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Helmut Mayrhofer is an Austrian lichenologist. He is known for his expertise on the lichen family
Physciaceae The Physciaceae are a family of mostly lichen-forming fungi belonging to the class Lecanoromycetes in the division Ascomycota. A 2016 estimate placed 19 genera and 601 species in the family. Genera This is a list of the genera contained withi ...
and his studies of the lichen
flora Flora (: floras or florae) is all the plant life present in a particular region or time, generally the naturally occurring (indigenous (ecology), indigenous) native plant, native plants. The corresponding term for animals is ''fauna'', and for f ...
of the
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, the
Alps The Alps () ; german: Alpen ; it, Alpi ; rm, Alps ; sl, Alpe . are the highest and most extensive mountain range system that lies entirely in Europe, stretching approximately across seven Alpine countries (from west to east): France, ...
, and other regions.


Biography

Helmut Mayrhofer was born in 1953 in Aich, a village in Austria. He went to secondary school in
Gröbming Gröbming is a municipality in the district of Liezen in Styria, Austria Austria, , bar, Östareich officially the Republic of Austria, is a country in the southern part of Central Europe, lying in the Eastern Alps. It is a federa ...
and
Graz Graz (; sl, Gradec) is the capital city of the Austrian state of Styria and second-largest city in Austria after Vienna. As of 1 January 2021, it had a population of 331,562 (294,236 of whom had principal-residence status). In 2018, the popu ...
, and then to the
University of Graz The University of Graz (german: link=no, Karl-Franzens-Universität Graz, ), located in Graz, Austria, is the largest and oldest university in Styria, as well as the second-largest and second-oldest university in Austria. History The univers ...
. He graduated from there in 1976 with a
master's degree A master's degree (from Latin ) is an academic degree awarded by universities or colleges upon completion of a course of study demonstrating mastery or a high-order overview of a specific field of study or area of professional practice.
, having submitted a thesis titled "Flechten auf ''Rhododendron ferrugineum''" ("Lichens on '' Rhododendron ferrugineum''"). Mayrhofer worked at the as a high school teacher before returning to the University of Graz for work as a scientific assistant, a position he held until 1988. While working as a schoolteacher, Mayrhofer was also working on his doctoral degree, under the supervision of
Josef Poelt Josef Poelt was a botanist, bryologist and lichenologist. He held the chair in Systematic Botany and Plant Geography at the Free University of Berlin (1965 - 1972) and then was head of the Botanical Institute and Botanical Garden of Graz Universi ...
. He defended his thesis, titled "Die saxicolen Arten der Flechtengattungen ''Rinodina'' und ''Rinodinella'' in der Alten Welt" ("The saxicolous species of the lichen genera ''Rinodina'' and ''Rinodinella'' in the Old World) in 1988, receiving his ''
venia docendi Doctor of Science ( la, links=no, Scientiae Doctor), usually abbreviated Sc.D., D.Sc., S.D., or D.S., is an academic research degree awarded in a number of countries throughout the world. In some countries, "Doctor of Science" is the degree used f ...
'' (Doctor of Science) in
plant systematics The history of plant systematics—the biological classification of plants—stretches from the work of ancient Greek to modern evolutionary biologists. As a field of science, plant systematics came into being only slowly, early plant lo ...
. That year he became a university lecturer; eventually he became an
associate professor Associate professor is an academic title with two principal meanings: in the North American system and that of the ''Commonwealth system''. Overview In the '' North American system'', used in the United States and many other countries, it is ...
in 1995, and held this position until he retired in 2018. As of that year, he had supervised 10 PhD students and 38 masters and diploma thesis candidates. Mayrhofer served twice (1999–2010 and 2013–2017) as the head of the Institute of Plant Sciences at the University of Graz. He was a
visiting scholar In academia, a visiting scholar, visiting researcher, visiting fellow, visiting lecturer, or visiting professor is a scholar from an institution who visits a host university to teach, lecture, or perform research on a topic for which the visitor ...
at the
University of Ljubljana The University of Ljubljana ( sl, Univerza v Ljubljani, , la, Universitas Labacensis), often referred to as UL, is the oldest and largest university in Slovenia. It has approximately 39,000 enrolled students. History Beginnings Although certain ...
in Slovenia from 2001 to 2003 and at the
University of Montenegro The University of Montenegro ( cnr, Универзитет Црнe Горe / Univerzitet Crne Gore) is a national public university of Montenegro. Its central administration and majority of constitutive faculties are located in the country's cap ...
in
Podgorica Podgorica (Cyrillic script, Cyrillic: Подгорица, ; Literal translation, lit. 'under the hill') is the Capital city, capital and List of cities and towns in Montenegro, largest city of Montenegro. The city was formerly known as Titograd ...
from 2004 to 2006. He served as a member of the editorial boards of the journals ''
Acta Botanica Croatica Acta or ACTA may refer to: Institutions * Anti-Counterfeiting Trade Agreement, an intellectual property trade agreement * Administrative Council for Terminal Attachments, a standards organization for terminal equipment such as registered jacks * ...
'', ''
Mycobiota Mycobiota (plural noun, no singular) are a group of all the fungi present in a particular geographic region (e.g. "the mycobiota of Ireland") or habitat type (e.g. "the mycobiota of cocoa"). Human mycobiota Mycobiota exist on the surface and in th ...
'', and ''
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'', and has been an editor of the journal ''
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''.


