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''Labocania'' is a
genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of living and fossil organisms as well as viruses. In the hierarchy of biological classification, genus comes above species and below family. In binomial n ...
of
carnivorous A carnivore , or meat-eater (Latin, ''caro'', genitive ''carnis'', meaning meat or "flesh" and ''vorare'' meaning "to devour"), is an animal or plant whose food and energy requirements derive from animal tissues (mainly muscle, fat and other ...
theropod Theropoda (; ), whose members are known as theropods, is a dinosaur clade that is characterized by hollow bones and three toes and claws on each limb. Theropods are generally classed as a group of saurischian dinosaurs. They were ancestrally ...
dinosaur Dinosaurs are a diverse group of reptiles of the clade Dinosauria. They first appeared during the Triassic period, between 243 and 233.23  million years ago (mya), although the exact origin and timing of the evolution of dinosaurs is t ...
of uncertain affinities from
Baja California Baja California (; 'Lower California'), officially the Free and Sovereign State of Baja California ( es, Estado Libre y Soberano de Baja California), is a state in Mexico. It is the northernmost and westernmost of the 32 federal entities of Mex ...
,
Mexico Mexico ( Spanish: México), officially the United Mexican States, is a country in the southern portion of North America. It is bordered to the north by the United States; to the south and west by the Pacific Ocean; to the southeast by Guate ...
, which lived 73 million years ago, in the
Campanian The Campanian is the fifth of six ages of the Late Cretaceous Epoch on the geologic timescale of the International Commission on Stratigraphy (ICS). In chronostratigraphy, it is the fifth of six stages in the Upper Cretaceous Series. Campani ...
stage Stage or stages may refer to: Acting * Stage (theatre), a space for the performance of theatrical productions * Theatre, a branch of the performing arts, often referred to as "the stage" * ''The Stage'', a weekly British theatre newspaper * Sta ...
of the late
Cretaceous The Cretaceous ( ) is a geological period that lasted from about 145 to 66 million years ago (Mya). It is the third and final period of the Mesozoic Era, as well as the longest. At around 79 million years, it is the longest geological period of ...
Period. In the summer of 1970, the
National Geographic Society The National Geographic Society (NGS), headquartered in Washington, D.C., United States, is one of the largest non-profit scientific and educational organizations in the world. Founded in 1888, its interests include geography, archaeology, ...
and the
Los Angeles County Museum of Natural History The Natural History Museum of Los Angeles County is the largest natural and historical museum in the western United States. Its collections include nearly 35 million specimens and artifacts and cover 4.5 billion years of history. This large colle ...
organized a joint
paleontological Paleontology (), also spelled palaeontology or palæontology, is the scientific study of life that existed prior to, and sometimes including, the start of the Holocene epoch (roughly 11,700 years before present). It includes the study of fossi ...
expedition, led by
geologist A geologist is a scientist who studies the solid, liquid, and gaseous matter that constitutes Earth and other terrestrial planets, as well as the processes that shape them. Geologists usually study geology, earth science, or geophysics, alt ...
William J. Morris, to the Arroyo del Rosario in Baja California. While prospecting, volunteer Harley James Garbani discovered the skeleton of a theropod north of
Punta Baja Punta is an Afro-indigenous dance and cultural music originating in the Caribbean Island of Saint Vincent And The Grenadines by the Garifuna people before being exiled from the island. Which is also known as Yurumei. It has African and Araw ...
near Cerro Rayado. Garbani excavated the site in 1970 and 1971. The
type species In zoological nomenclature, a type species (''species typica'') is the species name with which the name of a genus or subgenus is considered to be permanently taxonomically associated, i.e., the species that contains the biological type specimen( ...
, ''Labocania anomala'', was described and named by
Ralph Molnar Ralph E. Molnar is a paleontologist who had been Curator of Mammals at the Queensland Museum and more recently associated with the Museum of Northern Arizona. He is also a research associate at the Texas natural Science Centre. He co-authored descr ...
in 1974. The generic name references the
La Bocana Roja Formation La Bocana Roja Formation is a geological formation Formation may refer to: Linguistics * Back-formation, the process of creating a new lexeme by removing or affixes * Word formation, the creation of a new word by adding affixes Mathematics an ...
