Keuper Marls
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The Keuper is a
lithostratigraphic Lithostratigraphy is a sub-discipline of stratigraphy, the geological science associated with the study of strata or rock layers. Major focuses include geochronology, comparative geology, and petrology. In general, strata are primarily igneous ...
unit (a sequence of
rock strata In geology and related fields, a stratum (: strata) is a layer of rock or sediment characterized by certain lithologic properties or attributes that distinguish it from adjacent layers from which it is separated by visible surfaces known as ...
) in the
subsurface In geology, bedrock is solid rock that lies under loose material (regolith) within the crust of Earth or another terrestrial planet. Definition Bedrock is the solid rock that underlies looser surface material. An exposed portion of bedro ...
of large parts of west and central
Europe Europe is a continent located entirely in the Northern Hemisphere and mostly in the Eastern Hemisphere. It is bordered by the Arctic Ocean to the north, the Atlantic Ocean to the west, the Mediterranean Sea to the south, and Asia to the east ...
. The Keuper consists of dolomite,
shale Shale is a fine-grained, clastic sedimentary rock formed from mud that is a mix of flakes of Clay mineral, clay minerals (hydrous aluminium phyllosilicates, e.g., Kaolinite, kaolin, aluminium, Al2Silicon, Si2Oxygen, O5(hydroxide, OH)4) and tiny f ...
s or
claystone Mudrocks are a class of fine-grained siliciclastic sedimentary rocks. The varying types of mudrocks include siltstone, claystone, mudstone and shale. Most of the particles of which the stone is composed are less than and are too small to ...
s and
evaporite An evaporite () is a water- soluble sedimentary mineral deposit that results from concentration and crystallization by evaporation from an aqueous solution. There are two types of evaporite deposits: marine, which can also be described as oce ...
s that were deposited during the
Middle Middle or The Middle may refer to: * Centre (geometry), the point equally distant from the outer limits. Places * Middle (sheading), a subdivision of the Isle of Man * Middle Bay (disambiguation) * Middle Brook (disambiguation) * Middle Creek ...
and
Late Triassic The Late Triassic is the third and final epoch (geology), epoch of the Triassic geologic time scale, Period in the geologic time scale, spanning the time between annum, Ma and Ma (million years ago). It is preceded by the Middle Triassic Epoch a ...
epoch In chronology and periodization, an epoch or reference epoch is an instant in time chosen as the origin of a particular calendar era. The "epoch" serves as a reference point from which time is measured. The moment of epoch is usually decided b ...
s (about ). The Keuper lies on top of the
Muschelkalk The Muschelkalk (German for "shell-bearing limestone"; ) is a sequence of sedimentary rock, sedimentary rock strata (a lithostratigraphy, lithostratigraphic unit) in the geology of central and western Europe. It has a Middle Triassic (240 to 230 m ...
and under the predominantly
Lower Jurassic The Early Jurassic Epoch (in chronostratigraphy corresponding to the Lower Jurassic Series) is the earliest of three epochs of the Jurassic Period. The Early Jurassic starts immediately after the Triassic–Jurassic extinction event, 201.3 Ma&nb ...
Lias or other Early Jurassic strata. The Keuper together with the Muschelkalk and the
Buntsandstein The Buntsandstein (German for ''coloured'' or ''colourful sandstone'') or Bunter sandstone is a lithostratigraphy, lithostratigraphic and allostratigraphy, allostratigraphic unit (a sequence of rock strata) in the Subsurface (geology), subsurface ...
form the Germanic Trias Group, a characteristic sequence of rock strata that gave the
Triassic The Triassic ( ; sometimes symbolized 🝈) is a geologic period and system which spans 50.5 million years from the end of the Permian Period 251.902 million years ago ( Mya), to the beginning of the Jurassic Period 201.4 Mya. The Triassic is t ...
its name. "Muschelkalk (geology)", Britannica Online Encyclopedia, October 2010, webpage:
EB-39


Exposure

The Upper Triassic is well exposed in
Swabia Swabia ; , colloquially ''Schwabenland'' or ''LĂ€ndle''; archaic English also Suabia or Svebia is a cultural, historic and linguistic region in southwestern Germany. The name is ultimately derived from the medieval Duchy of Swabia, one of ...
