island these came from, and the term was also used for other
species A species () is often defined as the largest group of organisms in which any two individuals of the appropriate sexes or mating types can produce fertile offspring, typically by sexual reproduction. It is the basic unit of Taxonomy (biology), ...
with "solitary" habits, such as the Réunion blue swamphen and the
Réunion sacred ibis Réunion (; ; ; known as before 1848) is an island in the Indian Ocean that is an overseas department and region of France. Part of the Mascarene Islands, it is located approximately east of the island of Madagascar and southwest of the isla ...
. At one point it was even believed that Réunion was the home of not only a white dodo, but also a white solitaire. In 1786, sub-fossil bones were discovered in a cave which confirmed Leguat's descriptions, but at this time no living residents of Rodrigues remembered having seen living birds. The star
constellation A constellation is an area on the celestial sphere in which a group of visible stars forms Asterism (astronomy), a perceived pattern or outline, typically representing an animal, mythological subject, or inanimate object. The first constellati ...
Turdus Solitarius Turdus Solitarius (Latin for ''solitary thrush'') was a constellation created by French astronomer Pierre Charles Le Monnier in 1776 from stars of Hydra's tail. It was named after the Rodrigues solitaire, an extinct flightless bird that was ende ...
was named after this bird. Today 17 endemic bird species survive on the islands. Two species – the
Mascarene paradise flycatcher The Mascarene paradise flycatcher (''Terpsiphone bourbonnensis'') is a species of bird in the monarch-flycatcher family Monarchidae. It is endemic to the Mascarene islands of Mauritius and Réunion. There are two subspecies recognized: the nomin ...
(''Terpsiphone bourbonnensis'') and
Mascarene swiftlet The Mascarene swiftlet or Mauritius swiftlet (''Aerodramus francicus'') is a species of swift in the family Apodidae. It is found in Mauritius and Réunion, and the populations on the two islands have recently been confirmed to differ subspecifica ...
(''Aerodramus francicus'') – inhabit both Mauritius and Réunion. Eight species are endemic to Mauritius – the
Mauritius grey white-eye The Mauritius grey white-eye (''Zosterops mauritianus'') is a species of bird in the family Zosteropidae. It is one of two white-eye species endemic to the island of Mauritius, the other being the rare and localized Mauritius olive white-eye. It ...
, (''Zosterops mauritianus''), Mauritius cuckooshrike (''Lalage typica''),
Mauritius kestrel The Mauritius kestrel (''Falco punctatus'') is a bird of prey from the family Falconidae endemic to the forests of Mauritius, where it is restricted to the southwestern plateau's forests, cliffs, and ravines. It is the most distinct of the India ...
(''Falco punctatus''),
Mauritius fody The Mauritius fody (''Foudia rubra'') is a rare species of bird in the weaver family. It is endemic to the island of Mauritius. It is classified by BirdLife International as being endangered. It is also on the United States' Endangered Species L ...
(''Foudia rubra''),
Mauritius bulbul The Mauritius bulbul (''Hypsipetes olivaceus''), is a songbird belonging to the bulbul family (biology), family, Pycnonotidae. It is endemic to Mauritius. Taxonomy and systematics The Mauritius bulbul was the type species of the obsolete genus ' ...
(''Hypsipetes olivaceus''), Mauritius parakeet (''Psittacula eques''),
Mauritius olive white-eye The Mauritius olive white-eye (''Zosterops chloronothos'', often mistakenly spelled ''Zosterops chloronothus''.) is a very rare and localized passerine from the family of white-eyes (''Zosteropidae''). It is one of two white-eye species endemic ...
(''Zosterops chloronothos''), and
pink pigeon The pink pigeon (''Nesoenas mayeri'') is a species of pigeon in the family Columbidae Endemic (ecology), endemic to Mauritius. The pink pigeon nearly became extinct in the 1970s and the 1990s and is still very rare. It is the only Mascarene pigeo ...
(''Nesoenas mayeri''). The
Réunion grey white-eye The Réunion grey white-eye (''Zosterops borbonicus'') is a small passerine from the family Zosteropidae, which is native to the islands of Réunion.Gill F. and Donsker D. (eds), Family Zosteropidae, in IOC World Bird Names (ver 6.2), Internatio ...
(''Zosterops borbonicus''),
Réunion cuckooshrike The Réunion cuckooshrike (''Lalage newtoni'') is a passerine bird in the cuckooshrike family. It is endemic to the island of Réunion, where it is restricted to two areas of mountain forest in the north of the island. Males are dark grey above a ...
(''Lalage newtoni''),
Réunion stonechat The Réunion stonechat (''Saxicola tectes'') is a species of stonechat, endemic to the French Indian Ocean island of Réunion. It is found across the island in forests, shrublands and artificial environments such as gardens and plantations. This ...
(''Saxicola tectes''),
Réunion olive white-eye The Réunion olive white-eye (''Zosterops olivaceus'') is a species of bird in the family Zosteropidae. It is found on Réunion. Its natural habitats are boreal forests and subtropical or tropical high-altitude grassland. Description like the ...
(''Zosterops olivaceus''), and
Réunion bulbul The Réunion bulbul (''Hypsipetes borbonicus'') is a species of the bulbul family of passerine birds. It is endemic to Réunion. Taxonomy and systematics The Réunion bulbul was originally described in the genus ''Turdus''. Formerly, some author ...
(''Hypsipetes borbonicus'') are endemic to Réunion. The
Rodrigues warbler The Rodrigues warbler (''Acrocephalus rodericanus'') is a species of Old World warbler in the family Acrocephalidae. It is found only on the island of Rodrigues (which belongs to Mauritius) and used to be plentiful there. Their population soon g ...
(''Acrocephalus rodericanus'') and
Rodrigues fody The Rodrigues fody (''Foudia flavicans'') is a rare species of bird in the weaver family. It is endemic to Rodrigues, an island of Mauritius. It is classified by BirdLife International as being vulnerable. It is also on the United States' Endang ...
(''Foudia flavicans'') are found only on Rodrigues.


