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Giant Tortoise
Giant tortoises are any of several species of various large land tortoises, which include a number of extinct species, as well as two extant species with multiple subspecies formerly common on the islands of the western Indian Ocean and on the Galápagos Islands. History As of February 2024, two different species of giant tortoise are found on two remote groups of tropical islands: Aldabra Atoll and Fregate Island in the Seychelles and the Galápagos Islands in Ecuador. These tortoises can weigh as much as and can grow to be long. Giant tortoises originally made their way to islands from the mainland via oceanic dispersal. Tortoises are aided in such dispersal by their ability to float with their heads up and to survive for up to six months without food or fresh water. Giant tortoises were once all placed in a single genus (often referred to as ''Testudo'' or ''Geochelone''), but more recent studies have shown that giant tortoises represent several distinct lineages that ...
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Tortoise
Tortoises ( ) are reptiles of the family Testudinidae of the order Testudines (Latin for "tortoise"). Like other turtles, tortoises have a shell to protect from predation and other threats. The shell in tortoises is generally hard, and like other members of the suborder Cryptodira, they retract their necks and heads directly backward into the shell to protect them. Tortoises can vary in size with some species, such as the Galápagos giant tortoise, growing to more than in length, whereas others like the Speckled cape tortoise have shells that measure only long. Several lineages of tortoises have independently evolved very large body sizes in excess of , including the Galapagos giant tortoise and the Aldabra giant tortoise. They are usually diurnal animals with tendencies to be crepuscular depending on the ambient temperatures. They are generally reclusive animals. Tortoises are the longest-living land animals in the world, although the longest-living species of tortois ...
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Manouria
''Manouria'' is a genus of tortoises in the family Testudinidae. The genus was erected by John Edward Gray in 1854. Transitional adaptations ''Manouria'' is either the most basal or second most basal member of the family Testudinidae (its phylogenetic position is adjacent to that of '' Gopherus''). As a result, while tortoises are primarily terrestrial, the genus ''Manouria'' is a model for the evolutionary transition to terrestriality, as it still has an affinity for aquatic environments, and has retained some ancestral adaptations for an aquatic lifestyle while losing others. The Asian forest tortoise (''Manouria emys'') has been observed attempting to feed on submerged food items, albeit unsuccessfully. Additionally, the species grasps food item with its jaws as in aquatic or semiaquatic taxa, as opposed to first making contact with the tongue as with all other tortoise species (with the possible exception of ''Gopherus''). Species The following five species are recognized ...
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Cenozoic
The Cenozoic Era ( ; ) is Earth's current geological era, representing the last 66million years of Earth's history. It is characterized by the dominance of mammals, insects, birds and angiosperms (flowering plants). It is the latest of three geological eras of the Phanerozoic Eon, preceded by the Mesozoic and Paleozoic. The Cenozoic started with the Cretaceous–Paleogene extinction event, when many species, including the non-avian dinosaurs, became extinct in an event attributed by most experts to the impact of a large asteroid or other celestial body, the Chicxulub impactor. The Cenozoic is also known as the Age of Mammals because the terrestrial animals that dominated both hemispheres were mammalsthe eutherians ( placentals) in the Northern Hemisphere and the metatherians (marsupials, now mainly restricted to Australia and to some extent South America) in the Southern Hemisphere. The extinction of many groups allowed mammals and birds to greatly diversify so that large m ...
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Quaternary Extinction Event
The Late Pleistocene to the beginning of the Holocene saw the extinction of the majority of the world's megafauna, typically defined as animal species having body masses over , which resulted in a collapse in faunal density and diversity across the globe. The extinctions during the Late Pleistocene are differentiated from previous extinctions by their extreme size bias towards large animals (with small animals being largely unaffected), and widespread absence of ecological succession to replace these extinct megafaunal species, and the regime shift of previously established faunal relationships and habitats as a consequence. The timing and severity of the extinctions varied by region and are generally thought to have been driven by humans, climatic change, or a combination of both. Human impact on megafauna populations is thought to have been driven by hunting ("overkill"), as well as possibly environmental alteration. The relative importance of human vs climatic factors in the ...
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Island Gigantism
Island gigantism, or insular gigantism, is a biological phenomenon in which the size of an animal species isolated on an island increases dramatically in comparison to its mainland relatives. Island gigantism is one aspect of the more general "island effect" or "Foster's rule", which posits that when mainland animals colonize islands, small species tend to evolve larger bodies, and large species tend to evolve smaller bodies (insular dwarfism). This is itself one aspect of the more general phenomenon of island syndrome which describes the differences in morphology, ecology, physiology and behaviour of insular (island) species compared to their continental counterparts. Following the arrival of humans and associated introduced predators (dogs, cats, rats, pigs), many giant as well as other island endemics have become extinct (e.g. the dodo and Rodrigues solitaire, giant flightless pigeons related to the Nicobar pigeon). A similar size increase, as well as increased woodiness, has b ...
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Titanochelon
''Titanochelon'' is an extinct genus of giant tortoises known from the Early Miocene to the beginning of the Pleistocene in Europe, extending from the Iberian Peninsula to Anatolia, as well as possibly North Africa. Some members of the genus were larger than extant giant tortoises, with a shell length of up to . Taxonomy There are approximately 10 known species in the genus, most of which were originally assigned to '' Testudo'' (a genus which formally encompassed almost all fossil tortoises) or '' Cheirogaster,'' the type species of which, ''Cheirogaster maurini'' is known from the Eocene of France and is quite different to the species assigned to ''Titanochelon''. After a major systematic revision in 2014, the genus ''Titanochelon'' was created to house these related species.Hans Volker Karl, Ulrich Staesche, Amtyaz Safi New findings of neogene tortoises ''Titanochelon kayadibiensis'' sp. nov. and ''Protestudo bessarabica'' (Riabinin, 1918) (Testudinidae) from the Miocene of w ...