,
Franconia Franconia ( ; ; ) is a geographical region of Germany, characterised by its culture and East Franconian dialect (). Franconia is made up of the three (governmental districts) of Lower Franconia, Lower, Middle Franconia, Middle and Upper Franco ...
,
Alsace Alsace (, ; ) is a cultural region and a territorial collectivity in the Grand Est administrative region of northeastern France, on the west bank of the upper Rhine, next to Germany and Switzerland. In January 2021, it had a population of 1,9 ...
and
Lorraine Lorraine, also , ; ; Lorrain: ''Louréne''; Lorraine Franconian: ''Lottringe''; ; ; is a cultural and historical region in Eastern France, now located in the administrative region of Grand Est. Its name stems from the medieval kingdom of ...
and
Luxembourg Luxembourg, officially the Grand Duchy of Luxembourg, is a landlocked country in Western Europe. It is bordered by Belgium to the west and north, Germany to the east, and France on the south. Its capital and most populous city, Luxembour ...
; it extends from
Basel Basel ( ; ), also known as Basle ( ), ; ; ; . is a city in northwestern Switzerland on the river Rhine (at the transition from the High Rhine, High to the Upper Rhine). Basel is Switzerland's List of cities in Switzerland, third-most-populo ...
on the east side of the
Rhine The Rhine ( ) is one of the List of rivers of Europe, major rivers in Europe. The river begins in the Swiss canton of GraubĂŒnden in the southeastern Swiss Alps. It forms part of the Swiss-Liechtenstein border, then part of the Austria–Swit ...
into
Hanover Hanover ( ; ; ) is the capital and largest city of the States of Germany, German state of Lower Saxony. Its population of 535,932 (2021) makes it the List of cities in Germany by population, 13th-largest city in Germany as well as the fourth-l ...
, and through
England England is a Countries of the United Kingdom, country that is part of the United Kingdom. It is located on the island of Great Britain, of which it covers about 62%, and List of islands of England, more than 100 smaller adjacent islands. It ...
into
Scotland Scotland is a Countries of the United Kingdom, country that is part of the United Kingdom. It contains nearly one-third of the United Kingdom's land area, consisting of the northern part of the island of Great Britain and more than 790 adjac ...
and north-east
Ireland Ireland (, ; ; Ulster Scots dialect, Ulster-Scots: ) is an island in the North Atlantic Ocean, in Northwestern Europe. Geopolitically, the island is divided between the Republic of Ireland (officially Names of the Irish state, named Irelan ...
; it appears flanking the central plateau of
France France, officially the French Republic, is a country located primarily in Western Europe. Overseas France, Its overseas regions and territories include French Guiana in South America, Saint Pierre and Miquelon in the Atlantic Ocean#North Atlan ...
and in the
Pyrenees The Pyrenees are a mountain range straddling the border of France and Spain. They extend nearly from their union with the Cantabrian Mountains to Cap de Creus on the Mediterranean coast, reaching a maximum elevation of at the peak of Aneto. ...
and
Sardinia Sardinia ( ; ; ) is the Mediterranean islands#By area, second-largest island in the Mediterranean Sea, after Sicily, and one of the Regions of Italy, twenty regions of Italy. It is located west of the Italian Peninsula, north of Tunisia an ...
. The Keuper sequence is linked by name to the
Keuper Uplands The Keuper Uplands ( or, more rarely ''Keuperwaldberge'') are part of the South German Scarplands and cover an area of about 3,200 square kilometres. The following regions belong to the Keuper Uplands (from southwest to northeast): the Kleiner Heu ...
area of southern Germany. In south
Sweden Sweden, formally the Kingdom of Sweden, is a Nordic countries, Nordic country located on the Scandinavian Peninsula in Northern Europe. It borders Norway to the west and north, and Finland to the east. At , Sweden is the largest Nordic count ...
