Hugh Falconer
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Hugh Falconer MD FRS (29 February 1808 – 31 January 1865) was a Scottish
geologist A geologist is a scientist who studies the structure, composition, and History of Earth, history of Earth. Geologists incorporate techniques from physics, chemistry, biology, mathematics, and geography to perform research in the Field research, ...
,
botanist Botany, also called plant science, is the branch of natural science and biology studying plants, especially Plant anatomy, their anatomy, Plant taxonomy, taxonomy, and Plant ecology, ecology. A botanist or plant scientist is a scientist who s ...
,
palaeontologist Paleontology, also spelled as palaeontology or palæontology, is the scientific study of the life of the past, mainly but not exclusively through the study of fossils. Paleontologists use fossils as a means to classify organisms, measure geolo ...
, and paleoanthropologist. He studied the
flora Flora (: floras or florae) is all the plant life present in a particular region or time, generally the naturally occurring (indigenous (ecology), indigenous) native plant, native plants. The corresponding term for animals is ''fauna'', and for f ...
,
fauna Fauna (: faunae or faunas) is all of the animal life present in a particular region or time. The corresponding terms for plants and fungi are ''flora'' and '' funga'', respectively. Flora, fauna, funga and other forms of life are collectively ...
, and
geology Geology (). is a branch of natural science concerned with the Earth and other astronomical objects, the rocks of which they are composed, and the processes by which they change over time. Modern geology significantly overlaps all other Earth ...
of
India India, officially the Republic of India, is a country in South Asia. It is the List of countries and dependencies by area, seventh-largest country by area; the List of countries by population (United Nations), most populous country since ...
,
Assam Assam (, , ) is a state in Northeast India, northeastern India, south of the eastern Himalayas along the Brahmaputra Valley, Brahmaputra and Barak River valleys. Assam covers an area of . It is the second largest state in Northeast India, nor ...
,
Burma Myanmar, officially the Republic of the Union of Myanmar; and also referred to as Burma (the official English name until 1989), is a country in northwest Southeast Asia. It is the largest country by area in Mainland Southeast Asia and ha ...
, and most of the
Mediterranean islands The Mediterranean Sea basin is supposed to host more than 10,000 islands, with 2,217 islands larger than 0.01 km2. The two main island countries in the region are Malta and Cyprus, while other countries with islands in the Mediterranean Sea in ...
and was the first to suggest the modern evolutionary theory of
punctuated equilibrium In evolutionary biology, punctuated equilibrium (also called punctuated equilibria) is a Scientific theory, theory that proposes that once a species appears in the fossil record, the population will become stable, showing little evolution, evol ...
. He studied the Siwalik fossil beds, and may also have been the first person to discover a
fossil A fossil (from Classical Latin , ) is any preserved remains, impression, or trace of any once-living thing from a past geological age. Examples include bones, shells, exoskeletons, stone imprints of animals or microbes, objects preserve ...
ape Apes (collectively Hominoidea ) are a superfamily of Old World simians native to sub-Saharan Africa and Southeast Asia (though they were more widespread in Africa, most of Asia, and Europe in prehistory, and counting humans are found global ...
.