, named after ''la Bocana Roja'', "the red
estuary An estuary is a partially enclosed coastal body of brackish water with one or more rivers or streams flowing into it, and with a free connection to the open sea. Estuaries form a transition zone between river environments and maritime environm ...
". The specific name means "anomalous" in
Latin Latin (, or , ) is a classical language belonging to the Italic branch of the Indo-European languages. Latin was originally a dialect spoken in the lower Tiber area (then known as Latium) around present-day Rome, but through the power ...
, in reference to the distinctive build. The
holotype A holotype is a single physical example (or illustration) of an organism, known to have been used when the species (or lower-ranked taxon) was formally described. It is either the single such physical example (or illustration) or one of seve ...
, LACM 20877, was found in a layer of the La Bocana Roja Formation, dating from the late Campanian, about 73 million years old. It consists of a very fragmentary skeleton with
skull The skull is a bone protective cavity for the brain. The skull is composed of four types of bone i.e., cranial bones, facial bones, ear ossicles and hyoid bone. However two parts are more prominent: the cranium and the mandible. In humans, t ...
elements, including a right quadrate, a left frontal, a piece of the left
maxilla The maxilla (plural: ''maxillae'' ) in vertebrates is the upper fixed (not fixed in Neopterygii) bone of the jaw formed from the fusion of two maxillary bones. In humans, the upper jaw includes the hard palate in the front of the mouth. The ...
, a fragment of the dentarium, a
chevron Chevron (often relating to V-shaped patterns) may refer to: Science and technology * Chevron (aerospace), sawtooth patterns on some jet engines * Chevron (anatomy), a bone * ''Eulithis testata'', a moth * Chevron (geology), a fold in rock lay ...
, the upper parts of both
ischia Ischia ( , , ) is a volcanic island in the Tyrrhenian Sea. It lies at the northern end of the Gulf of Naples, about from Naples. It is the largest of the Phlegrean Islands. Roughly trapezoidal in shape, it measures approximately east to wes ...
, the middle shaft of the right pubis, most of the second right
metatarsal The metatarsal bones, or metatarsus, are a group of five long bones in the foot, located between the tarsal bones of the hind- and mid-foot and the phalanges of the toes. Lacking individual names, the metatarsal bones are numbered from the med ...
, a pedal
phalanx The phalanx ( grc, φάλαγξ; plural phalanxes or phalanges, , ) was a rectangular mass military formation, usually composed entirely of heavy infantry armed with spears, pikes, sarissas, or similar pole weapons. The term is particular ...
and several loose
teeth A tooth ( : teeth) is a hard, calcified structure found in the jaws (or mouths) of many vertebrates and used to break down food. Some animals, particularly carnivores and omnivores, also use teeth to help with capturing or wounding prey, te ...
. The elements were not articulated, dispersed over a surface of about two square metres, and strongly weathered. The remains were mixed with the ribs of
Hadrosauroidea Hadrosauroidea is a clade or superfamily of ornithischian dinosaurs that includes the "duck-billed" dinosaurs, or hadrosaurids, and all dinosaurs more closely related to them than to '' Iguanodon''. Their remains have been recovered in Asia, Euro ...
. Though its exact size is hard to establish, ''Labocania'' was probably a medium-sized carnivore, about long.
Gregory S. Paul Gregory Scott Paul (born December 24, 1954) is an American freelance researcher, author and illustrator who works in paleontology, and more recently has examined sociology and theology. He is best known for his work and research on theropod dino ...
estimated its length at 7 meters (23 ft), its weight at 1.5 tonnes (1.65 short tons) in 2010. In 2016 Molina-Pérez and Larramendi gave a higher estimation of 8.2 meters (27 ft) and 2.6 tonnes (2.86 short tons). The cranial elements are very robust, and the frontals in particular are strongly thickened. The teeth of the maxilla are gradually recurving and rather flat; those of the
premaxilla The premaxilla (or praemaxilla) is one of a pair of small cranial bones at the very tip of the upper jaw of many animals, usually, but not always, bearing teeth. In humans, they are fused with the maxilla. The "premaxilla" of therian mammal has ...
do not have a D-shaped cross-section. Because ''Labocania'' is based on fragmentary material, its affinities are uncertain. Molnar noted certain similarities between ''Labocania'' and
tyrannosaurids Tyrannosauridae (or tyrannosaurids, meaning " tyrant lizards") is a family of coelurosaurian theropod dinosaurs that comprises two subfamilies containing up to thirteen genera, including the eponymous '' Tyrannosaurus''. The exact number of ...