Research

Mayrhofer started research on crustose lichens beginning with his master's thesis. Through comparative light microscopical studies of lichens in '' Rinodina'', he developed a typology of
ascospore An ascus (; ) is the sexual spore-bearing cell produced in ascomycete fungi. Each ascus usually contains eight ascospores (or octad), produced by meiosis followed, in most species, by a mitotic cell division. However, asci in some genera o ...
s that proved to be useful in the identification of species in that genus, and later in the
taxonomy Taxonomy is the practice and science of categorization or classification. A taxonomy (or taxonomical classification) is a scheme of classification, especially a hierarchical classification, in which things are organized into groups or types. ...
of the entire family. As a result of his research, ''Rinodina'' went from being one of the least understood crustose lichen genera in the 1980s to one of the most studied. He later included other characters such as secondary chemistry,
ascus An ascus (; ) is the sexual spore-bearing cell produced in ascomycete fungi. Each ascus usually contains eight ascospores (or octad), produced by meiosis followed, in most species, by a mitotic cell division. However, asci in some genera o ...
shapes, ascospore
ontogeny Ontogeny (also ontogenesis) is the origination and development of an organism (both physical and psychological, e.g., moral development), usually from the time of fertilization of the egg to adult. The term can also be used to refer to the stu ...
, and
molecular phylogenetics Molecular phylogenetics () is the branch of phylogeny that analyzes genetic, hereditary molecular differences, predominantly in DNA sequences, to gain information on an organism's evolutionary relationships. From these analyses, it is possible to ...
to help resolve taxonomic issues in the Physciaceae. Another research interest of Mayrhofer has been documenting lichen
biodiversity Biodiversity or biological diversity is the variety and variability of life on Earth. Biodiversity is a measure of variation at the genetic ('' genetic variability''), species ('' species diversity''), and ecosystem ('' ecosystem diversity' ...
in various locations. Together with his students, and sometimes in collaboration with other colleagues, he has catalogued the lichen
biota Biota may refer to: * Biota (ecology), the plant and animal life of a region * Biota (plant), common name for a coniferous tree, ''Platycladus orientalis'' * Biota, Cinco Villas, a municipality in Aragon, Spain * Biota (band), a band from Color ...
of several countries in the
Balkan Peninsula The Balkans ( ), also known as the Balkan Peninsula, is a geographical area in southeastern Europe with various geographical and historical definitions. The region takes its name from the Balkan Mountains that stretch throughout the whol ...
, the Mediterranean islands of
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and
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, as well as the Asian countries of
Armenia Armenia (), , group=pron officially the Republic of Armenia,, is a landlocked country in the Armenian Highlands of Western Asia.The UNbr>classification of world regions places Armenia in Western Asia; the CIA World Factbook , , and ...
and
Tajikistan Tajikistan (, ; tg, Тоҷикистон, Tojikiston; russian: Таджикистан, Tadzhikistan), officially the Republic of Tajikistan ( tg, Ҷумҳурии Тоҷикистон, Jumhurii Tojikiston), is a landlocked country in Centr ...
. Mayrhofer has explored extensively several mountainous areas in Austria and Slovenia and has compiled an annotated inventory of lichens in the
Alps The Alps () ; german: Alpen ; it, Alpi ; rm, Alps ; sl, Alpe . are the highest and most extensive mountain range system that lies entirely in Europe, stretching approximately across seven Alpine countries (from west to east): France, ...
.