Early life

Falconer was the youngest son of David Falconer of
Forres Forres (; ) is a town and former royal burgh in the north of Scotland on the County of Moray, Moray coast, approximately northeast of Inverness and west of Elgin, Moray, Elgin. Forres has been a winner of the Scotland in Bloom award on several ...
, Elginshire. In 1826 Hugh Falconer graduated at the
University of Aberdeen The University of Aberdeen (abbreviated ''Aberd.'' in List of post-nominal letters (United Kingdom), post-nominals; ) is a public university, public research university in Aberdeen, Scotland. It was founded in 1495 when William Elphinstone, Bis ...
, where he studied
natural history Natural history is a domain of inquiry involving organisms, including animals, fungi, and plants, in their natural environment, leaning more towards observational than experimental methods of study. A person who studies natural history is cal ...
. Afterward, he studied medicine in the
University of Edinburgh The University of Edinburgh (, ; abbreviated as ''Edin.'' in Post-nominal letters, post-nominals) is a Public university, public research university based in Edinburgh, Scotland. Founded by the City of Edinburgh Council, town council under th ...
, taking the degree of MD in 1829.Arnold, David (2006) ''The Tropics and the Traveling Gaze: India, Landscape, and Science, 1800–1856'' University of Washington Press, Seattle
pp. 156–157
During this period he zealously attended the botanical classes of Prof. R. Graham (1786–1845), and those on geology by Prof.
Robert Jameson image:Robert Jameson.jpg, Robert Jameson Robert Jameson Fellow of the Royal Society, FRS FRSE (11 July 1774 – 19 April 1854) was a Scottish natural history, naturalist and mineralogist. As Regius Professor of Natural History at the Univers ...
, the teacher of
Charles Darwin Charles Robert Darwin ( ; 12 February 1809 – 19 April 1882) was an English Natural history#Before 1900, naturalist, geologist, and biologist, widely known for his contributions to evolutionary biology. His proposition that all speci ...
. Falconer became an assistant-surgeon on the
Bengal Bengal ( ) is a Historical geography, historical geographical, ethnolinguistic and cultural term referring to a region in the Eastern South Asia, eastern part of the Indian subcontinent at the apex of the Bay of Bengal. The region of Benga ...
establishment of the
British East India Company The East India Company (EIC) was an English, and later British, joint-stock company that was founded in 1600 and dissolved in 1874. It was formed to Indian Ocean trade, trade in the Indian Ocean region, initially with the East Indies (South A ...
in 1830. Upon his arrival in Bengal he made an examination of the
fossil A fossil (from Classical Latin , ) is any preserved remains, impression, or trace of any once-living thing from a past geological age. Examples include bones, shells, exoskeletons, stone imprints of animals or microbes, objects preserve ...
bones from Ava, upper Burma in the possession of the
Asiatic Society of Bengal The Asiatic Society is an organisation founded during the Company rule in India to enhance and further the cause of " Oriental research" (in this case, research into India and the surrounding regions). It was founded by the philologist Will ...
. His description of the fossils, published soon afterward, gave him a recognized position among the scientists of
India India, officially the Republic of India, is a country in South Asia. It is the List of countries and dependencies by area, seventh-largest country by area; the List of countries by population (United Nations), most populous country since ...
. Early in 1831 he was posted to the army station at
Meerut Meerut (, ISO 15919, ISO: ''Mēraṭh'') is a city in the western region of the States and union territories of India, Indian state of Uttar Pradesh. Located in the Meerut district, it is northeast of the national capital, New Delhi, and is ...
, India, then in the North Western Provinces, and now in the state of
Uttar Pradesh Uttar Pradesh ( ; UP) is a States and union territories of India, state in North India, northern India. With over 241 million inhabitants, it is the List of states and union territories of India by population, most populated state in In ...
.