, the lower portion contains
coal Coal is a combustible black or brownish-black sedimentary rock, formed as rock strata called coal seams. Coal is mostly carbon with variable amounts of other Chemical element, elements, chiefly hydrogen, sulfur, oxygen, and nitrogen. Coal i ...
-bearing strata, as in the
Himalayas The Himalayas, or Himalaya ( ), is a mountain range in Asia, separating the plains of the Indian subcontinent from the Tibetan Plateau. The range has some of the Earth's highest peaks, including the highest, Mount Everest. More than list of h ...
,
Japan Japan is an island country in East Asia. Located in the Pacific Ocean off the northeast coast of the Asia, Asian mainland, it is bordered on the west by the Sea of Japan and extends from the Sea of Okhotsk in the north to the East China Sea ...
,
Tibet Tibet (; ''Böd''; ), or Greater Tibet, is a region in the western part of East Asia, covering much of the Tibetan Plateau and spanning about . It is the homeland of the Tibetan people. Also resident on the plateau are other ethnic groups s ...
,
Burma Myanmar, officially the Republic of the Union of Myanmar; and also referred to as Burma (the official English name until 1989), is a country in northwest Southeast Asia. It is the largest country by area in Mainland Southeast Asia and ha ...
, eastern
Siberia Siberia ( ; , ) is an extensive geographical region comprising all of North Asia, from the Ural Mountains in the west to the Pacific Ocean in the east. It has formed a part of the sovereign territory of Russia and its predecessor states ...
and in
Spitsbergen Spitsbergen (; formerly known as West Spitsbergen; Norwegian language, Norwegian: ''Vest Spitsbergen'' or ''Vestspitsbergen'' , also sometimes spelled Spitzbergen) is the largest and the only permanently populated island of the Svalbard archipel ...
. The upper portion of the
Karoo Supergroup The Karoo Supergroup is the most widespread stratigraphic unit in Africa south of the Kalahari Desert. The supergroup consists of a sequence of units, mostly of nonmarine origin, deposited between the Late Carboniferous and Early Jurassic, a per ...
of
South Africa South Africa, officially the Republic of South Africa (RSA), is the Southern Africa, southernmost country in Africa. Its Provinces of South Africa, nine provinces are bounded to the south by of coastline that stretches along the Atlantic O ...
and part of the Otapiri stage of
New Zealand New Zealand () is an island country in the southwestern Pacific Ocean. It consists of two main landmasses—the North Island () and the South Island ()—and List of islands of New Zealand, over 600 smaller islands. It is the List of isla ...
are probably of Rhaetian age.


Germany

In Germany and adjacent parts of western and central Europe, the Keuper unit is divided into three groups: * Upper Keuper :The upper part of this division is often a grey dolomite known as the Grenz dolomite; the impure
coal Coal is a combustible black or brownish-black sedimentary rock, formed as rock strata called coal seams. Coal is mostly carbon with variable amounts of other Chemical element, elements, chiefly hydrogen, sulfur, oxygen, and nitrogen. Coal i ...
beds Lettenkohle are aggregated towards the base. The upper Keuper, Rhaetic or Avicula contorta zone in Germany is mainly sandy with dark grey
shale Shale is a fine-grained, clastic sedimentary rock formed from mud that is a mix of flakes of Clay mineral, clay minerals (hydrous aluminium phyllosilicates, e.g., Kaolinite, kaolin, aluminium, Al2Silicon, Si2Oxygen, O5(hydroxide, OH)4) and tiny f ...
s and
marl Marl is an earthy material rich in carbonate minerals, Clay minerals, clays, and silt. When Lithification, hardened into rock, this becomes marlstone. It is formed in marine or freshwater environments, often through the activities of algae. M ...
s; it is seldom more than . The sandstones are used for building purposes at
Bayreuth Bayreuth ( or ; High Franconian German, Upper Franconian: Bareid, ) is a Town#Germany, town in northern Bavaria, Germany, on the Red Main river in a valley between the Franconian Jura and the Fichtel Mountains. The town's roots date back to 11 ...
