Kuehneosauridae is an

extinct Extinction is the termination of an organism by the death of its Endling, last member. A taxon may become Functional extinction, functionally extinct before the death of its last member if it loses the capacity to Reproduction, reproduce and ...

family of small, lizard-like gliding

diapsid Diapsids ("two arches") are a clade of sauropsids, distinguished from more primitive eureptiles by the presence of two holes, known as temporal fenestrae, in each side of their skulls. The earliest traditionally identified diapsids, the araeosc ...

s known from the

Triassic The Triassic ( ; sometimes symbolized 🝈) is a geologic period and system which spans 50.5 million years from the end of the Permian Period 251.902 million years ago ( Mya), to the beginning of the Jurassic Period 201.4 Mya. The Triassic is t ...

period of

Europe Europe is a continent located entirely in the Northern Hemisphere and mostly in the Eastern Hemisphere. It is bordered by the Arctic Ocean to the north, the Atlantic Ocean to the west, the Mediterranean Sea to the south, and Asia to the east ...

and

North America North America is a continent in the Northern Hemisphere, Northern and Western Hemisphere, Western hemispheres. North America is bordered to the north by the Arctic Ocean, to the east by the Atlantic Ocean, to the southeast by South Ameri ...

Description and systematics

They are distinguished from other diapsids by their 'wings' formed by elongated ribs. These allowed the animal to glide and parachute similar to living gliding lizards. They were most likely insectivorous, judging from their pin-like teeth. They are often, but not always, placed in the group

Lepidosauromorpha Lepidosauromorpha (in PhyloCode known as Pan-Lepidosauria) is a group of reptiles comprising all diapsids closer to lizards than to archosaurs (which include crocodiles and birds). The only living sub-group is the Lepidosauria, which contains tw ...

, though other studies have recovered them in other positions within

Sauria Sauria is the clade of diapsids containing the most recent common ancestor of Archosauria (which includes crocodilians and birds) and Lepidosauria (which includes squamates and the tuatara), and all its descendants. Since most molecular phyl ...

, including

Archosauromorpha Archosauromorpha ( Greek for "ruling lizard forms") is a clade of diapsid reptiles containing all reptiles more closely related to archosaurs (such as crocodilians and dinosaurs, including birds) than to lepidosaurs (such as tuataras, lizards, ...

. The oldest and most primitive known member is '' Pamelina'' from the

Early Triassic The Early Triassic is the first of three epochs of the Triassic Period of the geologic timescale. It spans the time between 251.9 Ma and Ma (million years ago). Rocks from this epoch are collectively known as the Lower Triassic Series, which ...

(

Olenekian In the geologic timescale, the Olenekian is an age (geology), age in the Early Triassic epoch (geology), epoch; in chronostratigraphy, it is a stage (stratigraphy), stage in the Lower Triassic series (stratigraphy), series. It spans the time betw ...

stage) of

Poland Poland, officially the Republic of Poland, is a country in Central Europe. It extends from the Baltic Sea in the north to the Sudetes and Carpathian Mountains in the south, bordered by Lithuania and Russia to the northeast, Belarus and Ukrai ...

, which already has vertebrae with characteristics consistent with gliding or parachuting. ''

Icarosaurus ''Icarosaurus'' (meaning "Icarus lizard") is an extinct genus of kuehneosaurid reptile from the Late Triassic (Norian age) Lower Lockatong Formation of New Jersey.Berg, T.M., ''et al''. (1983). Stratigraphic Correlation Chart of Pennsylvania: G ...

'' is known from a single specimen from the

Carnian The Carnian (less commonly, Karnian) is the lowermost stage (stratigraphy), stage of the Upper Triassic series (stratigraphy), Series (or earliest age (geology), age of the Late Triassic Epoch (reference date), Epoch). It lasted from 237 to 227.3 ...

-aged

Lockatong Formation The Triassic Lockatong Formation is a mapped bedrock unit in Pennsylvania, New Jersey, and New York. It is named after the Lockatong Creek in Hunterdon County, New Jersey. Description The Lockatong is defined as a light to dark gray, greenish-g ...

New Jersey New Jersey is a U.S. state, state located in both the Mid-Atlantic States, Mid-Atlantic and Northeastern United States, Northeastern regions of the United States. Located at the geographic hub of the urban area, heavily urbanized Northeas ...

. The

Late Triassic The Late Triassic is the third and final epoch (geology), epoch of the Triassic geologic time scale, Period in the geologic time scale, spanning the time between annum, Ma and Ma (million years ago). It is preceded by the Middle Triassic Epoch a ...

(

Norian The Norian is a division of the Triassic geological period, Period. It has the rank of an age (geology), age (geochronology) or stage (stratigraphy), stage (chronostratigraphy). It lasted from ~227.3 to Mya (unit), million years ago. It was prec ...

stage) kuehneosaurids from

England England is a Countries of the United Kingdom, country that is part of the United Kingdom. It is located on the island of Great Britain, of which it covers about 62%, and List of islands of England, more than 100 smaller adjacent islands. It ...

, '' Kuehneosaurus'' and ''

Kuehneosuchus ''Kuehneosuchus'' is an extinct genus of Late Triassic kuehneosaurid reptile known from the Late Triassic (Norian stage) of southwest England. It was named by Pamela Lamplugh Robinson in 1967 and the type and only species is ''Kuehneosuchus la ...

'', are very similar and can be distinguished from one another primarily on the length of their "wing" ribs, relatively short and massive in ''Kuehneosaurus'' but longer and more gracile in ''Kuehneosuchus. Kuehneosaurus'' was likely only capable of parachuting, while ''Kuehneosuchus'' could probably glide. '' Rhabdopelix'' may have been a kuehneosaurid; however, the fossils were lost, and the characteristics described are not entirely consistent with the other family members. Kuehneosaurids were not the first flying vertebrates. Weigeltisaurids ("Coelurosauravidae") that lived as long ago as the

Late Permian Late or LATE may refer to: Everyday usage * Tardy, or late, not being on time * Late (or the late) may refer to a person who is dead Music * ''Late'' (The 77s album), 2000 * Late (Alvin Batiste album), 1993 * Late!, a pseudonym used by Dave Groh ...

also had bone-supported membranous wings. But the anatomy of these groups is too different to consider them close relatives, although both weigeltisaurids and kuehneosaurids belong to diapsid reptiles.

Phylogeny

The

cladogram A cladogram (from Greek language, Greek ''clados'' "branch" and ''gramma'' "character") is a diagram used in cladistics to show relations among organisms. A cladogram is not, however, an Phylogenetic tree, evolutionary tree because it does not s ...

below follows a 2009 analysis by paleontologists Susan E. Evans and Magdalena Borsuk−Białynicka.

References

External links

* Prehistoric reptile families Triassic reptiles Olenekian first appearances Norian extinctions Taxa named by Alfred Romer {{triassic-reptile-stub

Description and systematics

Phylogeny

See also

References

External links