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Gunnar Bror Fritiof Degelius (
The birth name is the name of the person given upon their birth. The term may be applied to the surname, the given name or to the entire name. Where births are required to be officially registered, the entire name entered onto a births registe ...
Nilsson until 1932; 27 January 1903 – 22 July 1993) was a Swedish lichenologist. Between the publications of his first and final scientific papers, Degelius had a 70-year-long research career. While he was best known for his expertise on the lichen genus ''
Collema ''Collema'' (jelly lichen) is a genus of lichens in the family Collemataceae. The photobiont is the cyanobacterium genus ''Nostoc''.Dobson, F.S. (2000) Lichens, an illustrated guide to the British and Irish species. 4th edition. Richmond publish ...
'', he also wrote important papers on lichen biology and ecology,
floristic {{Short pages monitor ''Did you mean: "Floristics Floristics is the study of plants of geographical regions. It is a branch of phytogeography, which technically makes it a branch of botany, geography, and a subbranch of biogeography. Harvard Un ...
studies of the
Nordic countries The Nordic countries (also known as the Nordics or ''Norden''; ) are a geographical and cultural region in Northern Europe, as well as the Arctic Ocean, Arctic and Atlantic Ocean, North Atlantic oceans. It includes the sovereign states of Denm ...
and various other areas around the world, and lichen
succession Succession is the act or process of following in order or sequence. Governance and politics *Order of succession, in politics, the ascension to power by one ruler, official, or monarch after the death, resignation, or removal from office of ...
. Degelius described 124 new
taxa In biology, a taxon (back-formation from ''taxonomy''; : taxa) is a group of one or more populations of an organism or organisms seen by taxonomists to form a unit. Although neither is required, a taxon is usually known by a particular name and ...
(mostly species), and published about 130 scientific papers. In 1992 he was one of the first to be awarded the
Acharius Medal __NOTOC__ The Acharius Medal is awarded by the International Association for Lichenology (IAL) for lifetime achievement in lichenology. The organization resolved at its 1990 meeting that it would simultaneously honor professional achievement and ...
for his lifetime contributions to lichenology. Fifteen species and three
genera Genus (; : genera ) is a taxonomic rank above species and below family as used in the biological classification of living and fossil organisms as well as viruses. In binomial nomenclature, the genus name forms the first part of the binomial s ...
have been named in honour of Degelius.