Reptiles

Mauritius, Réunion, and Rodrigues were also once each home to one or more species of
giant tortoise Giant tortoises are any of several species of various large land tortoises, which include a number of extinct species, as well as two extant species with multiple subspecies formerly common on the islands of the western Indian Ocean and on the ...
s, now extinct, which comprised the genus ''
Cylindraspis ''Cylindraspis'' is a genus of recently extinct giant tortoises. All of its species lived in the Mascarene Islands (Mauritius, Rodrigues and Réunion) in the Indian Ocean and all are now extinct due to hunting and introduction of non-native pred ...
''. There are 13 living endemic reptile species, including a number of species of day
gecko Geckos are small, mostly carnivorous lizards that have a wide distribution, found on every continent except Antarctica. Belonging to the infraorder Gekkota, geckos are found in warm climates. They range from . Geckos are unique among lizards ...
es (
genus Genus (; : genera ) is a taxonomic rank above species and below family (taxonomy), family as used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In bino ...
''
Phelsuma ''Phelsuma'' is a large genus of geckos in the Family (biology), family Gekkonidae. Species in the genus ''Phelsuma'' are commonly referred to as day geckos. Some day geckos are seriously endangered and some are common, but all ''Phelsuma'' spec ...
'').


Mammals

The islands' only native mammals are
bat Bats are flying mammals of the order Chiroptera (). With their forelimbs adapted as wings, they are the only mammals capable of true and sustained flight. Bats are more agile in flight than most birds, flying with their very long spread-out ...
s, including the
Mauritian flying fox The Mauritian flying fox (''Pteropus niger''), also known as Greater Mascarene flying fox or Mauritius fruit bat is a large megabat species endemic to Mauritius and La Réunion LA most frequently refers to Los Angeles, the second most populou ...
(''Pteropus niger''), which is endemic to Mauritius and Réunion, and the
Rodrigues flying fox The Rodrigues flying fox or Rodrigues fruit bat (''Pteropus rodricensis'') is a species of bat in the family Pteropodidae, the flying foxes or fruit bats. It is endemic to Rodrigues, an island in the Indian Ocean belonging to Mauritius. Its nat ...
(''Pteropus rodricensis'') found only on Rodrigues. The lesser Mascarene flying fox (''Pteropus subniger'') is extinct.