, especially in the form of the
ischium The ischium () form ...
which features a low triangular obturator process and a circular lateral scar on the upper end, but he did not assign ''Labocania'' to any family, placing it as
Theropoda Theropoda (; ), whose members are known as theropods, is a dinosaur clade that is characterized by hollow bones and three toes and claws on each limb. Theropods are generally classed as a group of saurischian dinosaurs. They were ancestrally ca ...
''
incertae sedis ' () or ''problematica'' is a term used for a taxonomic group where its broader relationships are unknown or undefined. Alternatively, such groups are frequently referred to as "enigmatic taxa". In the system of open nomenclature, uncertain ...
''. Molnar especially compared ''Labocania'' with ''
Indosaurus ''Indosaurus'' () is a genus of carnivorous theropod dinosaur that lived in what is now India, about 69 to 66 million years ago during the Maastrichtian The Maastrichtian () is, in the ICS geologic timescale, the latest age (uppermost stage ...
'' and "
Chilantaisaurus ''Chilantaisaurus'' (" lizard") is a genus of large theropod dinosaur, possibly a neovenatorid or a primitive coelurosaur, from the Late Cretaceous Ulansuhai Formation of China (Turonian age, about 92 million years ago). The type species, ''C ...
" ''maortuensis'', later made the separate genus ''
Shaochilong ''Shaochilong'' (meaning "shark toothed dragon") is an extinct genus of carcharodontosaurid dinosaur from the mid-Cretaceous (Late Cenomanian to the end of the Turonian stage) Ulansuhai Formation of China (about 92 million years ago). The type ...
''. ''Labocania'' was considered as a possible tyrannosauroid in the 2004 review of the group by
Thomas R. Holtz Jr. Thomas Richard Holtz Jr. (born September 13, 1965) is an American vertebrate palaeontologist, author, and principal lecturer at the University of Maryland's Department of Geology. He has published extensively on the phylogeny, morphology, ecomorp ...
, who, however, pointed out that the similarities with the Tyrannosauridae were shared with the
Coelurosauria Coelurosauria (; from Greek, meaning "hollow tailed lizards") is the clade containing all theropod dinosaurs more closely related to birds than to carnosaurs. Coelurosauria is a subgroup of theropod dinosaurs that includes compsognathids, ty ...
in general—no tyrannosauroid
synapomorphies In phylogenetics, an apomorphy (or derived trait) is a novel character or character state that has evolved from its ancestral form (or plesiomorphy). A synapomorphy is an apomorphy shared by two or more taxa and is therefore hypothesized to hav ...
were present—and that ''Labocania'' also showed some
abelisaurid Abelisauridae (meaning "Abel's lizards") is a family (or clade) of ceratosaurian theropod dinosaurs. Abelisaurids thrived during the Cretaceous period, on the ancient southern supercontinent of Gondwana, and today their fossil remains are fou ...
traits such as the thick frontals and a reclining quadrate. On the other hand, the L-shaped chevron and the flattened outer side of the second metatarsal indicated a position in the
Tetanurae Tetanurae (/ˌtɛtəˈnjuːriː/ or "stiff tails") is a clade that includes most Theropoda, theropod dinosaurs, including Megalosauroidea, megalosauroids, Allosauroidea, allosauroids, Tyrannosauroidea, tyrannosauroids, Ornithomimosauria, ornitho ...
.


References

{{Taxonbar, from=Q1827222 Prehistoric neotheropods Campanian genus first appearances Campanian genus extinctions Late Cretaceous dinosaurs of North America Cretaceous Mexico Fossils of Mexico Fossil taxa described in 1974 Taxa named by Ralph Molnar