, Culmbach and
Bamberg Bamberg (, , ; East Franconian German, East Franconian: ''BambÀrch'') is a town in Upper Franconia district in Bavaria, Germany, on the river Regnitz close to its confluence with the river Main (river), Main. Bamberg had 79,000 inhabitants in ...
. In Swabia and the Wesergebirge are several
bone A bone is a rigid organ that constitutes part of the skeleton in most vertebrate animals. Bones protect the various other organs of the body, produce red and white blood cells, store minerals, provide structure and support for the body, ...
-beds, thicker than those in the middle Keuper, which contain a rich assemblage of fossil remains of fish, reptiles and the mammalian teeth of '' Microlestes antiquus'' and ''Triglyptzas Fraasi''. The name "Rhaetic" is derived from the Rhaetic Alps where the beds are well developed; they occur also in central France, the Pyrenees and England. In South Tirol and the Judicarian Mountains, the Rhaetic is represented by the Kossenei beds. In the Alpine region, the presence of coral beds gives rise to the so-called Lithodendron Kalk. *'' Hauptkeuper'' or ''Gipskeuper'', the middle : The middle division is thicker than either of the others (at Göttingen, ); it consists of a marly series below, grey, red and green marls, with
gypsum Gypsum is a soft sulfate mineral composed of calcium sulfate Hydrate, dihydrate, with the chemical formula . It is widely mined and is used as a fertilizer and as the main constituent in many forms of plaster, drywall and blackboard or sidewalk ...
and dolomite—this is the gypskeuper in its restricted sense. The higher part of the series is
sand Sand is a granular material composed of finely divided mineral particles. Sand has various compositions but is usually defined by its grain size. Sand grains are smaller than gravel and coarser than silt. Sand can also refer to a textural ...
y, hence called the Steinmergel; it is comparatively free from gypsum. To this division belong the Myophoria beds (
M. Raibliana M, or m, is the thirteenth letter of the English alphabet. M may also refer to: Companies and products * M (fragrance), a 2008 fragrance by Mariah Carey * M-series rangefinder, a series of Leica camera models * M (virtual assistant), a form ...
) with
galena Galena, also called lead glance, is the natural mineral form of lead(II) sulfide (PbS). It is the most important ore of lead and an important source of silver. Galena is one of the most abundant and widely distributed sulfide minerals. It crysta ...
in places; the Estheria beds (
E. laxilesta E is the fifth letter of the Latin alphabet. E or e may also refer to: Computing and computation * E (1970s text editor), a text editor developed at the Stanford AI Lab in the 1970s * E (complexity), a set of decision problems solvable by a ...
); the Schelfsandstein, used as a building-stone; the Lehrherg and Berg-gyps beds;
Semionotus ''Semionotus'' (from , 'mark' and , 'back') is an extinct genus of ray-finned fish found throughout Northern Pangaea (North America and Europe) during the late Triassic, becoming extinct in the Early Jurassic The Jurassic ( ) is a Geological ...
beds ( S. Bergen) with building-stone of
Coburg Coburg ( , ) is a Town#Germany, town located on the Itz (river), Itz river in the Upper Franconia region of Bavaria, Germany. Long part of one of the Thuringian states of the Ernestine duchies, Wettin line, it joined Bavaria by popular vote only ...