Siwalik Hills

In 1832, Falconer became Superintendent of the Saharanpur botanical garden, India, succeeding John Royle. Falconer remained at Saharanpur until 1842, during which time he became widely known for his study of fossil
mammals A mammal () is a vertebrate animal of the class Mammalia (). Mammals are characterised by the presence of milk-producing mammary glands for feeding their young, a broad neocortex region of the brain, fur or hair, and three middle e ...
in the Siwalik Hills. Falconer and his associates may have made the first discovery of a fossil ape, in the 1830s in the
Neogene The Neogene ( ,) is a geologic period and system that spans 20.45 million years from the end of the Paleogene Period million years ago ( Mya) to the beginning of the present Quaternary Period million years ago. It is the second period of th ...
deposits in the Siwalik Hills. In the
Tertiary Tertiary (from Latin, meaning 'third' or 'of the third degree/order..') may refer to: * Tertiary period, an obsolete geologic period spanning from 66 to 2.6 million years ago * Tertiary (chemistry), a term describing bonding patterns in organic ch ...
strata In geology and related fields, a stratum (: strata) is a layer of Rock (geology), rock or sediment characterized by certain Lithology, lithologic properties or attributes that distinguish it from adjacent layers from which it is separated by v ...
of the Siwalik Hills in 1831 Falconer discovered bones of
crocodile Crocodiles (family (biology), family Crocodylidae) or true crocodiles are large, semiaquatic reptiles that live throughout the tropics in Africa, Asia, the Americas and Australia. The term "crocodile" is sometimes used more loosely to include ...
s,
tortoise Tortoises ( ) are reptiles of the family Testudinidae of the order Testudines (Latin for "tortoise"). Like other turtles, tortoises have a shell to protect from predation and other threats. The shell in tortoises is generally hard, and like o ...
s and other animals. With others, he later brought to light a sub-tropical
fossil A fossil (from Classical Latin , ) is any preserved remains, impression, or trace of any once-living thing from a past geological age. Examples include bones, shells, exoskeletons, stone imprints of animals or microbes, objects preserve ...
fauna Fauna (: faunae or faunas) is all of the animal life present in a particular region or time. The corresponding terms for plants and fungi are ''flora'' and '' funga'', respectively. Flora, fauna, funga and other forms of life are collectively ...
of unexampled extent and richness, including remains of ''
Mastodon A mastodon, from Ancient Greek μαστός (''mastós''), meaning "breast", and ὀδούς (''odoús'') "tooth", is a member of the genus ''Mammut'' (German for 'mammoth'), which was endemic to North America and lived from the late Miocene to ...
'', the colossal
ruminant Ruminants are herbivorous grazing or browsing artiodactyls belonging to the suborder Ruminantia that are able to acquire nutrients from plant-based food by fermenting it in a specialized stomach prior to digestion, principally through microb ...
''
Sivatherium ''Sivatherium'' ("Shiva's beast", from Shiva and ''therium'', Latinized form of Ancient Greek θηρίον -'' thēríon'') is an extinct genus of giraffid that ranged throughout Africa and Eurasia. The species ''Sivatherium giganteum'' is, by ...
'', and the enormous extinct
tortoise Tortoises ( ) are reptiles of the family Testudinidae of the order Testudines (Latin for "tortoise"). Like other turtles, tortoises have a shell to protect from predation and other threats. The shell in tortoises is generally hard, and like o ...
'' Colossochelys atlas''. Falconer also published a geological description of the Siwálik Hills in 1834. For these valuable discoveries he and Proby Cautley (1802–1871) together received the Wollaston Medal from the
Geological Society of London The Geological Society of London, known commonly as the Geological Society, is a learned society based in the United Kingdom. It is the oldest national geological society in the world and the largest in Europe, with more than 12,000 Fellows. Fe ...
, its highest award, in 1837. In 1834 Falconer was asked by a Commission of Bengal to investigate the commercial feasibility of growing tea in India. On his recommendation tea plants were introduced, and the resultant
black tea Black tea (also literally translated as red tea from various East Asian languages) is a type of tea that is more tea processing, oxidized than oolong, yellow tea, yellow, white tea, white, and green tea, green teas. Black tea is generally st ...
became competitive with Chinese teas. Falconer returned from India in 1842 because of ill health. He brought back 70 large chests of dried plants and 48 cases of fossils, bones and geological specimens. He then travelled throughout Europe making geological observations, and was elected Fellow of the
Royal Society The Royal Society, formally The Royal Society of London for Improving Natural Knowledge, is a learned society and the United Kingdom's national academy of sciences. The society fulfils a number of roles: promoting science and its benefits, re ...
in 1845. Continuing in the service of the British East India Company as a naturalist, he pursued research at the
British Museum The British Museum is a Museum, public museum dedicated to human history, art and culture located in the Bloomsbury area of London. Its permanent collection of eight million works is the largest in the world. It documents the story of human cu ...
and
East India House East India House was the London headquarters of the East India Company, from which much of Company rule in India, British India was governed until the British government took control of the company's possessions in India in 1858. It was locate ...
and prepared casts of the most remarkable fossils for the leading museums of Europe.


