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The latticed heath (''Chiasmia clathrata'') is a
moth Moths are a paraphyletic group of insects that includes all members of the order Lepidoptera that are not butterflies, with moths making up the vast majority of the order. There are thought to be approximately 160,000 species of moth, many of ...
of the family
Geometridae The geometer moths are moths belonging to the family Geometridae of the insect order Lepidoptera, the moths and butterflies. Their scientific name derives from the Ancient Greek ''geo'' γεω (derivative form of or "the earth"), and ''metr ...
, belonging to the subfamily
Ennominae Ennominae is the largest subfamily of the geometer moth family (Geometridae) with some 9,700 described species in 1,100 genera. They are usually a fairly small moths, though some (such as the peppered moth) grow to be considerably large. This s ...
, placed in the tribe
Macariini The Macariini are a tribe of geometer moths in the subfamily Ennominae. Though they share many traits with the Sterrhinae, this is probably plesiomorphic rather than indicative of a close relationship, and DNA sequence data points to the Boarm ...
. The genus was erected by
Carl Linnaeus Carl Linnaeus (; 23 May 1707 – 10 January 1778), also known after his ennoblement in 1761 as Carl von Linné Blunt (2004), p. 171. (), was a Swedish botanist, zoologist, taxonomist, and physician who formalised binomial nomenclature, ...
in his 1758 10th edition of ''Systema Naturae''.


Taxonomy

The current placement of the latticed heath in the genus ''
Chiasmia ''Chiasmia'' is a genus of moths in the family Geometridae. It was described by Jacob Hübner in 1823. Species Species of this genus include: *''Chiasmia abnormata'' (Prout, 1917) *'' Chiasmia abyssinica'' Krüger, 2001 *'' Chiasmia acutiapex'' ...
'' follows from the revision by Malcolm J. Scoble (2002) of the tribe Macariini, when he showed that true ''
Semiothisa ''Semiothisa'' is a genus of moths in the family Geometridae. It was erected by Jacob Hübner in 1818. Species Some species of this genus are: * '' Semiothisa albivia'' (Prout, 1915) * '' Semiothisa arenosa'' (Butler, 1875) * '' Semiothisa assi ...
'' species were restricted to the Americas. There are a number of described subspecies. Molecular work has confirmed the placement of the species within ''Chiasmia''.


Subspecies

Subspecies include: * ''Chiasmia clathrata clathrata'' (Linné) * ''Chiasmia clathrata centralasiae'' (Krulikowski, 1911) * ''Chiasmia clathrata djakonovi'' (Kardakoff, 1928) * ''Chiasmia clathrata kurilata'' (Bryk, 1942)


Distribution and habitat

This species can be found throughout Europe, the Near East,
North Africa North Africa, or Northern Africa is a region encompassing the northern portion of the African continent. There is no singularly accepted scope for the region, and it is sometimes defined as stretching from the Atlantic shores of Mauritania in t ...
, and east through
Russia Russia (, , ), or the Russian Federation, is a transcontinental country spanning Eastern Europe and Northern Asia. It is the largest country in the world, with its internationally recognised territory covering , and encompassing one-eigh ...
,
Siberia Siberia ( ; rus, Сибирь, r=Sibir', p=sʲɪˈbʲirʲ, a=Ru-Сибирь.ogg) is an extensive geographical region, constituting all of North Asia, from the Ural Mountains in the west to the Pacific Ocean in the east. It has been a part o ...
,
Amur River The Amur (russian: река́ Аму́р, ), or Heilong Jiang (, "Black Dragon River", ), is the world's tenth longest river, forming the border between the Russian Far East and Northeastern China ( Inner Manchuria). The Amur proper is long ...
, northern
Iran Iran, officially the Islamic Republic of Iran, and also called Persia, is a country located in Western Asia. It is bordered by Iraq and Turkey to the west, by Azerbaijan and Armenia to the northwest, by the Caspian Sea and Turkm ...
,
Kazakhstan Kazakhstan, officially the Republic of Kazakhstan, is a transcontinental country located mainly in Central Asia and partly in Eastern Europe. It borders Russia Russia (, , ), or the Russian Federation, is a transcontinental coun ...
, China,
Korea Korea ( ko, 한국, or , ) is a peninsular region in East Asia. Since 1945, it has been divided at or near the 38th parallel, with North Korea (Democratic People's Republic of Korea) comprising its northern half and South Korea (Republi ...
to Japan. It is a fairly common species in the
British Isles The British Isles are a group of islands in the North Atlantic Ocean off the north-western coast of continental Europe, consisting of the islands of Great Britain, Ireland, the Isle of Man, the Inner and Outer Hebrides, the Northern Isles (O ...
.Skinner, B., 1984. ''Colour Identification Guide to Moths of the British Isles'' These moths inhabit a range of open areas, including grassland, moorland, and waste ground.


