Juglandoideae is a subfamily of the walnut family

Juglandaceae The Juglandaceae are a plant family known as the walnut family. They are trees, or sometimes shrubs, in the order Fagales. Members of this family are native to the Americas, Eurasia, and Southeast Asia. The nine or ten genera in the family have ...

. This

clade In biology, a clade (), also known as a Monophyly, monophyletic group or natural group, is a group of organisms that is composed of a common ancestor and all of its descendants. Clades are the fundamental unit of cladistics, a modern approach t ...

was first described by Koidzumi in 1937 by the name "Drupoideae," based on the

drupe In botany, a drupe (or stone fruit) is a type of fruit in which an outer fleshy part (exocarp, or skin, and mesocarp, or flesh) surrounds a single shell (the ''pip'' (UK), ''pit'' (US), ''stone'', or ''pyrena'') of hardened endocarp with a seed ...

-like fruits of ''

Juglans Walnut trees are any species of tree in the plant genus ''Juglans'', the type genus of the family (biology), family Juglandaceae, the seeds of which are referred to as walnuts. All species are deciduous trees, tall, with pinnate leaves , with ...

'' and ''

Carya Hickory is a common name for trees composing the genus ''Carya'', which includes 19 species accepted by ''Plants of the World Online''. Seven species are native to southeast Asia in China, Indochina, and northeastern India (Assam), and twelve ...

''. This name was rejected because it was not based on the name of the type genus. Leroy (1955) and Melchior (1964) both published descriptions of the clade using the name "Jugandoideae," but both were deemed invalid due to technicalities. The first valid publication of the name was by Manning (1978). * Subfamily Juglandoideae ** Tribe Caryeae *** ''

'' – hickory and pecan *** '' Annamocarya'' *** '' Platycarya'' ** Tribe

Juglandeae Juglandeae is a Tribe (biology), tribe of the Juglandoideae subfamily (biology), subfamily, in the Juglandaceae family (biology), family. Walnut tree species comprise the ''Juglans'' genus (biology), genus, which belong to the Juglandeae tribe. ...

*** '' Cyclocarya'' – wheel wingnut *** ''

'' – walnut *** ''

Pterocarya ''Pterocarya'', often called wingnuts in English, are trees in the walnut family Juglandaceae. They are native to Asia. The botanic name is from Ancient Greek (''pteron'') "wing" + (''karyon'') "Nut (fruit), nut". Description ''Pterocarya'' a ...

'' – wingnut Manos and Stone (2001) proposed the following reorganization to reflect a more probable

phylogenetic In biology, phylogenetics () is the study of the evolutionary history of life using observable characteristics of organisms (or genes), which is known as phylogenetic inference. It infers the relationship among organisms based on empirical dat ...

relationship that shows that ''Platycarya'' is sister to the rest of the subfamily, while Manchester (1987) addressed the fossil record of the subfamily: Subfamily Juglandoideae * Tribe Platycaryeae ** †'' Hooleya'' ** †'' Palaeoplatycarya'' ** '' Platycarya'' ** †'' Platycarypollenites'' ** †'' Pterocaryopsis'' ** †'' Vinea'' * Tribe

** Subtribe Caryinae *** '' Annamocarya'' *** ''

'' – hickory and pecan *** †'' Caryapollenites'' *** †'' Eucaryoxylon'' *** †'' Juglandicarya'' ** Subtribe Juglandinae *** '' Cyclocarya'' – wheel wingnut *** ''

'' – walnut *** ''

'' – wingnut *** †'' Pterocaryoxylon'' * Tribe ''

Incertae sedis or is a term used for a taxonomy (biology), taxonomic group where its broader relationships are unknown or undefined. Alternatively, such groups are frequently referred to as "enigmatic taxa". In the system of open nomenclature, uncertainty ...

'' ** †'' Boreocarya'' ** †'' Cruciptera'' ** †'' Globocarya'' ** †'' Polyptera''

References

Rosid subfamilies {{Fagales-stub