; and the Burgand Stubensandstein. *'' Kohlenkeuper, Unterkeuper'' or ''Lettenkohle'', the lower :The lower division consists mainly of grey
clay Clay is a type of fine-grained natural soil material containing clay minerals (hydrous aluminium phyllosilicates, e.g. kaolinite, ). Most pure clay minerals are white or light-coloured, but natural clays show a variety of colours from impuriti ...
s and schieferletten with white, grey and brightly colored
sandstone Sandstone is a Clastic rock#Sedimentary clastic rocks, clastic sedimentary rock composed mainly of grain size, sand-sized (0.0625 to 2 mm) silicate mineral, silicate grains, Cementation (geology), cemented together by another mineral. Sand ...
and dolomitic
limestone Limestone is a type of carbonate rock, carbonate sedimentary rock which is the main source of the material Lime (material), lime. It is composed mostly of the minerals calcite and aragonite, which are different Polymorphism (materials science) ...
. The
salt In common usage, salt is a mineral composed primarily of sodium chloride (NaCl). When used in food, especially in granulated form, it is more formally called table salt. In the form of a natural crystalline mineral, salt is also known as r ...
, which is associated with gypsum, is exploited in south Germany at Dreuze, Pettoncourt, as well as in Vie in the
Lorraine Lorraine, also , ; ; Lorrain: ''Louréne''; Lorraine Franconian: ''Lottringe''; ; ; is a cultural and historical region in Eastern France, now located in the administrative region of Grand Est. Its name stems from the medieval kingdom of ...
region of France. A 4-metre (13 ft) coal is found on this horizon in the
Ore Mountains The Ore Mountains (, or ; ) lie along the Czech–German border, separating the historical regions of Bohemia in the Czech Republic and Saxony in Germany. The highest peaks are the Klínovec in the Czech Republic (German: ''Keilberg'') at ab ...
on the border between Germany and the
Czech Republic The Czech Republic, also known as Czechia, and historically known as Bohemia, is a landlocked country in Central Europe. The country is bordered by Austria to the south, Germany to the west, Poland to the northeast, and Slovakia to the south ...
, and another, 2 metres thick (7 ft), has been mined in
Upper Silesia Upper Silesia ( ; ; ; ; Silesian German: ; ) is the southeastern part of the historical and geographical region of Silesia, located today mostly in Poland, with small parts in the Czech Republic. The area is predominantly known for its heav ...
, now in
Poland Poland, officially the Republic of Poland, is a country in Central Europe. It extends from the Baltic Sea in the north to the Sudetes and Carpathian Mountains in the south, bordered by Lithuania and Russia to the northeast, Belarus and Ukrai ...
.


Great Britain

In Great Britain the 'Keuper' is no longer a formally recognised geological division. The one-time Keuper Marls are now redesignated as the
Mercia Mudstone Group The Mercia Mudstone Group is an early Triassic lithostratigraphy, lithostratigraphic group (stratigraphy), group (a sequence of rock strata) which is widespread in Britain, especially in the English Midlands—the name is derived from the ancient ...
. The underlying Keuper Sandstone is now the Helsby Sandstone Formation at the top of the
Sherwood Sandstone Group The Sherwood Sandstone Group is a Triassic lithostratigraphic group (a sequence of rock strata) which is widespread in Britain, especially in the English Midlands. The name is derived from Sherwood Forest in Nottinghamshire which is underlain by ...
. Traditionally it contained the following subdivisions: * Rhaetic or Penarth :Grey, red and green marls, black shales, and so-called white (). Upper Keuper marl, red and grey marls and shales with rock salt, . As in Germany, there are one or more bone beds in the English Rhaetic with a similar assemblage of fossils. * Lower Keuper :Sandstone, marls and thin sandstones at the top, red and white sandstones (including the so-called waterstones) below, with
breccia Breccia ( , ; ) is a rock composed of large angular broken fragments of minerals or Rock (geology), rocks cementation (geology), cemented together by a fine-grained matrix (geology), matrix. The word has its origins in the Italian language ...
s and conglomerates at the base, . *Basal conglomerate :A shore or scree breccia derived from local materials; it is well developed in the Mendip district. The rocksalt beds vary from 25 cm (1 inch) to in thickness; they are extensively worked (mined and pumped) in
Cheshire Cheshire ( ) is a Ceremonial counties of England, ceremonial county in North West England. It is bordered by Merseyside to the north-west, Greater Manchester to the north-east, Derbyshire to the east, Staffordshire to the south-east, and Shrop ...