Eponymy

Several lichen species have been named in Mayrhofer's honour, including: ''
Fuscidea mayrhoferi ''Fuscidea'' is a genus of crustose lichens in the family Fuscideaceae. It has about 40 species. The genus was circumscribed in 1972 by lichenologists Volkmar Wirth and Antonín Vězda, with '' Fuscidea aggregatilis'' assigned as the type spec ...
'' ; ''
Lecanora mayrhoferi ''Lecanora'' is a genus of lichen commonly called rim lichens.Field Guide to California Lichens, Stephen Sharnoff, Yale University Press, 2014, Lichens in the genus '' Squamarina'' are also called rim lichens. Members of the genus have roughly c ...
'' ; '' Rinodina mayrhoferi'' (1984); ''
Skyttea mayrhoferi ''Skyttea'' is a genus of lichenicolous (lichen-dwelling) fungi in the family Cordieritidaceae. The genus was circumscribed in 1981 by lichenologists Martha Allen Sherwood, David L. Hawksworth, and Brian J. Coppins, with ''Skyttea nitschkei'' as ...
'' ; ''
Unguiculariopsis helmutii ''Unguiculariopsis'' is a genus of lichenicolous fungi in the family Cordieritidaceae. It has 29 species. Species * '' Unguiculariopsis acerina'' * '' Unguiculariopsis acrocordiae'' * '' Unguiculariopsis adirondacensis'' * ''Unguiculariopsis ...
'' ; '' Xanthoparmelia mayrhoferi'' ; '' Acarospora mayrhoferi'' ; ''
Cratiria mayrhoferi ''Cratiria'' is a genus of lichenized fungi in the family Caliciaceae. The genus has a widespread distribution, especially in tropical regions, and contains about 20 species. The genus was circumscribed by Austrian lichenologist Bernhard Marb ...
'' ; '' Dimelaena mayrhoferiana'' ; '' Lecanora helmutii'' ; ''
Stigmidium mayrhoferi ''Stigmidium'' is a genus of lichenicolous (lichen-eating) fungi in the family Mycosphaerellaceae. The genus was circumscribed by Italian botanist Vittore Benedetto Antonio Trevisan de Saint-Léon in 1860, with ''Stigmidium schaereri'' assig ...
'' ''
Tremella mayrhoferi ''Tremella'' is a genus of fungi in the family Tremellaceae. All ''Tremella'' species are parasites of other fungi and most produce anamorphic yeast states. Basidiocarps (fruit bodies), when produced, are gelatinous and are colloquially cl ...
'' ; and ''
Usnea mayrhoferi ''Usnea'' is a genus of mostly pale grayish-green fruticose lichens that grow like leafless mini-shrubs or tassels anchored on bark or twigs.Field Guide to California Lichens, Stephen Sharnoff, Yale University Press, 2014, The genus is in the f ...
'' .


Selected publications

A compilation of Mayrhofer's scientific publications up until 2018 was given in the Bilovitz et al. tribute article. Some of his major works include: * * * * * *


See also

* :Taxa named by Helmut Mayrhofer


References

{{DEFAULTSORT:Mayrhofer, Helmut 1953 births Living people Austrian lichenologists Austrian taxonomists People from Liezen District University of Graz alumni