Description


Adult

''Chiasmia clathrata'' has a
wingspan The wingspan (or just span) of a bird or an airplane is the distance from one wingtip to the other wingtip. For example, the Boeing 777–200 has a wingspan of , and a wandering albatross (''Diomedea exulans'') caught in 1965 had a wingspan ...
of 20–25 mm.''UKMoths''
/ref> In this rather variable species the background colour of the wings varies from yellowish to white, with a network of brown lines (hence its common name). These lines vary in thickness and sometimes the wings are almost entirely dark brown. File:Chiasmia clathrata - Géomètre à barreaux MHNT.jpg , Female – dorsal and ventral sides File: Chiasma clathrata02.jpg, Spread wings File: Semiothisa clathrata 02.jpg, Underside, at night


Larva

The final
instar An instar (, from the Latin '' īnstar'', "form", "likeness") is a developmental stage of arthropods, such as insects, between each moult (''ecdysis''), until sexual maturity is reached. Arthropods must shed the exoskeleton in order to grow o ...
larva A larva (; plural larvae ) is a distinct juvenile form many animals undergo before metamorphosis into adults. Animals with indirect development such as insects, amphibians, or cnidarians typically have a larval phase of their life cycle. Th ...
is pale green with white lines, including a strong lateral line and thin dorsal lines along the body. Abdominal segments A1 through A5 have a strong white line across the rear end of each segment that ends just above the lateral line.Porter, J., 1997. ''The Colour Identification Guide to Caterpillars of the British Isles''. Viking Press, Harmondsworth, Middlesex. xii + 275 pp.


Ecology

In the British Isles there are one or two generations annually, with adults seen at any time from May to September. These moths are diurnal but they can also be observed at night where they are attracted to
artificial light Lighting or illumination is the deliberate use of light to achieve practical or aesthetic effects. Lighting includes the use of both artificial light sources like lamps and light fixtures, as well as natural illumination by capturing dayligh ...
.Truxa, C., Fiedler, K. (2012), Attraction to light - from how far do moths (Lepidoptera) return to weak artificial sources of light? ''European Journal of Entomology'', 109 (1):77-84. Larvae feed on bedstraws (''
Galium mollugo ''Galium mollugo'', common name hedge bedstraw or false baby's breath, is a herbaceous perennial plant of the family Rubiaceae. It shares the name hedge bedstraw with the related European species, Galium album. Description ''Galium mollugo'' can ...
'', ''
Galium verum ''Galium verum'' (lady's bedstraw or yellow bedstraw) is a herbaceous perennial plant of the family Rubiaceae. It is widespread across most of Europe, North Africa, and temperate Asia from Israel, Lebanon and Turkey to Japan and Kamchatka. It ...
'') and various
legumes A legume () is a plant in the family Fabaceae (or Leguminosae), or the fruit or seed of such a plant. When used as a dry grain, the seed is also called a pulse. Legumes are grown agriculturally, primarily for human consumption, for livestock fo ...
such as clovers (''
Trifolium medium ''Trifolium medium'', the zigzag clover, is a flowering plant species in the bean family Fabaceae. It is similar in appearance to red clover, ''Trifolium pratense'', but the leaflets are narrower and have no white markings and the narrow stipule ...
'', ''
Trifolium pratense ''Trifolium pratense'', the red clover, is a herbaceous species of flowering plant in the bean family Fabaceae, native to Europe, Western Asia, and northwest Africa, but planted and naturalized in many other regions. Description Red clov ...
''),
trefoil A trefoil () is a graphic form composed of the outline of three overlapping rings, used in architecture and Christian symbolism, among other areas. The term is also applied to other symbols with a threefold shape. A similar shape with four ring ...
s, lucerne (''
Medicago sativa Alfalfa () (''Medicago sativa''), also called lucerne, is a perennial flowering plant in the legume family Fabaceae. It is cultivated as an important forage crop in many countries around the world. It is used for grazing, hay, and silage, as ...
'') and
meadow vetchling ''Lathyrus pratensis'' or meadow vetchling, yellow pea, meadow pea and meadow pea-vine, is a perennial legume that grows to 1.2 m in height. The hermaphrodite flowers are pollinated by bees. As a perennial, this plant reproduces itself over man ...
, primarily in June and July and from mid-August through September, though in Ireland and northern Britain larvae occur in July and August. The species overwinters as a
pupa A pupa ( la, pupa, "doll"; plural: ''pupae'') is the life stage of some insects undergoing transformation between immature and mature stages. Insects that go through a pupal stage are holometabolous: they go through four distinct stages in their ...
.


Bibliography

* , 1986. ''Collins Guide to the Insects of Britain and Western Europe'' (Reprinted 1991) * , 2011. Phylogenetic relationships of selected European Ennominae (Lepidoptera: Geometridae). ''European Journal of Entomology'' 108: 267–273. * , 1997. ''The Colour Identification Guide to Caterpillars of the British Isles''. Viking Press, Harmondsworth, Middlesex. xii + 275 pp. * , 1984. ''Colour Identification Guide to Moths of the British Isles'' * , 2002. A review of the genera of Macariini with a revised classification of the tribe (Geometridae: Ennominae). ''Zoological Journal of the Linnean Society'' 134 (3): 257–315.


References


External links

* Paolo Mazzei, Daniel Morel, Raniero Panfil
''Moths and Butterflies of Europe and North Africa''''Lepiforum e.V.''
*Heiner Ziegle
''Schmetterlinge der Schweiz''
{{Taxonbar, from=Q25779 Macariini Moths described in 1758 Moths of Asia Moths of Europe Moths of Africa Taxa named by Carl Linnaeus Articles containing video clips