,
Middlesbrough Middlesbrough ( ), colloquially known as Boro, is a port town in the Borough of Middlesbrough, North Yorkshire, England. Lying to the south of the River Tees, Middlesbrough forms part of the Teesside Built up area, built-up area and the Tees Va ...
and
County Antrim County Antrim (named after the town of Antrim, County Antrim, Antrim, ) is one of the six counties of Northern Ireland, located within the historic Provinces of Ireland, province of Ulster. Adjoined to the north-east shore of Lough Neagh, the c ...
. The Keuper covers a large area in the
Midlands The Midlands is the central region of England, to the south of Northern England, to the north of southern England, to the east of Wales, and to the west of the North Sea. The Midlands comprises the ceremonial counties of Derbyshire, Herefor ...
and around the flanks of the Pennine range; it reaches southward to the east
Devon Devon ( ; historically also known as Devonshire , ) is a ceremonial county in South West England. It is bordered by the Bristol Channel to the north, Somerset and Dorset to the east, the English Channel to the south, and Cornwall to the west ...
coast, northeastward into
Yorkshire Yorkshire ( ) is an area of Northern England which was History of Yorkshire, historically a county. Despite no longer being used for administration, Yorkshire retains a strong regional identity. The county was named after its county town, the ...
and northwestward into Northern Ireland and southernmost Scotland.


Fossils

The Keuper is not rich in
fossil A fossil (from Classical Latin , ) is any preserved remains, impression, or trace of any once-living thing from a past geological age. Examples include bones, shells, exoskeletons, stone imprints of animals or microbes, objects preserve ...
s; the principal plants are
cypress Cypress is a common name for various coniferous trees or shrubs from the ''Cupressus'' genus of the '' Cupressaceae'' family, typically found in temperate climates and subtropical regions of Asia, Europe, and North America. The word ''cypress'' ...
like
conifer Conifers () are a group of conifer cone, cone-bearing Spermatophyte, seed plants, a subset of gymnosperms. Scientifically, they make up the phylum, division Pinophyta (), also known as Coniferophyta () or Coniferae. The division contains a sin ...
s (''
Walchia ''Walchia'' is a primitive fossil conifer found in upper Pennsylvanian (geology), Pennsylvanian (Carboniferous) and lower Permian (about 310-290 Mya (unit), Mya) rocks of Europe and North America. A forest of In situ, in-situ Walchia tree-stumps ...
'', '' Voltzia'') and a few
calamites ''Calamites'' is a genus of Extinction, extinct arborescent (tree-like) horsetails to which the modern horsetails (genus ''Equisetum'') are closely related. Unlike their Herbaceous plant, herbaceous modern cousins, these plants were medium-size ...
with such forms as ''Equisetum arenaceum'' and ''Pterophyllum jaegeri''. ''Avicula contorta'', ''Protocardium rhaeticum'', ''
Terebratula ''Terebratula'' is a modern genus of brachiopods with a fossil record dating back to the Late Devonian. These brachiopods are stationary epifaunal suspension feeders and have a worldwide distribution. Description ''Terebratula'' species have bi ...
gregaria'', ''Myophoria costata'', ''M. goldfassi'', '' Lingula tenuessima'', and ''Anoplophoria lettica'' may be mentioned among the invertebrates. The fish include ''
Ceratodus ''Ceratodus'' (from , 'horn' and 'tooth') is an extinct genus of freshwater lungfish that was found worldwide during the Mesozoic Era. It has been described as a "catch all", and a " form genus" used to refer to the remains (typically tooth ...
'', ''
Hybodus ''Hybodus'' (from , 'crooked' and 'tooth') is an extinct genus of Hybodontiformes, hybodont. Species closely related to the type species ''Hybodus reticulatus'' lived during the Early Jurassic epoch. Numerous species have been assigned to ''Hyb ...