Freshwater

The three largest islands have many freshwater streams which descend from the mountains to the sea. The upper reaches, particularly on Réunion, are fast-flowing and steep with numerous waterfalls. Tamarin Falls or Sept Cascades (293 m) on Mauritius is the highest waterfall on Mauritius, while Cascade Trou de Fer (725 m, ranks 23rd worldwide) on Réunion is the highest of the Mascarenes."Mascarenes". Freshwater Ecoregions of the World. Accessed 21 August 2021

The Grand River South East (34 km) is the longest river on Mauritius, followed by Rivière du Poste (23 km), Grand River North West (22 km), Rivière La Chaux (22 km), and Rivière des Créoles (20 km). Mauritius has two natural crater lakes, Grand Bassin and Bassin Blanc. There are about twenty species of fish which spend at least part of their lives in freshwater. The islands have five endemic freshwater fish species, all goby, gobies (Gobiiformes) – ''
Hypseleotris cyprinoides ''Hypseleotris cyprinoides'', the tropical carp-gudgeon or tropical bitterling-gudgeon, is a species of fish in the family Eleotridae found in fresh, brackish, and marine coastal waters from Africa through southern Asia to the Pacific Islands. T ...
,
Cotylopus acutipinnis ''Cotylopus'' is a genus of gobies native to streams and rivers in Mauritius, Réunion, Mayotte, and Anjouan in the Comoros, which are all islands in the Western Indian Ocean off Africa. Species The recognized species in this genus are: * ''Cot ...
, Glossogobius kokius, Gobius commersonii'', and '' Oxyurichthys guibei''. ''
Cotylopus acutipinnis ''Cotylopus'' is a genus of gobies native to streams and rivers in Mauritius, Réunion, Mayotte, and Anjouan in the Comoros, which are all islands in the Western Indian Ocean off Africa. Species The recognized species in this genus are: * ''Cot ...
'', which is endemic to Réunion, hatches in the sea before migrating to the rainforest streams where it lives its adult life.


Marine

The Mascarene Islands are surrounded by approximately 750 square km of coral reef. Rodrigues has nearly continuous
fringing reef A fringing reef is one of the three main types of coral reef. It is distinguished from the other main types, barrier reefs and atolls, in that it has either an entirely shallow backreef zone (lagoon) or none at all. If a fringing reef grows direc ...
s bounding an extensive lagoon, 7 to 20 km wide, with deep channels. Mauritius is also surrounded by a fringing reef. In contrast, Réunion has very short stretches of narrow fringing reef along the western and southwestern coasts only. The islets of the Cargados Carajos Shoals, which have a very depauperate terrestrial biota owing to being so low-lying and swamped during cyclones, are bound to the east by an extensive arc of fringing reef, which accounts for ~30% of the reefs of the Mascarene Islands. Lagoon reefs and reef flats are dominated by scleractinian corals such as branching and tabular ''
Acropora ''Acropora'' is a genus of small polyp stony coral in the phylum Cnidaria. Some of its species are known as table coral, elkhorn coral, and staghorn coral. Over 149 species are described. ''Acropora'' species are some of the major reef corals ...
'', ''
Porites ''Porites'' is a genus of stony coral; they are small polyp stony (SPS) corals. (Also referred to as finger coral or hump coral) They are characterised by a finger-like morphology. Members of this genus have widely spaced calices, a well-de ...
'' massives, foliaceous ''
Montipora ''Montipora'' is a genus of Scleractinian corals in the phylum Cnidaria. Members of the genus ''Montipora'' may exhibit many different growth morphologies. With eighty five known species, ''Montipora'' is the second most species rich coral genus ...
'' and '' Pavona'', and sand consolidated with beds of seagrass such ''
Halophila ''Halophila'' is a genus of seagrasses in the family '' Hydrocharitaceae'', the tape-grasses. It was described as a genus in 1806. The number of its contained species, and its own placement in the order Alismatales, has evolved. Description T ...
'' (
Hydrocharitaceae Hydrocharitaceae is a flowering plant family which includes 14 accepted genera and a total of ca 135 known species (Christenhusz & Byng 2016). The family holds a number of species of aquatic plants, including tape-grass, the well-known Canadian ...
). Among coral reef fishes, wrasses ( Labridae), damselfish (
Pomacentridae Pomacentridae is a family of ray-finned fish, comprising the damselfishes and clownfishes. This family were formerly placed in the order Perciformes or as indeterminate percomorphs, but are now considered basal blenniiforms. They are primaril ...
), carnivorous groupers (
Serranidae Serranidae is a large family (biology), family of fishes belonging to the order Perciformes. The family contains about 450 species in 65 genera, including the sea basses and the groupers (subfamily Epinephelinae). Although many species are small, ...
), and surgeonfishes (Acanthuridae) have many species.