Calcutta

In 1847 Falconer became superintendent of the Calcutta Botanical Garden and professor of botany in the
Medical College A medical association or medical college is a trade association that brings together practitioners of a particular geographical area (a country, region, province). In common-law countries, they are often grouped by medical specialties ( cardiolog ...
,
Calcutta Kolkata, also known as Calcutta (List of renamed places in India#West Bengal, its official name until 2001), is the capital and largest city of the Indian States and union territories of India, state of West Bengal. It lies on the eastern ba ...
, near his older brother, Alexander Falconer, a Calcutta merchant. Hugh Falconer served as an advisor to the Indian government and the Agricultural and Horticultural Society of Bengal, the ''de facto'' colonial "Department of Agriculture". He prepared an important report on the
teak Teak (''Tectona grandis'') is a tropical hardwood tree species in the family Lamiaceae. It is a large, deciduous tree that occurs in mixed hardwood forests. ''Tectona grandis'' has small, fragrant white flowers arranged in dense clusters (panic ...
forests of Tenasserim, and this saved them from destruction by reckless felling. Through his recommendation, the cultivation of the
cinchona ''Cinchona'' (pronounced or ) is a genus of flowering plants in the family Rubiaceae containing at least 23 species of trees and shrubs. All are native to the Tropical Andes, tropical Andean forests of western South America. A few species are ...
in the Indian empire was introduced for the medicinal use of its bark in the treatment of
malaria Malaria is a Mosquito-borne disease, mosquito-borne infectious disease that affects vertebrates and ''Anopheles'' mosquitoes. Human malaria causes Signs and symptoms, symptoms that typically include fever, Fatigue (medical), fatigue, vomitin ...
.


Punctuated equilibrium

Falconer was originally a creationist who denied the fact of
evolution Evolution is the change in the heritable Phenotypic trait, characteristics of biological populations over successive generations. It occurs when evolutionary processes such as natural selection and genetic drift act on genetic variation, re ...
.Gould, Stephen Jay. (2007). ''Punctuated Equilibrium''. Harvard University Press. pp. 14–18. In November 1859,
Charles Darwin Charles Robert Darwin ( ; 12 February 1809 – 19 April 1882) was an English Natural history#Before 1900, naturalist, geologist, and biologist, widely known for his contributions to evolutionary biology. His proposition that all speci ...
sent Falconer a copy of his ''
On the Origin of Species ''On the Origin of Species'' (or, more completely, ''On the Origin of Species by Means of Natural Selection, or the Preservation of Favoured Races in the Struggle for Life'')The book's full original title was ''On the Origin of Species by M ...
'' with a letter which stated "I am fully convinced that you will become, year after year, less fixed in your belief in the immutability of species". In June 1861, Falconer expressed respect in a letter to Darwin for receiving the book. By the early 1860s he reassessed his worldview and came to embrace evolution from his studies of the fossil record. In 1863, Falconer authored a monograph ''On the American Fossil elephant of the regions bordering the Gulf of Mexico''. A year before he had privately sent the work to Darwin who was delighted to read it. In the work he observed long periods of evolutionary stasis in fossil mammals with short periods of rapid evolutionary change throughout geological time. This research shows great foresight. Niles Eldredge and
Stephen Jay Gould Stephen Jay Gould ( ; September 10, 1941 – May 20, 2002) was an American Paleontology, paleontologist, Evolutionary biology, evolutionary biologist, and History of science, historian of science. He was one of the most influential and widely re ...
developed the same basic theory a century later, a theory known as
punctuated equilibrium In evolutionary biology, punctuated equilibrium (also called punctuated equilibria) is a Scientific theory, theory that proposes that once a species appears in the fossil record, the population will become stable, showing little evolution, evol ...
. According to Gould, the work "anticipates a primary inference of punctuated equilibrium— that a local pattern of abrupt replacement does not signify macromutational transformation in situ, but an origin of the later species from an ancestral population living elsewhere, followed by migration into the local region."


