''Maghriboselache'' is an extinct

genus Genus (; : genera ) is a taxonomic rank above species and below family (taxonomy), family as used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In bino ...

cladoselachid Cladoselachidae is an extinct family of cartilaginous fishes closely related to, and possibly nested within, Symmoriiformes. They are characterized by having an elongated body with a spine in each of the two dorsal fins. In 2023, a paper describi ...

symmoriiform Symmoriiformes is an extinct order of cartilaginous fish. Originally named Symmoriida by Zangerl (1981), the name has since been corrected to Symmoriiformes to avoid confusion with a family. The symmoriiform fossils record begins during the late ...

fish that lived 369 million years ago during the Late

Devonian The Devonian ( ) is a period (geology), geologic period and system (stratigraphy), system of the Paleozoic era (geology), era during the Phanerozoic eon (geology), eon, spanning 60.3 million years from the end of the preceding Silurian per ...

(middle

Famennian The Famennian is the later of two faunal stages in the Late Devonian epoch. The most recent estimate for its duration is that it lasted from around 371.1 to 359.3 million years ago. An earlier 2012 estimate, still used by the International Commis ...

stage) of

Morocco Morocco, officially the Kingdom of Morocco, is a country in the Maghreb region of North Africa. It has coastlines on the Mediterranean Sea to the north and the Atlantic Ocean to the west, and has land borders with Algeria to Algeria–Morocc ...

. The genus contains a single species, ''M. mohmezanei''. ''Maghriboselache'' represents the first cladoselachid to be discovered with significant details of the jaws,

braincase In human anatomy, the neurocranium, also known as the braincase, brainpan, brain-pan, or brainbox, is the upper and back part of the skull, which forms a protective case around the brain. In the human skull, the neurocranium includes the calv ...

and even

endocranium The endocranium in comparative anatomy is a part of the skull base in vertebrates and it represents the basal, inner part of the cranium. The term is also applied to the outer layer of the dura mater in human anatomy. Structure Structurally, t ...

. Along with ''

Cladoselache ''Cladoselache'' ("branch shark") is an Extinction, extinct genus of shark-like chondrichthyan (cartilaginous fish) from the Late Devonian (Famennian) of North America. It was similar in body shape to modern Lamnidae, lamnid sharks (such as Isuru ...

'', it shares a unique and distinctive tooth and upper jaw

morphology Morphology, from the Greek and meaning "study of shape", may refer to: Disciplines *Morphology (archaeology), study of the shapes or forms of artifacts *Morphology (astronomy), study of the shape of astronomical objects such as nebulae, galaxies, ...

Discovery and naming

''Maghriboselache'' was discovered in the eastern parts of the Anti-Atlas mountain range. Most specimens were found in the Thylacocephalan layer. The genus was described by Klug, Coates, Frey, Greif, Jobbins, Pohle, Lagnaoui, Haouz and Ginter in 2023. The generic name, "''Maghriboselache''", is derived from "''al Maghrib''", the

Arabic Arabic (, , or , ) is a Central Semitic languages, Central Semitic language of the Afroasiatic languages, Afroasiatic language family spoken primarily in the Arab world. The International Organization for Standardization (ISO) assigns lang ...

word for Morocco, combined with "σέλαχος" ("''selachos''"), the

Greek Greek may refer to: Anything of, from, or related to Greece, a country in Southern Europe: *Greeks, an ethnic group *Greek language, a branch of the Indo-European language family **Proto-Greek language, the assumed last common ancestor of all kno ...

word for cartilaginous fish. The

specific name Specific name may refer to: * in Database management systems, a system-assigned name that is unique within a particular database In taxonomy, either of these two meanings, each with its own set of rules: * Specific name (botany), the two-part (bino ...

, "''mohamezanei''", honors Moha Mezane, a French amateur geologist and fossil hunter, who discovered some ''Maghriboselache'' fossils.

Description

''Maghriboselache'' is known from multiple preserved fossil specimens preserving most of its skeletal features. Some of these fossils are preserved three-dimensionally. It is unique from other cartilaginous fish (

Chondrichthyes Chondrichthyes (; ) is a class of jawed fish that contains the cartilaginous fish or chondrichthyans, which all have skeletons primarily composed of cartilage. They can be contrasted with the Osteichthyes or ''bony fish'', which have skeleto ...

) due to its broad snout and nostril placement. The genus has distally broad, flat, and strap-like radial

pectoral fins Fins are moving appendages protruding from the body of fish that interact with water to generate thrust and help the fish swim. Apart from the tail or caudal fin, fish fins have no direct connection with the back bone and are supported only b ...

and a cleaver-shaped

palatoquadrate In some fishes, the palatoquadrate is the dorsal component of the mandibular arch, the ventral one being Meckel's cartilage. The palatoquadrate forms from splanchnocranium in various chordates including placoderms and acanthodians. See also * H ...

with an optic process shorter than

palatine process In human anatomy of the mouth, the palatine process of maxilla (palatal process), is a thick, horizontal process of the maxilla. It forms the anterior three quarters of the hard palate, the horizontal plate of the palatine bone making up the rest ...

. The size of ''Maghriboselache'' specimens ranges between in length, but most are slightly more than long.

Teeth

The tooth fossils of ''Maghriboselache'' are only partially visible. The teeth on the

Meckel's cartilage In humans, the cartilaginous bar of the mandibular arch is formed by what are known as Meckel's cartilages (right and left; also known as Meckelian cartilages), above which the incus and malleus are located. Meckel's cartilage arises from the fir ...

of this genus are nearly equally distanced with a 10% gap between each tooth.

Classification

Klug ''et al''. (2023) recovered ''Maghriboselache'' as a

member of the chondrichthyian clade

Symmoriiformes Symmoriiformes is an extinct order of cartilaginous fish. Originally named Symmoriida by Zangerl (1981), the name has since been corrected to Symmoriiformes to avoid confusion with a family. The symmoriiform fossils record begins during the late ...

, as the

sister taxon In phylogenetics, a sister group or sister taxon, also called an adelphotaxon, comprises the closest relative(s) of another given unit in an evolutionary tree. Definition The expression is most easily illustrated by a cladogram: Taxon A and ...

to ''

''. The Cladoselachidae is in turn sister to the clade containing

Holocephali Holocephali (Sometimes spelled Holocephala; Romanization of Greek, Greek for "complete head" in reference to the fusion of Palatoquadrate, upper jaw with the rest of the skull) is a Subclass (biology), subclass of Chondrichthyes, cartilaginous fi ...

and

Symmoriida Symmoriiformes is an extinct order of cartilaginous fish. Originally named Symmoriida by Zangerl (1981), the name has since been corrected to Symmoriiformes to avoid confusion with a family. The symmoriiform fossils record begins during the late ...

. The results of their

phylogenetic analyses In biology, phylogenetics () is the study of the evolutionary history of life using observable characteristics of organisms (or genes), which is known as Computational phylogenetics, phylogenetic inference. It infers the relationship among organ ...

are shown in the

cladogram A cladogram (from Greek language, Greek ''clados'' "branch" and ''gramma'' "character") is a diagram used in cladistics to show relations among organisms. A cladogram is not, however, an Phylogenetic tree, evolutionary tree because it does not s ...

below:

References

{{Taxonbar, from=Q117789893 Symmoriiformes Monotypic prehistoric cartilaginous fish genera Fossil taxa described in 2023