Luciola
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''Luciola'' is a
genus Genus (; : genera ) is a taxonomic rank above species and below family (taxonomy), family as used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In bino ...
of flashing
fireflies The Lampyridae are a family of elateroid beetles with more than 2,000 described species, many of which are light-emitting. They are soft-bodied beetles commonly called fireflies, lightning bugs, or glowworms for their conspicuous production ...
in the
family Family (from ) is a Social group, group of people related either by consanguinity (by recognized birth) or Affinity (law), affinity (by marriage or other relationship). It forms the basis for social order. Ideally, families offer predictabili ...
Lampyridae. They are especially well known from Japan and are often called Japanese fireflies, but their members range farther into Asia and reach southern Europe (Italy, France, Spain, Portugal, among other countries) and Africa. This genus is traditionally held to extend to Australia, but these
species A species () is often defined as the largest group of organisms in which any two individuals of the appropriate sexes or mating types can produce fertile offspring, typically by sexual reproduction. It is the basic unit of Taxonomy (biology), ...
do not seem to belong herein (see below for details).


Biological description

Unlike some other fireflies, the females of ''Luciola'' are fully winged.
Ecological Ecology () is the natural science of the relationships among living organisms and their environment. Ecology considers organisms at the individual, population, community, ecosystem, and biosphere levels. Ecology overlaps with the closely re ...
ly, the genus is remarkable in that the
larva A larva (; : larvae ) is a distinct juvenile form many animals undergo before metamorphosis into their next life stage. Animals with indirect development such as insects, some arachnids, amphibians, or cnidarians typically have a larval phase ...
e of several species are
aquatic Aquatic means relating to water; living in or near water or taking place in water; does not include groundwater, as "aquatic" implies an environment where plants and animals live. Aquatic(s) may also refer to: * Aquatic animal, either vertebrate ...
. The adults of the typical ''Luciola'' are similar to those of '' Atyphella'' which was formerly included in this genus. They can be easily distinguished by the males'
aedeagus An aedeagus ( or aedeagi) is a reproductive organ of male arthropods through which they secrete sperm from the testes during copulation (zoology), copulation with a female. It can be thought of as the insect equivalent of a mammal's penis, th ...
however, which in ''Luciola'' proper has large
lateral Lateral is a geometric term of location which may also refer to: Biology and healthcare * Lateral (anatomy), a term of location meaning "towards the side" * Lateral cricoarytenoid muscle, an intrinsic muscle of the larynx * Lateral release ( ...
lobes that do not taper and have elongated, slender and flattened smaller lobes along their
ventral Standard anatomical terms of location are used to describe unambiguously the anatomy of humans and other animals. The terms, typically derived from Latin or Greek roots, describe something in its standard anatomical position. This position prov ...
margin. The middle lobe of the males' aedeagus is very slim and has a point before the tip. By contrast, the
Australia Australia, officially the Commonwealth of Australia, is a country comprising mainland Australia, the mainland of the Australia (continent), Australian continent, the island of Tasmania and list of islands of Australia, numerous smaller isl ...
n group resembles '' Pyrophanes'' in its lateral lobes of the aedeagus being small and not visible from beneath. But unlike in ''Pyrophanes'', the sheath of the aedeagus lacks paraprocts, and the seventh ventrite of the
abdomen The abdomen (colloquially called the gut, belly, tummy, midriff, tucky, or stomach) is the front part of the torso between the thorax (chest) and pelvis in humans and in other vertebrates. The area occupied by the abdomen is called the abdominal ...
lacks the hairy lobes and points at its hind margin.Fu & Ballantyne (2008) Two Japanese species of ''Luciola'', collectively known as ''hotaru'' (蛍), are significant in
Japanese culture Japanese culture has changed greatly over the millennia, from the country's prehistoric Jōmon period, to its contemporary modern culture, which absorbs influences from Asia and other regions of the world. Since the Jomon period, ancestral ...
and
folklore Folklore is the body of expressive culture shared by a particular group of people, culture or subculture. This includes oral traditions such as Narrative, tales, myths, legends, proverbs, Poetry, poems, jokes, and other oral traditions. This also ...