Wildlife, threats, and preservation

Mauritius has one of the highest population densities in the world. All of the islands have experienced significant losses of habitats. Many of the surviving endemic species are still threatened with extinction and have little protection. Less than 40 percent of Réunion is covered with natural vegetation, only about 5 percent of Mauritius, and almost none of Rodrigues. On Réunion, forest has been cleared for agriculture and then overtaken by introduced plants. Mauritius was largely converted to sugar cane, tea, and conifer plantations. On Rodrigues, shifting cultivation has caused the damage.


Launch of Saint Brandon Conservation Trust

On 8 May 2024,
Saint Brandon Conservation Trust In Christian belief, a saint is a person who is recognized as having an exceptional degree of holiness, likeness, or closeness to God. However, the use of the term ''saint'' depends on the context and denomination. In Anglican, Oriental Ortho ...
, an independent Mauritian NGO, was internationally launched at the
Corporate Council on Africa The Corporate Council on Africa (CCA) is a trade association focusing on strengthening commercial relationships between the United States and the African continent. History The Corporate Council on Africa (CCA), based in Washington D.C., is a nonp ...
in Dallas, Texas, to protect St. Brandon from pollution and to protect surviving flora and fauna in the
Cargados Carajos Saint Brandon (), also known as the Cargados Carajos Shoals, is a southwest Indian Ocean archipelago of sand banks, shoals and islets belonging to the Republic of Mauritius. It lies about northeast of the island of Mauritius. It consists of fiv ...
.


Protected areas

Protected areas constitute 40.6% of the Mascarene Islands' land area. Terrestrial protected areas on Mauritius include
Black River Gorges National Park Black River Gorges National Park is a national park in the hilly south-western part of Mauritius. It was proclaimed on June 15, 1994 and is managed by the National Parks and Conservation Service. It covers an area of 67.54 km2 including humi ...
, Bras d'Eau National Park, Perrier Nature Reserve, Corps de Garde Nature Reserve, Le Pouce Nature Reserve, Cabinet Nature Reserve, Gouly Pere Nature Reserve, and Bois Sec Nature Reserve. There are 32 designated protected areas on Réunion, including
Réunion National Park Réunion National Park () is a National Park of France located on the island of Réunion, an overseas department in the western Indian Ocean. Established on 5 March 2007, the park protects the endemic ecosystems of ''Les Hauts'', Réunion's moun ...
(1055.15 km2), which cover 63.13% of the island.UNEP-WCMC (2021). Protected Area Profile for Réunion from the World Database of Protected Areas. Accessed 18 August 2021


