''Lusovenator'' (meaning "Portuguese hunter") is a genus of carcharodontosaurian

theropod Theropoda (; from ancient Greek , (''therion'') "wild beast"; , (''pous, podos'') "foot"">wiktionary:ποδός"> (''pous, podos'') "foot" is one of the three major groups (clades) of dinosaurs, alongside Ornithischia and Sauropodom ...

dinosaur Dinosaurs are a diverse group of reptiles of the clade Dinosauria. They first appeared during the Triassic Geological period, period, between 243 and 233.23 million years ago (mya), although the exact origin and timing of the #Evolutio ...

from the

Late Jurassic The Late Jurassic is the third Epoch (geology), epoch of the Jurassic Period, and it spans the geologic time scale, geologic time from 161.5 ± 1.0 to 143.1 ± 0.8 million years ago (Ma), which is preserved in Upper Jurassic stratum, strata.Owen ...

(

Kimmeridgian In the geologic timescale, the Kimmeridgian is an age in the Late Jurassic Epoch and a stage in the Upper Jurassic Series. It spans the time between 154.8 ±0.8 Ma and 149.2 ±0.7 Ma (million years ago). The Kimmeridgian follows the Oxfordian ...

) Praia de Amoreira Porto-Novo Member and the Late Jurassic-

Early Cretaceous The Early Cretaceous (geochronology, geochronological name) or the Lower Cretaceous (chronostratigraphy, chronostratigraphic name) is the earlier or lower of the two major divisions of the Cretaceous. It is usually considered to stretch from 143.1 ...

(

Tithonian In the geological timescale, the Tithonian is the latest age (geology), age of the Late Jurassic Epoch and the uppermost stage (stratigraphy), stage of the Upper Jurassic Series. It spans the time between 149.2 ±0.7 annum, Ma and 143.1 ±0.6 (mi ...

Berriasian In the geological timescale, the Berriasian is an age/ stage of the Early/Lower Cretaceous. It is the oldest subdivision in the entire Cretaceous. It has been taken to span the time between 143.1 ±0.6 Ma and 137.05 ± 0.2 (million years ago) ...

) Assenta Member of the

Lourinhã Formation The Lourinhã Formation () is a fossil-rich geological formation in western Portugal, named for the municipality of Lourinhã. The formation is mostly Late Jurassic in age (Kimmeridgian/Tithonian), with the top of the formation extending into the ...

in present-day

Portugal Portugal, officially the Portuguese Republic, is a country on the Iberian Peninsula in Southwestern Europe. Featuring Cabo da Roca, the westernmost point in continental Europe, Portugal borders Spain to its north and east, with which it share ...

. It includes one species, ''Lusovenator santosi''.

Discovery and naming

The

holotype A holotype (Latin: ''holotypus'') is a single physical example (or illustration) of an organism used when the species (or lower-ranked taxon) was formally described. It is either the single such physical example (or illustration) or one of s ...

was discovered during the 1980s by José Joaquim dos Santos, and he donated his fossil collection to the Sociedade de História Natural around thirty years later. The unnamed holotype was described and placed in the

Allosauroidea Carnosauria is an extinct group of carnivorous theropod dinosaurs that lived during the Jurassic and Cretaceous periods. While Carnosauria was historically considered largely synonymous with Allosauroidea, some recent studies have revived Carn ...

in 2017. The neotype, was described in 2019 and both specimens were placed within Carcharodontosauria. The species ''Lusovenator santosi'' was named and described in 2020. The known remains consist of the holotype SHN.036 - "a partial postcranial skeleton preserving the odontoid, the atlantal intercentrum, a cervical vertebra, isolated cervical neural spines, dorsal vertebrae, fragments of sacral vertebrae, caudal vertebrae, chevrons, fragments of cervical and dorsal ribs, the right ilium, both pubes, and ischia", thought to represent a juvenile individual - and the neotype SHN.019, "a partial skeleton represented by a series of articulated caudal vertebrae and an almost complete right pes".

Etymology

The generic name ''Lusovenator'' is derived from

Lusitania Lusitania (; ) was an ancient Iberian Roman province encompassing most of modern-day Portugal (south of the Douro River) and a large portion of western Spain (the present Extremadura and Province of Salamanca). Romans named the region after th ...

, the province of the

Roman Empire The Roman Empire ruled the Mediterranean and much of Europe, Western Asia and North Africa. The Roman people, Romans conquered most of this during the Roman Republic, Republic, and it was ruled by emperors following Octavian's assumption of ...

that roughly matches present-day Portugal, affixed with the Latin ''venator'', meaning "hunter". The specific epithet, ''santosi'', refers to José Joaquim dos Santos, who discovered the remains.

Classification

The describers ran a phylogenetic analysis and recovered ''Lusovenator'' within the clade Carcharodontosauria, outside

Carcharodontosauridae Carcharodontosauridae (carcharodontosaurids; from the Greek καρχαροδοντόσαυρος, ''carcharodontósauros'': "shark-toothed lizards") is a group of carnivorous theropod dinosaurs. In 1931, Ernst Stromer named Carcharodontosaurida ...

and

Neovenatoridae Carnosauria is an extinct group of carnivorous theropod dinosaurs that lived during the Jurassic and Cretaceous periods. While Carnosauria was historically considered largely synonymous with Allosauroidea, some recent studies have revived Carn ...

. Their

cladogram A cladogram (from Greek language, Greek ''clados'' "branch" and ''gramma'' "character") is a diagram used in cladistics to show relations among organisms. A cladogram is not, however, an Phylogenetic tree, evolutionary tree because it does not s ...

is shown below: Cau (2024) found ''Lusovenator'' to be in the family

, forming a clade with '' Sauroniops'', '' Veterupristisaurus'', '' Eocarcharia'', and '' Concavenator''.

References

{{Taxonbar, from=Q97149714 Carnosauria Dinosaur genera Tithonian dinosaurs Lourinhã Formation Fossil taxa described in 2020 Dinosaurs of Portugal