. They are symbols of the ''
hitodama In Japanese folklore, hitodama ( Japanese ; meaning "human soul") are balls of fire that mainly float in the middle of the night.広辞苑 第五版 p.2255 「人魂」 They are said to be "souls of the dead that have separated from their bodies", ...
'' (人魂 or 人玉), the
soul The soul is the purported Mind–body dualism, immaterial aspect or essence of a Outline of life forms, living being. It is typically believed to be Immortality, immortal and to exist apart from the material world. The three main theories that ...
s of the newly-dead. See also the explanations at the article on the movie ''
Hotaru no Haka is a 1988 Japanese animated war film written and directed by Isao Takahata, and produced by Studio Ghibli. It is based on the 1967 semi-autobiographical short story of the same name by Akiyuki Nosaka. Set in the city of Kobe, Japan, it tells ...
'' ("Grave of the Fireflies") for a discussion of the cultural significance of the ''hotaru''.


Systematics

''Luciola'' in the narrow sense appears to be most closely related to a rather basal group of Luciolinae, including '' Bourgeoisia'' and '' Lampyroidea'' and '' Hotaria''. Their relationships are not well resolved and ''Hotaria'' is sometimes merged with ''Luciola''. On the other hand, the genera '' Atyphella'' and '' Pygoluciola'', which for some time were included in ''Luciola'', is now again recognized as distinct. The internal
systematics Systematics is the study of the diversification of living forms, both past and present, and the relationships among living things through time. Relationships are visualized as evolutionary trees (synonyms: phylogenetic trees, phylogenies). Phy ...
of ''Luciola'' are somewhat in need of revision, and it is not certain whether these species are all correctly assigned to this genus. For example, the
Australia Australia, officially the Commonwealth of Australia, is a country comprising mainland Australia, the mainland of the Australia (continent), Australian continent, the island of Tasmania and list of islands of Australia, numerous smaller isl ...
n species appear to be separate and not very closely related to the other members of this genus, but rather to '' Colophotia'', '' Pteroptyx'' and '' Pyrophanes''. Whether the
larva A larva (; : larvae ) is a distinct juvenile form many animals undergo before metamorphosis into their next life stage. Animals with indirect development such as insects, some arachnids, amphibians, or cnidarians typically have a larval phase ...
e of these can be recognized by the share soft
terga A ''tergum'' (Latin for "the back"; : ''terga'', associated adjective tergal) is the dorsal ('upper') portion of an arthropod segment other than the head. The anterior edge is called the 'base' and posterior edge is called the 'apex' or 'margin'. ...
lacking paranota, or whether this is a
plesiomorph In phylogenetics, a plesiomorphy ("near form") and symplesiomorphy are synonyms for an ancestral Phenotypic trait, character shared by all members of a clade, which does not distinguish the clade from other clades. Plesiomorphy, symplesiomorph ...
ic trait also present in other Luciolinae, remains to be determined. ''L. deplanata'' might be worthy of separation in a
monotypic In biology, a monotypic taxon is a taxonomic group (taxon) that contains only one immediately subordinate taxon. A monotypic species is one that does not include subspecies or smaller, infraspecific taxa. In the case of genera, the term "unisp ...
genus '' Photuroluciola''. But considering the fact that many species remain essentially unstudied, a 2008 review of the genus concluded that:
"A phylogenetic analysis ..may give indications for subdivisions within ''Luciola''. However, there are elements of the cart and the horse here – which should come first? Break up ''Luciola'' based on narrow analyses, or wait until more analyses are available? It is our contention that those Luciolinae species which do not fit elsewhere should either be described as ''Luciola'' sp. or remain undescribed until such time as phylogenetic analyses indicate a repeatable pattern of subdivisions with ''Luciola''. Any other action would be, in our opinion, premature."
In 2010, ''leii'', ''ficta'', ''hydrophila'' and ''lateralis'' were transferred to the genus '' Aquatica''.


Species

These species belong to the genus ''Luciola'': * '' Luciola affinis'' Ritsema, 1875 (Australia) * '' Luciola amplipennis'' Fairmaire, 1880 (Africa) * '' Luciola angusticollis'' E. Olivier, 1886 (Southern Asia) * '' Luciola antennalis'' Bourgeois, 1905 (Southern Asia) * '' Luciola antica'' Boisduval, 1835 (Oceania) * '' Luciola antipodum'' Bourgeois, 1884 (Australia) * ''
Luciola apicalis ''Luciola'' is a genus of flashing fireflies in the family Lampyridae. They are especially well known from Japan and are often called Japanese fireflies, but their members range farther into Asia and reach southern Europe (Italy, France, Spain, ...