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Solitudo
''Solitudo'' is an extinct genus of tortoise that was found during the Pliocene and Pleistocene on the Mediterranean islands of Menorca, Malta and Sicily. The genus includes three described species, ''Solitudo robusta'', ''Solitudo gymnesica'' and ''Solitudo sicula'' as well as a likely fourth, undescribed species from Monte Pellegrino in Sicily. ''Solitudo sicula'', the youngest of the species, died out approximately 12.5 thousand years Before Present, BP. The largest species, ''Solitudo gymnesica'', has been estimated to have reached a carapace length of . History The oldest discoveries of fossil turtles now included under ''Solitudo'' were made in the 19th century, with Leith-Adams describing remains from Zebbug Cave (Malta) as ''Testudo robusta''. In 1914 ''Testudo gymnesica'' was described based on Pliocene material found on Menorca. Younger material was later discovered in the Zubbio di Cozzo San Pietro cave in northern Sicily. These remains, which include a femur, phalanx ...
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Megalochelys
''Megalochelys'' ("great turtle") is an extinct genus of tortoises that lived from the Miocene to Pleistocene. They are noted for their giant size, the largest known for any tortoise, with a maximum carapace length of over 2 m (6.5 ft) in ''M. atlas''. The genus ranged from western India and Pakistan to as far east as Sulawesi and Timor in Indonesia, though the island specimens likely represent distinct species. Extinct animals of Indonesia Description One species of ''Megalochelys,'' ''M. atlas'', is the largest known tortoise, with shells of in length having been reported. Popular weight estimates for this taxon have varied greatly with the highest estimates reaching up to in some instances.Orenstein, R. 2001. Survivors in Armor: Turtles, Tortoises, and Terrapins. Key Porter Books Ltd. However, weights based on volumetric displacement of the skeleton suggest a mass of around .Brown, B. 1931. The Largest Known Land Tortoise. Nat. Hist. Vol. 31:184–187. ''M. atl ...
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Before Present
Before Present (BP) or "years before present (YBP)" is a time scale used mainly in archaeology, geology, and other scientific disciplines to specify when events occurred relative to the origin of practical radiocarbon dating in the 1950s. Because the "present" time changes, standard practice is to use 1January 1950 as the commencement date (epoch) of the age scale, with 1950 being labelled as the "standard year". The abbreviation "BP" has been interpreted retrospectively as "Before Physics", which refers to the time before nuclear weapons testing artificially altered the proportion of the carbon isotopes in the atmosphere, which scientists must account for when using radiocarbon dating for dates of origin that may fall after this year. In a convention that is not always observed, many sources restrict the use of BP dates to those produced with radiocarbon dating; the alternative notation "RCYBP" stands for the explicit "radio carbon years before present". Usage The BP scale is s ...
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Hesperotestudo
''Hesperotestudo'' ("Western turtle") is an extinct genus of tortoise native to North and Central America (ranging as far south as Costa Rica) from the Early Miocene to the Late Pleistocene. Species of ''Hesperotestudo'' varied widely in size, with a large undescribed specimen from the Late Pleistocene of El Salvador reaching in carapace length, larger than that of extant giant tortoises. Historically considered a subgenus of ''Geochelone'', it is now considered to be distantly related to that genus. Its relationships with other tortoises are uncertain. The exposed areas of the bodies of ''Hesperotestudo'' species were extensively covered with large dermal ossicles, which in life were covered in keratin. It has been suggested that species of ''Hesperotestudo'' were relatively tolerant of cold weather. ''Hesperotestudo'' became extinct at the end of the Pleistocene roughly co-incident with the arrival of the first humans in North America. There is apparently a site in Florida wh ...
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Chelonoidis
''Chelonoidis'' is a genus of turtles in the tortoise family erected by Leopold Fitzinger in 1835. They are found in South America and the Galápagos Islands, and formerly had a wide distribution in the West Indies. The multiple subspecies of the Galápagos tortoise (''C. niger'') are among the largest extant terrestrial chelonians. Giant members of the genus, such as Lutz's giant tortoise (''C. lutzae'') were also present in mainland South America and the West Indies during the Pleistocene, and the latter into the Holocene. Taxonomy They were formerly assigned to '' Geochelone'', but a 2006 genetic analysis indicated that they were actually most closely related to hingeback tortoises. However, a more recent genetic analysis of mtDNA has found that they are actually most closely related to the lineage containing '' Centrochelys'' and '' Geochelone''. Their ancestors apparently floated across the Atlantic from Africa to South America in the Oligocene. This crossing was made ...
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African Spurred Tortoise
The African spurred tortoise (''Centrochelys sulcata''), also called the sulcata tortoise, is an endangered species of tortoise inhabiting the southern edge of the Sahara, Sahara Desert, the Sahel, in Africa. It is the largest mainland species of tortoise in Africa, and the third-largest in the world, after the Galapagos tortoise and Aldabra giant tortoise. It is the only living species in its genus, ''Centrochelys.'' Taxonomy and etymology In 1779 the English illustrator John Frederick Miller included a hand-coloured plate of the African spurred tortoise in his ''Icones animalium et plantarum'' and coined the binomial name ''Testudo sulcata''. Its Species, specific name ''sulcata'' is from the Latin word ''sulcus'' meaning "furrow" and refers to the furrows on the tortoise's scales. The species was subsequently given other binomial names. There are no recognized subspecies despite there being two separate populations, one in Western Africa and the other in Eastern Africa. Ther ...
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