See also

*
St. Brandon Saint Brandon (), also known as the Cargados Carajos Shoals, is a southwest Indian Ocean archipelago of sand banks, shoals and islets belonging to the Republic of Mauritius. It lies about northeast of the island of Mauritius. It consists of fiv ...
*
Île Raphael Île Raphael is an island in the Saint Brandon archipelago, a group of 30 outer islands of Mauritius. The island is named after Veuve Raphaël. Veuve Raphaël's husband was a sea captain and had installations on the corner of rue (route) des ...
*
L'île du Sud L'île du Sud (South Island, l'île Boisées) is an island located in the St. Brandon archipelago. It is one of the three islands (the others being Île Raphael and L'Île Coco) used as a base of operations for fishing activities by Raphael Fishin ...
*
Avocaré Island Avocaré Island (Avoquer, Avocaire, L'Avocaire) is an island located in the St Brandon archipelago. In the World Bank Report recommending the creation of the marine protected area of St. Brandon, Avocaré Island was classified as a Group 3 Isla ...
*
France Staub France Staub (September 29, 1920 – July 2, 2005) was a Mauritian ornithologist, herpetologist, botanist, and conservationist. Biography Staub was a descendant of French botanist Jacques Delisse (1773−1856). He obtained the diploma at the Ma ...
*
L'île du Gouvernement l'île du Gouvernement is an island in the St. Brandon archipelago. The island is uninhabited, and mostly functions as a bird and turtle sanctuary. This island was one of those recommended as part of the ''St. Brandon Marine Protected Area'' (M ...
*
ÃŽle Verronge ÃŽle Verronge is one of circa thirty isles, island and sandbars that is part of the St. Brandon archipelago in the Indian Ocean. It is constitutionally designated, under Mauritian law, as one of the outer islands of Mauritius. In the 1840s, the ...
*
Constitution of Mauritius The Constitution of the Republic of Mauritius () is the supreme law of Mauritius, according to Chapter I, Section 2 of the constitution, if any other law is inconsistent with this Constitution, that other law shall, to the extent of the inconsist ...
*
Mauritian Wildlife Foundation The Mauritian Wildlife Foundation (MWF) is an independent, non-governmental, non-profit conservation agency working in Mauritius and the Outer Islands to save threatened endemic local flora and fauna. The MWF is currently working to conserve th ...


References


Sources

* Quammen, David, (1996) ''The Song of the Dodo''. Touchstone,
New York New York most commonly refers to: * New York (state), a state in the northeastern United States * New York City, the most populous city in the United States, located in the state of New York New York may also refer to: Places United Kingdom * ...
. * Diamond, Jared, (1984) "Historic extinctions: A rosetta stone for understanding prehistoric extinctions". In: P. Martin and R. Klein (eds.) (1984) ''Quaternary Extinctions: A prehistoric revolution.'' University of Arizona Press,
Tucson Tucson (; ; ) is a city in Pima County, Arizona, United States, and its county seat. It is the second-most populous city in Arizona, behind Phoenix, Arizona, Phoenix, with a population of 542,630 in the 2020 United States census. The Tucson ...
.


External links


Scientific research application on the nature reserve of Mare-Longue (Réunion - Mascarene Islands)
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PDF from the site of
Biblioteca Estense The Biblioteca Estense (), was the family library of the marquises and dukes of the House of Este. The exact date of the library's birth is still under speculation, however it is known for certain that the library was in use during the fourteenth ...
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Modena Modena (, ; ; ; ; ) is a city and ''comune'' (municipality) on the south side of the Po Valley, in the Province of Modena, in the Emilia-Romagna region of northern Italy. It has 184,739 inhabitants as of 2025. A town, and seat of an archbis ...

del Cantino
Cantino planisphere image, zoomable to very high resolution, at ''Bibliothèque et Archives nationales du Québec''. (Click on ''1:1'' button to get maximum resolution. Click on ''Image plein écran'' to get full-screen view.) {{Authority control
Mascarene Islands The Mascarene Islands (, ) or Mascarenes or Mascarenhas Archipelago is a group of islands in the Indian Ocean east of Madagascar consisting of islands belonging to the Republic of Mauritius as well as the French department of Réunion. Their na ...
Afrotropical ecoregions Archipelagoes of Africa Archipelagoes of the Indian Ocean Former French colonies Former Portuguese colonies Former colonies in Africa Freshwater ecoregions of Africa Geography of East Africa Islands of France Islands of Mauritius Marine ecoregions Tropical and subtropical moist broadleaf forests Volcanic arc islands Western Indo-Pacific Ile Raphael Outer Islands of Mauritius Reefs of the Indian Ocean Fishing areas of the Indian Ocean Insular ecology Important Bird Areas of Mauritius Atolls of the Indian Ocean Biodiversity Quarantine facilities Fly fishing Fishing tournaments Environment of Mauritius Natural history of Mauritius