'' (Eschscholtz, 1822) (Southern Asia) * '' Luciola aquilaclara'' Ballantyne, 2013 (Australia) * '' Luciola atriceps'' Pic, 1916 (Africa) * '' Luciola atrimembris'' Pic, 1933 (Southern Asia) * '' Luciola atripes'' Pic, 1929 (Southern Asia) * ''
Luciola atritarsis ''Luciola'' is a genus of flashing fireflies in the family Lampyridae. They are especially well known from Japan and are often called Japanese fireflies, but their members range farther into Asia and reach southern Europe (Italy, France, Spain, ...
'' Pic, 1916 (Africa) * '' Luciola atriventris'' Pic, 1925 (Africa) * '' Luciola aurantiaca'' Pic, 1927 (Southern Asia) * '' Luciola auritula'' E. Olivier, 1910 (Southern Asia) * '' Luciola basipennis'' Pic, 1958 (Africa) * ''
Luciola bicoloriceps ''Luciola'' is a genus of flashing fireflies in the family Lampyridae. They are especially well known from Japan and are often called Japanese fireflies, but their members range farther into Asia and reach southern Europe (Italy, France, Spain, ...
'' Pic, 1924 (Southern Asia) * '' Luciola bimaculata'' Klug, 1855 (Africa) * '' Luciola bimaculicollis'' (Boheman, 1851) (Africa) * ''
Luciola binhana ''Luciola'' is a genus of flashing fireflies in the family Lampyridae. They are especially well known from Japan and are often called Japanese fireflies, but their members range farther into Asia and reach southern Europe (Italy, France, Spain, ...
'' Pic, 1927 (Southern Asia) * '' Luciola binotata'' Pic, 1951 (Africa) * '' Luciola biscutellata'' Fairmaire, 1896 (Africa) * '' Luciola bomansi'' Pic, 1956 (Africa) * '' Luciola bourgeoisi'' E. Olivier, 1895 (Southern Asia) * ''
Luciola burgeoni ''Luciola'' is a genus of flashing firefly, fireflies in the family (biology), family Lampyridae. They are especially well known from Japan and are often called Japanese fireflies, but their members range farther into Asia and reach southern Eur ...
'' Pic, 1931 (Africa) * '' Luciola caffra'' (Boheman, 1851) (Africa) * '' Luciola candezei'' E. Olivier, 1902 (Southern Asia) * '' Luciola capensis'' (Fabricius, 1775) (Africa) * '' Luciola carinaticollis'' Pic, 1952 (Africa) * '' Luciola cavifrons'' Fairmaire, 1900 (Africa) * '' Luciola chapaensis'' Pic, 1923 (Southern Asia) * '' Luciola christyi'' Pic, 1931 (Africa) * '' Luciola cincta'' (Fabricius, 1787) (Southern Asia) * '' Luciola cincticollis'' Klug, 1855 (Africa) * '' Luciola cingulata'' E. Olivier, 1885 (Southern Asia) * '' Luciola cisteloides'' Klug, 1855 (Africa) * '' Luciola clara'' E. Olivier, 1907 (Southern Asia) * '' Luciola collarti'' Pic, 1931 (Africa) * '' Luciola complanata'' Gorham, 1895 (Southern Asia) * '' Luciola completelimbata'' Pic, 1952 (Africa) * '' Luciola concreta'' E. Olivier, 1907 (Southern Asia) * '' Luciola congoana'' E. Olivier, 1895 (Africa) * ''
Luciola consobrina ''Luciola'' is a genus of flashing fireflies in the family Lampyridae. They are especially well known from Japan and are often called Japanese fireflies, but their members range farther into Asia and reach southern Europe (Italy, France, Spain, ...
