''Harpesaurus modiglianii'', also known commonly as Modigliani's nose-horned lizard, is a

species A species () is often defined as the largest group of organisms in which any two individuals of the appropriate sexes or mating types can produce fertile offspring, typically by sexual reproduction. It is the basic unit of Taxonomy (biology), ...

lizard Lizard is the common name used for all Squamata, squamate reptiles other than snakes (and to a lesser extent amphisbaenians), encompassing over 7,000 species, ranging across all continents except Antarctica, as well as most Island#Oceanic isla ...

in the

family Family (from ) is a Social group, group of people related either by consanguinity (by recognized birth) or Affinity (law), affinity (by marriage or other relationship). It forms the basis for social order. Ideally, families offer predictabili ...

Agamidae Agamidae is a family containing 582 species in 64 genera of iguanian lizards indigenous to Africa, Asia, Australia, and a few locations in Southern Europe. Many species are commonly called dragons or dragon lizards. Overview Phylogenetically ...

. The species is

endemic Endemism is the state of a species being found only in a single defined geographic location, such as an island, state, nation, country or other defined zone; organisms that are indigenous to a place are not endemic to it if they are also foun ...

Indonesia Indonesia, officially the Republic of Indonesia, is a country in Southeast Asia and Oceania, between the Indian Ocean, Indian and Pacific Ocean, Pacific oceans. Comprising over List of islands of Indonesia, 17,000 islands, including Sumatra, ...

. Once only known from the

holotype A holotype (Latin: ''holotypus'') is a single physical example (or illustration) of an organism used when the species (or lower-ranked taxon) was formally described. It is either the single such physical example (or illustration) or one of s ...

, a single specimen collected from

North Sumatra North Sumatra () is a Provinces of Indonesia, province of Indonesia located in the northern part of the island of Sumatra. Its capital and largest city is Medan on the east coast of the island. It borders Aceh to the northwest, Riau to the sou ...

in 1891, the species was rediscovered after a living population of the species was sighted in 2018.

Etymology

The

specific name Specific name may refer to: * in Database management systems, a system-assigned name that is unique within a particular database In taxonomy, either of these two meanings, each with its own set of rules: * Specific name (botany), the two-part (bino ...

, ''modiglianii'', is honor of Italian botanist

Elio Modigliani Elio Modigliani (13 June 1860 – 6 August 1932) was an Italian anthropologist, zoologist, explorer, and plant collector. Biography The son of a Florence, Florentine banker, he first made his name in the Italian scientific community at the age o ...

Geographic range

''H. modiglianii'' is indigenous to northern

Sumatra Sumatra () is one of the Sunda Islands of western Indonesia. It is the largest island that is fully within Indonesian territory, as well as the list of islands by area, sixth-largest island in the world at 482,286.55 km2 (182,812 mi. ...

, Indonesia.

Habitat

The preferred natural

habitat In ecology, habitat refers to the array of resources, biotic factors that are present in an area, such as to support the survival and reproduction of a particular species. A species' habitat can be seen as the physical manifestation of its ...

of ''H. modiglianii'' is

forest A forest is an ecosystem characterized by a dense ecological community, community of trees. Hundreds of definitions of forest are used throughout the world, incorporating factors such as tree density, tree height, land use, legal standing, ...

Description

The holotype specimen is a male, and the preservation process had caused it to turn blue. The discovery of live specimens in 2018 revealed that the species is naturally a luminous green colour in life. ''H. modiglianii'' has a sickle-shaped upturned nasal appendage. Not including this appendage, the

has a snout-to-vent length (SVL) of , and a tail length of . Its nasal appendage is long, roughly as long as the head. Vinciguerra (1933).

Reproduction

''H. modiglianii'' is

oviparous Oviparous animals are animals that reproduce by depositing fertilized zygotes outside the body (i.e., by laying or spawning) in metabolically independent incubation organs known as eggs, which nurture the embryo into moving offsprings kno ...

Etymology

Geographic range

Habitat

Description

Reproduction

References

Further reading