'' and ''
Lepidotus ''Lepidotes'' (from , 'covered with scales') (previously known as ''Lepidotus'') is an extinct genus of Mesozoic Actinopterygii, ray-finned fish. It has long been considered a wastebasket taxon, characterised by "general features, such as thick ...
''.
Labyrinthodont "Labyrinthodontia" (Greek, 'maze-toothed') is an informal grouping of extinct predatory amphibians which were major components of ecosystems in the late Paleozoic and early Mesozoic eras (about 390 to 150 million years ago). Traditionally conside ...
s represented by the footprints of ''Cheirotherium'' and the bones of ''
Mastodonsaurus ''Mastodonsaurus'' (meaning "teat tooth lizard") is an extinct genus of temnospondyl amphibian from the Middle Triassic of Europe. It belongs to a Triassic group of temnospondyls called Capitosauria, characterized by their large body size, large ...
'' (originally called ''Labyrinthodon'') and '' Capitosaurus''. Among the
reptile Reptiles, as commonly defined, are a group of tetrapods with an ectothermic metabolism and Amniotic egg, amniotic development. Living traditional reptiles comprise four Order (biology), orders: Testudines, Crocodilia, Squamata, and Rhynchocepha ...
s are ''
Hyperodapedon ''Hyperodapedon'' (from , 'above' and , 'pavement') is an extinct genus of rhynchosaur reptiles which lived during Triassic, Late Triassic period. Like other rhynchosaurs, it was an heavily built Archosauromorpha, archosauromorph, distantly rel ...
'', ''
Palaeosaurus ''Palaeosaurus'' (or ''Paleosaurus'') is a genus of indeterminate archosaur known from two teeth found in the Bromsgrove Sandstone Formation and also either the Magnesian Conglomerate or the Avon Fissure Fill of Clifton, Bristol, England (or ...
'', ''
Zanclodon ''Zanclodon'' ("scythe tooth") is an extinct genus of archosauriform from the Erfurt Formation in southern Germany. It was once a wastebasket taxon until a taxonomic revision by Schoch (2011) left only the paratype (SMNS 6045) within ''Zanclod ...
'', ''
Nothosaurus ''Nothosaurus'' ('false lizard', from the Ancient Greek and ) is an extinct genus of sauropterygian reptile from the Triassic period, approximately 245–228 million years ago, with fossils being distributed throughout the former Tethys Ocean, ...
'', ''
Henodus ''Henodus'' (from , 'one' and , 'tooth') is an extinct placodont of the Late Triassic period during the early Carnian age. Fossils of ''Henodus chelyops'' were found in the Estherienschichten Member of the Grabfeld Formation, near TĂŒbingen, ...
'' and ''
Belodon ''Belodon'' (meaning "arrow tooth") is a genus of phytosaur, a crocodile-like reptile that lived during the Triassic. Its fossils have been found in Europe, North American, and India, but their remains have not been found in the southern contine ...
''. The first fossil
mammal A mammal () is a vertebrate animal of the Class (biology), class Mammalia (). Mammals are characterised by the presence of milk-producing mammary glands for feeding their young, a broad neocortex region of the brain, fur or hair, and three ...
s also make their appearance at this time and the early
beetle Beetles are insects that form the Taxonomic rank, order Coleoptera (), in the superorder Holometabola. Their front pair of wings are hardened into wing-cases, elytra, distinguishing them from most other insects. The Coleoptera, with about 40 ...
''
Triamyxa ''Triamyxa'' is an extinct genus of myxophagan beetle in the monotypic family Triamyxidae from the Carnian stage of Late Triassic, approximately 230 million years ago. It was found in the Keuper Claystone of Poland. The type species is ''Tri ...
'' is also known from the Keuper.


Notes


References

* * {{Authority control Triassic System of Europe Lithostratigraphy Lithostratigraphy of Germany Stratigraphy of Europe