'' Hicker, 1942 (Africa) * '' Luciola costulata'' Fairmaire, 1886 (Africa) * '' Luciola coxalis'' E. Olivier, 1883 (Africa) * '' Luciola cruciata'' Motschulsky, 1854 genji-botaru (大端黑螢) (Europe & Asia) * ''
Luciola curticollis ''Luciola'' is a genus of flashing fireflies in the family Lampyridae. They are especially well known from Japan and are often called Japanese fireflies, but their members range farther into Asia and reach southern Europe (Italy, France, Spain, ...
'' Pic, 1927 (Southern Asia) * '' Luciola curtithorax'' Pic, 1928 (Southern Asia) * '' Luciola davidis'' E. Olivier, 1895 (Southern Asia) * '' Luciola delauneyi'' Bourgeois, 1890 (Southern Asia) * '' Luciola deleta'' Fairmaire, 1896 (Africa) * '' Luciola deplanata'' Pic, 1929 (Southern Asia) * '' Luciola dilatata'' Pic, 1929 (Southern Asia) * ''
Luciola dilecta ''Luciola'' is a genus of flashing fireflies in the family Lampyridae. They are especially well known from Japan and are often called Japanese fireflies, but their members range farther into Asia and reach southern Europe (Italy, France, Spain, ...
'' E. Olivier, 1908 (Africa) * '' Luciola discicollis'' Laporte, 1833 (Africa) * '' Luciola discobrunnescens'' Pic, 1951 (Africa) * '' Luciola discoidalis'' Pic, 1931 (Africa) * ''
Luciola dregei ''Luciola'' is a genus of flashing fireflies in the family Lampyridae. They are especially well known from Japan and are often called Japanese fireflies, but their members range farther into Asia and reach southern Europe (Italy, France, Spain, ...
'' (Motschulsky, 1853) (Africa) * '' Luciola errans'' E. Olivier, 1902 (Africa) * '' Luciola exigua'' (Gyllenhal, 1817) (Southern Asia) * '' Luciola exoleta'' Motschulsky, 1854 (Africa) * '' Luciola exstincta'' E. Olivier, 1886 (Southern Asia) * '' Luciola extricans'' (Walker, 1858) (Southern Asia) * '' Luciola falsa'' E. Olivier, 1907 (Africa) * ''
Luciola fasciiventris ''Luciola'' is a genus of flashing fireflies in the family Lampyridae. They are especially well known from Japan and are often called Japanese fireflies, but their members range farther into Asia and reach southern Europe (Italy, France, Spain, ...
'' Fairmaire, 1898 (Africa) * '' Luciola filiformis'' E. Olivier, 1913 (Southern Asia) * '' Luciola fissicollis'' Fairmaire, 1891 (Southern Asia) * '' Luciola flava'' Pic, 1929 (Southern Asia) * '' Luciola flavescens'' (Boisduval, 1835) (Oceania) * '' Luciola flebilis'' E. Olivier, 1909 (Southern Asia) * ''
Luciola fukiensis ''Luciola'' is a genus of flashing fireflies in the family Lampyridae. They are especially well known from Japan and are often called Japanese fireflies, but their members range farther into Asia and reach southern Europe (Italy, France, Spain, ...
'' Pic, 1955 (Southern Asia) * '' Luciola fusca'' (Motschulsky, 1854) (Africa) * '' Luciola fuscula'' (Boheman, 1851) (Africa) * '' Luciola gaiffei'' Alluaud, 1899 (Africa) * '' Luciola galactopyga'' Fairmaire, 1884 (Africa) * '' Luciola gaudens'' E. Olivier, 1910 (Africa) * ''
Luciola gemina ''Luciola'' is a genus of flashing fireflies in the family Lampyridae. They are especially well known from Japan and are often called Japanese fireflies, but their members range farther into Asia and reach southern Europe (Italy, France, Spain, ...
'' E. Olivier, 1905 (Africa) * '' Luciola geniculata'' E. Olivier, 1902 (Africa) * '' Luciola ghesquierei'' Pic, 1934 (Africa) * ''
Luciola gigantea ''Luciola'' is a genus of flashing fireflies in the family Lampyridae. They are especially well known from Japan and are often called Japanese fireflies, but their members range farther into Asia and reach southern Europe (Italy, France, Spain, ...