Later years

Having to leave India again in 1855 because of ill health, he spent the remainder of his life examining and comparing fossil species in England and the Continent to those he found in India, notably the species of mastodon, elephant, and rhinoceros. He also described some new
mammalia A mammal () is a vertebrate animal of the class Mammalia (). Mammals are characterised by the presence of milk-producing mammary glands for feeding their young, a broad neocortex region of the brain, fur or hair, and three middle ear bon ...
from the Purbeck strata of
Wessex The Kingdom of the West Saxons, also known as the Kingdom of Wessex, was an Anglo-Saxon Heptarchy, kingdom in the south of Great Britain, from around 519 until Alfred the Great declared himself as King of the Anglo-Saxons in 886. The Anglo-Sa ...
. Turning to the subject of human origins, he reported on the bone
caves Caves or caverns are natural voids under the Earth's surface. Caves often form by the weathering of rock and often extend deep underground. Exogene caves are smaller openings that extend a relatively short distance underground (such as rock ...
of
Sicily Sicily (Italian language, Italian and ), officially the Sicilian Region (), is an island in the central Mediterranean Sea, south of the Italian Peninsula in continental Europe and is one of the 20 regions of Italy, regions of Italy. With 4. ...
,
Malta Malta, officially the Republic of Malta, is an island country in Southern Europe located in the Mediterranean Sea, between Sicily and North Africa. It consists of an archipelago south of Italy, east of Tunisia, and north of Libya. The two ...
,
Gibraltar Gibraltar ( , ) is a British Overseas Territories, British Overseas Territory and British overseas cities, city located at the southern tip of the Iberian Peninsula, on the Bay of Gibraltar, near the exit of the Mediterranean Sea into the A ...
, Gower, and
Brixham Brixham is a coastal town and civil parish in the borough of Torbay in the county of Devon, in the south-west of England. As of the 2021 census, Brixham had a population of 16,825. It is one of the main three centres of the borough, along with ...
. He discovered the dwarf elephant species '' Palaeoloxodon falconeri'' on Sicily and Malta, and also found the bones of the flightless giant swan ''
Cygnus falconeri ''Cygnus falconeri'' is an extinct species of very large swan known from Middle Pleistocene-aged deposits from Malta and Sicily. Its dimensions are described as exceeding those of the living mute swan by one-third, which would give a bill-to-tai ...
'' in Malta at
Għar Dalam Għar Dalam (; "Cave of Dalam", Dalam being a fifteenth-century family name) is a 144-metre long phreatic tube and cave, located in the outskirts of Birżebbuġa, Malta. The cave contains the bones of animals that lived on Malta during the Plei ...
. Falconer served as vice-president of the
Royal Society The Royal Society, formally The Royal Society of London for Improving Natural Knowledge, is a learned society and the United Kingdom's national academy of sciences. The society fulfils a number of roles: promoting science and its benefits, re ...
1863–1864. Although suffering from exposure and overwork, Falconer returned hastily from Gibraltar to support
Charles Darwin Charles Robert Darwin ( ; 12 February 1809 – 19 April 1882) was an English Natural history#Before 1900, naturalist, geologist, and biologist, widely known for his contributions to evolutionary biology. His proposition that all speci ...
's claim to the
Copley Medal The Copley Medal is the most prestigious award of the Royal Society of the United Kingdom, conferred "for sustained, outstanding achievements in any field of science". The award alternates between the physical sciences or mathematics and the bio ...
in 1864. Falconer succumbed in London, England, on 31 January 1865 from rheumatic disease of the heart and lungs. He is buried in
Kensal Green Cemetery Kensal Green Cemetery is a cemetery in the Kensal Green area of North Kensington in the Royal Borough of Kensington and Chelsea and the London Borough of Hammersmith and Fulham in London, England. Inspired by Père Lachaise Cemetery in P ...
, London. The flower '' Rhododendron falconeri'' was named after Falconer by
Joseph Dalton Hooker Sir Joseph Dalton Hooker (30 June 1817 – 10 December 1911) was a British botanist and explorer in the 19th century. He was a founder of geographical botany and Charles Darwin's closest friend. For 20 years he served as director of the Ro ...
. Falconer's botanical notes, with 450 coloured drawings of Indian plants, were deposited in the library at
Kew Gardens Kew Gardens is a botanical garden, botanic garden in southwest London that houses the "largest and most diverse botany, botanical and mycology, mycological collections in the world". Founded in 1759, from the exotic garden at Kew Park, its li ...
, together with some of the specimens he collected. A marble bust was placed in the rooms of the
Royal Society The Royal Society, formally The Royal Society of London for Improving Natural Knowledge, is a learned society and the United Kingdom's national academy of sciences. The society fulfils a number of roles: promoting science and its benefits, re ...
of London, and another in the
Asiatic Society The Asiatic Society is an organisation founded during the Company rule in India to enhance and further the cause of " Oriental research" (in this case, research into India and the surrounding regions). It was founded by the philologist Will ...
of Bengal, Calcutta. A competitive Falconer scholarship of £100 per year was created for graduates in science or medicine of the University of Edinburgh. The Falconer Museum in
Forres Forres (; ) is a town and former royal burgh in the north of Scotland on the County of Moray, Moray coast, approximately northeast of Inverness and west of Elgin, Moray, Elgin. Forres has been a winner of the Scotland in Bloom award on several ...
, Scotland (), a "Victorian Gem", houses, amongst numerous other collections, the many Falconer finds that had not been sent to other institutions like the library of Kew Gardens or the British Museum, including Palaeolithic finds that enable "visitors of the Falconer Museum olook at three different species of humans at once, ''homo erectus, homo neanderthalensis'' and ''homo sapiens''".