'' Pic, 1916 (Africa) * '' Luciola gigas'' E. Olivier, 1888 (Africa and Southern Asia) * '' Luciola gowdeyi'' Pic, 1933 (Africa) * '' Luciola grandjeani'' Pic, 1931 (Africa) * '' Luciola guerini'' Laporte, 1833 (Australia) * '' Luciola hirticeps'' E. Olivier, 1888 (Southern Asia) * '' Luciola horni'' Bourgeois, 1905 (Southern Asia) * '' Luciola hypocrita'' E. Olivier, 1888 (Australia and Oceania) * '' Luciola ibukiyamana'' Nawa, 1921 (Southern Asia) * '' Luciola imbellis'' E. Olivier, 1908 (Africa) * '' Luciola imerinae'' Bourgeois, 1899 (Africa) * '' Luciola immarginata'' Bourgeois, 1890 (Southern Asia) * '' Luciola impedita'' E. Olivier, 1902 (Southern Asia) * '' Luciola incerta'' (Boisduval, 1835) (Oceania) * '' Luciola infuscata'' (Erichson, 1834) (Southern Asia) * '' Luciola inlateralis'' Pic, 1952 (Africa) * '' Luciola innotatipennis'' Pic, 1916 (Africa) * ''
Luciola insignis ''Luciola'' is a genus of flashing fireflies in the family Lampyridae. They are especially well known from Japan and are often called Japanese fireflies, but their members range farther into Asia and reach southern Europe (Italy, France, Spain, ...
'' E. Olivier, 1883 (Africa) * '' Luciola intermedia'' Fairmaire, 1896 (Africa) * ''
Luciola intricata ''Luciola intricata'', is a species of firefly beetle found in Sri Lanka. Body is about 6.1 mm long. Dorsum yellowish brown with dark tipped elytral apices. Head dingy yellowish brown with a darker brown median line. Labrum brown brownish. An ...
'' (Walker, 1858) (Southern Asia) * '' Luciola italica'' (Linnaeus, 1767) (Europe & Northern Asia) * '' Luciola japonica'' (Thunberg, 1784) (Africa and Southern Asia) * '' Luciola jeanvoinei'' Pic, 1927 (Southern Asia) * '' Luciola jengai'' Nada, 2019 (Southern Asia) * '' Luciola judaica'' E. Olivier, 1884 (Europe & Northern Asia) * '' Luciola kagiana'' Matsumura, 1928 (Southern Asia) * '' Luciola kervillei'' E. Olivier, 1909 (Europe & Northern Asia) * '' Luciola klapperichi'' Pic, 1955 (Southern Asia) * '' Luciola krugeri'' E. Olivier, 1905 (Africa) * '' Luciola kuroiwae'' Matsumura, 1918 (Southern Asia) * '' Luciola laeta'' Gerstaecker, 1871 (Africa) * '' Luciola lambkinae'' Keller & Ballantyne, 2021 (Southern Asia) * '' Luciola latediscoidalis'' Pic, 1952 (Africa) * '' Luciola latelimbata'' Pic, 1956 (Africa) * '' Luciola laticollis'' Gorham, 1883 (Southern Asia) * '' Luciola leroyi'' Pic, 1952 (Africa) * '' Luciola limbalis'' Fairmaire, 1889 (Southern Asia) * '' Luciola limbatipennis'' Pic, 1923 (Southern Asia) * '' Luciola longula'' E. Olivier, 1914 (Africa) * ''
Luciola lucernula ''Luciola'' is a genus of flashing fireflies in the family Lampyridae. They are especially well known from Japan and are often called Japanese fireflies, but their members range farther into Asia and reach southern Europe (Italy, France, Spain, ...
'' Reiche, 1850 (Africa) * '' Luciola lusitanica'' (Charpentier, 1825) (Europe & Northern Asia) * '' Luciola maculiscuta'' Fairmaire, 1884 (Africa) * '' Luciola madagascariensis'' (Guérin-Méneville, 1831) (Africa) * '' Luciola magambae'' McDermott, 1963 (Africa) * '' Luciola maindroni'' Pic, 1927 (Southern Asia) * '' Luciola major'' Pic, 1952 (Africa) * '' Luciola mareei'' Pic, 1952 (Africa) * '' Luciola mediodivisa'' Pic, 1952 (Africa) * '' Luciola mediojuncta'' Pic, 1952 (Africa) * '' Luciola melancholica'' E. Olivier, 1913 (Australia) * '' Luciola melanura'' Laporte, 1833 (Africa) * '' Luciola mendax'' E. Olivier, 1905 (Africa) * '' Luciola mocquerysi'' E. Olivier, 1902 (Africa) * '' Luciola mongolica'' Motschulsky, 1860 (Europe & Northern Asia) * '' Luciola multicostulata'' Pic, 1927 (Southern Asia) * '' Luciola mutata'' Pic, 1939 (Africa) * '' Luciola niah'' Jusoh, 2019 (Southern Asia) * '' Luciola nicolleri'' Bugnion, 1922 (Southern Asia) * ''
Luciola nigripes ''Luciola'' is a genus of flashing fireflies in the family Lampyridae. They are especially well known from Japan and are often called Japanese fireflies, but their members range farther into Asia and reach southern Europe (Italy, France, Spain, ...
'' Gorham, 1903 (Southern Asia) * '' Luciola nigritarsis'' Pic, 1927 (Southern Asia) * '' Luciola nitidicollis'' Pic, 1916 (Africa) * '' Luciola nitidula'' (Fabricius, 1781) (Africa) * '' Luciola noctivaga'' E. Olivier, 1907 (Southern Asia) * '' Luciola notaticollis'' Pic, 1914 (Southern Asia) * '' Luciola novaki'' Müller, 1946 (Europe & Northern Asia) * '' Luciola obscura'' Pic, 1928 (Southern Asia) * '' Luciola obscuripennis'' Klug, 1855 (Africa) * '' Luciola ochracea'' Gorham, 1895 (Southern Asia) * '' Luciola oculofissa'' Ballantyne, 2013 (Australia) * '' Luciola olivieri'' Bourgeois, 1908 (Africa) * '' Luciola overlaeti'' Pic, 1931 (Africa) * '' Luciola owadai'' Satô & Kimura, 1994 - Kumejima-botaru - (Southern Asia) * '' Luciola pallida'' Kolbe, 1883 (Africa) * '' Luciola pallidipes'' Pic, 1928 (Southern Asia) * '' Luciola papariensis'' Doi, 1932 (Southern Asia) * '' Luciola parvula'' Kiesenwetter, 1874 (Southern Asia) * ''
Luciola pattersoni ''Luciola'' is a genus of flashing fireflies in the family Lampyridae. They are especially well known from Japan and are often called Japanese fireflies, but their members range farther into Asia and reach southern Europe (Italy, France, Spain, ...
'' Pic, 1935 (Africa) * '' Luciola perpetiuscula'' Kolbe, 1887 (Africa) * '' Luciola perspicua'' E. Olivier, 1907 (Southern Asia) * '' Luciola picea'' Gorham, 1882 (Southern Asia) * '' Luciola pici'' McDermott, 1966 (Southern Asia) * '' Luciola picticollis'' Kiesenwetter, 1874 (Southern Asia) * '' Luciola pieli'' Pic, 1939 (Southern Asia) * '' Luciola pouilloni'' Pic, 1931 (Africa) * '' Luciola praestans'' E. Olivier, 1896 (Africa) * '' Luciola praetermissa'' E. Olivier, 1909 (Africa) * '' Luciola pumila'' (Boheman, 1851) (Africa) * '' Luciola puncticollis'' Laporte, 1833 (Africa) * '' Luciola quadrimaculata'' Pic, 1952 (Africa) * ''
Luciola recticollis ''Luciola'' is a genus of flashing fireflies in the family Lampyridae. They are especially well known from Japan and are often called Japanese fireflies, but their members range farther into Asia and reach southern Europe (Italy, France, Spain, ...