Selected publications

* Hugh Falconer and Proby T. Cautley, ''Fauna Antiqua Sivalensis, being the Fossil Zoology of the Sewalik Hills, in the North of India, Part I, Proboscidea,'' London (1846), with a series of 107 plates by G. H. Ford appearing between 1846 and 1849. * ''On the American Fossil Elephant of the Regions Bordering the Gulf of Mexico, (E. Columbi, Falc.): With General Observations on the Living and Extinct Species'' (1863) * ''Palæontological memoirs and notes of the late Hugh Falconer,'' edited, with a biographical sketch, by Charles Murchison, M.D., 2 vols., London, R. Hardwicke (1868). OCLC: 2847098
vol. II online at archive.org


* Falconer's works were documented in the Royal Society's ''Catalogue of Scientific Papers,'' vol. ii (1968).


Notes


Literature

* Patrick J. Boylan, ''The Falconer papers, Forres,'' Leicester: Leicestershire Museums, Art Galleries and Records Service (1977). * Grace, Lady Prestwich, ''Essays descriptive and biographical,'' Edinburgh and London, William Blackwood (1901). * Charles T. Gaudin, "Modifications apportées par M. Falconer à la faune du Val d'Arno", ''Bulletin des Séances de la Société Vaudoise des Sciences Naturelles'' 6: 130–1 (1859). * Patrick J. Boylan, "The controversy of the Moulin-Quignon jaw: the role of Hugh Falconer," in ''Images of the Earth: Essays in the History of the Environmental Sciences,'' Ludmilla J. Jordanova and Roy S. Porter, eds., Chalfont St. Giles, Bucks., British Society for the History of Science. (1979) * Leonard G. Wilson, "Brixham Cave and Sir Charles Lyell's . . . the Antiquity of man: the roots of Hugh Falconer's attack on Lyell," ''Archives of Natural History'' 23: 79–97 (1996). * * Kenneth A. R. Kennedy and Russell L. Ciochon, "A canine tooth from the Siwaliks: first recorded discovery of a fossil ape?" ''Journal of Human Evolution,'' Vol. 14, No. 3 (July, 1999). (Print) (Online). * Anne O'Connor, "Hugh Falconer, Joseph Prestwich and the Gower caves", ''Studies in Speleology,'' Vol. 14, pp 75 – 79 (2006). * Tim Murray (archaeologist), "Hugh Falconer: Botanist, palaeontologist, controversialist", in ''Life-writing in the History of Archaeology: Critical perspectives'', 2023, Clare Lewis and Gabriel Moshenskapp eds., pp 265-280
Hugh Falconer: Botanist, palaeontologist, controversialist


External links


The Falconer Museum
*
Friends Blog

Prof. Tim Murray on Hugh Falconer and his special role in the history of science
* {{DEFAULTSORT:Falconer, Hugh People from Forres 1808 births 1865 deaths Botanists with author abbreviations Alumni of the University of Aberdeen Alumni of the University of Edinburgh Scottish palaeontologists Scottish geologists 19th-century Scottish botanists Botanists active in India Fellows of the Royal Society Wollaston Medal winners Burials at Kensal Green Cemetery Punctuated equilibrium