'' E. Olivier, 1900 (Southern Asia) * '' Luciola robusticeps'' Pic, 1928 (Southern Asia) * '' Luciola roseicolllis'' Pic, 1933 (Southern Asia) * '' Luciola ruficollis'' (Guérin-Méneville, 1830) (Australia) * '' Luciola rugiceps'' E. Olivier, 1886 (Southern Asia) * '' Luciola satoi'' Jeng & Yang, 2003 (Southern Asia) * '' Luciola scheitzae'' Pic, 1952 (Africa) * '' Luciola semiventralis'' Fairmaire, 1899 (Africa) * '' Luciola semivittata'' Pic, 1952 (Africa) * '' Luciola septemmaculata'' E. Olivier, 1888 (Africa) * '' Luciola singapura'' Jusoh & Ballantyne, 2021 (Singapore) * '' Luciola sordida'' E. Olivier, 1905 (Southern Asia) * '' Luciola spectralis'' Gorham, 1880 (Australia) * '' Luciola spilodera'' Fairmaire, 1898 (Africa) * '' Luciola stanleyi'' E. Olivier, 1913 (Africa) * '' Luciola stigmaticollis'' Fairmaire, 1887 (Southern Asia) * '' Luciola subunicolor'' Pic, 1931 (Africa) * '' Luciola sudra'' Gorham, 1903 (Southern Asia) * '' Luciola syriaca'' (Costa, 1875) (Southern Asia) * '' Luciola tabida'' Gorham, 1880 (Africa) * ''
Luciola taeniaticollis ''Luciola'' is a genus of flashing fireflies in the family Lampyridae. They are especially well known from Japan and are often called Japanese fireflies, but their members range farther into Asia and reach southern Europe (Italy, France, Spain, ...
'' Fairmaire, 1886 (Africa) * '' Luciola tenuicornis'' E. Olivier, 1885 (Australia) * '' Luciola testaceipes'' Pic, 1916 (Africa) * '' Luciola testaceiventris'' Pic, 1916 (Africa) * '' Luciola tetrasticta'' Fairmaire, 1888 (Africa) * '' Luciola tincticollis'' Gorham, 1895 (Southern Asia) * '' Luciola tiomana'' Ballantyne, 2019 (Southern Asia) * '' Luciola trimaculata'' Pic, 1916 (Africa) * '' Luciola trivandrensis'' Raj, 1947 (Southern Asia) * '' Luciola tsushimana'' Nakane, 1970 (Southern Asia) * '' Luciola tuberculata'' Yiu, 2017 (Southern Asia) * '' Luciola turneri'' Pic, 1934 (Africa) * '' Luciola umbratica'' E. Olivier, 1907 (Africa) * '' Luciola unmunsana'' Doi, 1931 (Southern Asia) * '' Luciola varia'' E. Olivier, 1908 (Southern Asia) * '' Luciola vittata'' Laporte, 1833 (Southern Asia) * '' Luciola vleeschouwersi'' Pic, 1952 (Africa) * '' Luciola xanthochroa'' Fairmaire, 1904 (Africa) * '' Luciola xanthura'' Gorham, 1880 (Southern Asia)


Footnotes


References

* (2006): A phylogenetic reassessment of the rare S. E. Asian firefly genus ''Pygoluciola'' Wittmer (Coleoptera: Lampyridae: Luciolinae). ''Raffles Bulletin of Zoology'' 53(1): 21–48
PDF fulltext
* (2002): A description of larvae and redescription of adults of the firefly ''Pteroptyx valida'' Olivier in Selangor, Malaysia (Coleoptera: Lampyridae: Luciolinae), with notes of Luciolinae larvae. ''Raffles Bulletin of Zoology'' 50(1): 101–109
PDF fulltext
* (2008): Taxonomy and behaviour of lucioline fireflies (Coleoptera: Lampyridae: Luciolinae) with redefinition and new species of ''Pygoluciola'' Wittmer from mainland China and review of ''Luciola'' LaPorte. ''
Zootaxa ''Zootaxa'' is a peer-reviewed scientific mega journal for animal taxonomists. It is published by Magnolia Press (Auckland, New Zealand). The journal was established by Zhi-Qiang Zhang in 2001 and new issues are published multiple times a week ...
'' 1733: 1–44
PDF abstract
* (2010): ''Aquatica'' gen. nov. from mainland China with a description of ''Aquatica Wuhana'' sp. nov. (Coleoptera: Lampyridae: Luciolinae). ''Zootaxa'' 2530: 1–18
PDF abstract
* (2007): Phylogeny of North American fireflies (Coleoptera: Lampyridae): Implications for the evolution of light signals. '' Mol. Phylogenet. Evol.'' 45(1): 33–49. (HTML abstract) {{Authority control Lampyridae Lampyridae genera Bioluminescent insects Taxa named by François-Louis Laporte, comte de Castelnau