Early life and education

Born in
Uppsala Uppsala ( ; ; archaically spelled ''Upsala'') is the capital of Uppsala County and the List of urban areas in Sweden by population, fourth-largest city in Sweden, after Stockholm, Gothenburg, and Malmö. It had 177,074 inhabitants in 2019. Loc ...
on 27 January 1903, Degelius spent much of his youth in
Mariestad Mariestad () is a Urban areas in Sweden, locality and the seat of Mariestad Municipality, Västra Götaland County, Sweden. It had 16,611 inhabitants in 2019. Until 1997 it was the capital of the former Skaraborg County and an episcopal see in the ...
. He was introduced to
botany Botany, also called plant science, is the branch of natural science and biology studying plants, especially Plant anatomy, their anatomy, Plant taxonomy, taxonomy, and Plant ecology, ecology. A botanist or plant scientist is a scientist who s ...
at an early age by his father, Bror Nilsson, who was a pharmacist. By the age of six, Gunnar had a sizeable collection of
seed plant A seed plant or spermatophyte (; New Latin ''spermat-'' and Greek ' (phytón), plant), also known as a phanerogam (taxon Phanerogamae) or a phaenogam (taxon Phaenogamae), is any plant that produces seeds. It is a category of embryophyte (i.e. la ...
s, and he collected his first
lichen A lichen ( , ) is a hybrid colony (biology), colony of algae or cyanobacteria living symbiotically among hypha, filaments of multiple fungus species, along with yeasts and bacteria embedded in the cortex or "skin", in a mutualism (biology), m ...
– the dog lichen, ''
Peltigera canina ''Peltigera canina'', commonly known as the dog lichen, is a widely distributed species of foliose lichen in the family Peltigeraceae. It was originally described by Carl Linnaeus in his 1753 work ''Species Plantarum''. German botanist Carl Ludw ...
'' – at age 12. After his family moved to
Gothenburg Gothenburg ( ; ) is the List of urban areas in Sweden by population, second-largest city in Sweden, after the capital Stockholm, and the fifth-largest in the Nordic countries. Situated by the Kattegat on the west coast of Sweden, it is the gub ...
, he met the lichenologists Captain Carl Stenholm and
Adolf Hugo Magnusson Adolf Hugo Magnusson (1 March 188514 July 1964) was a Swedish naturalist who specialized in lichenology. He was a school teacher in Gothenburg from 1909 to 1948, but spent his spare time on the study of lichens. Magnusson published many monograph ...
, both of whom encouraged his early interest in lichens. Degelius attended school with Carl H. Lindroth, who later became a notable
entomologist Entomology (from Ancient Greek ἔντομον (''éntomon''), meaning "insect", and -logy from λόγος (''lógos''), meaning "study") is the branch of zoology that focuses on insects. Those who study entomology are known as entomologists. In ...
. They shared an early interest in
natural history Natural history is a domain of inquiry involving organisms, including animals, fungi, and plants, in their natural environment, leaning more towards observational than experimental methods of study. A person who studies natural history is cal ...
, and they would often go on trips together to collect plants, beetles, and lichens in
Mölndal Mölndal () is the seat and administrative centre of Mölndal Municipality and a part of the Gothenburg urban area on the west coast of Sweden. About 40,000 of the municipality's 60,000 inhabitants live in Mölndal proper. Geography Mölndal is l ...
, south of Gothenburg. Degelius
matriculated Matriculation is the formal process of entering a university, or of becoming eligible to enter by fulfilling certain academic requirements such as a matriculation examination. Australia In Australia, the term ''matriculation'' is seldom used now ...
in 1923, the same year he published his first scientific paper (under the name Nilsson) on the lichen ''
Arthonia spadicea ''Diarthonis'' is a fungal genus in the family Graphidaceae. Originally created in 1909 but long unused, the genus was revived in 2020 to accommodate ''D. spadicea'' when molecular studies showed this species fell outside the main '' Artho ...
''. Soon after he began studies at
Uppsala University Uppsala University (UU) () is a public university, public research university in Uppsala, Sweden. Founded in 1477, it is the List of universities in Sweden, oldest university in Sweden and the Nordic countries still in operation. Initially fou ...
. His professor of
systematic botany ''Systematic Botany'' is a peer-reviewed scientific journal covering the study of systematic botany. It is published quarterly by the American Society of Plant Taxonomists. According to the ''Journal Citation Reports'', the journal has a 2010 i ...
became aware of Degelius because of his tendency to frequently miss lectures; after he called him into his office for an explanation, it became evident that Degelius already knew the subject matter quite well, having essentially memorized the textbook used for the course,
Eduard Strasburger Eduard Adolf Strasburger (1 February 1844 – 18 May 1912) was a Polish- German professor and one of the most famous botanists of the 19th century. He discovered mitosis in plants. Life Eduard Strasburger was born in Warsaw, Congress Poland, t ...
's ''Lehrbuch der Botanik''. Degelius obtained a filosofie kandidat degree in 1927, a
filosofie magister A master's degree (from Latin ) is a postgraduate academic degree awarded by universities or colleges upon completion of a course of study demonstrating mastery or a high-order overview of a specific field of study or area of professional practi ...
in 1929, and a filosofie licentiat in 1933. It was at Uppsala university that Degelius associated with the professor
Rutger Sernander Johan ''Rutger'' Sernander (2November 186627October 1944) was a Swedish botanist, geologist and archaeologist. He was one of the founders of the study of palynology which would later be developed by Lennart von Post, as well as a pioneer in the e ...
at the Växtbiologiska Institutionen (Institute of Botany), from where he graduated in 1935 with a dissertation titled ''Das ozeanische Element der Strauch-und Laubflechtenflora von Skandinavien'' ("The oceanic element of the shrub and lichen flora of Scandinavia").


Career

From 1935 until 1961, Degelius was a
docent The term "docent" is derived from the Latin word , which is the third-person plural present active indicative of ('to teach, to lecture'). Becoming a docent is often referred to as habilitation or doctor of science and is an academic qualifi ...
(an academic appointment below the rank of Professor) in botany at Uppsala University. He worked under the supervision of Professor
Gustaf Einar Du Rietz Gustaf Einar Du Rietz (25 April 1895 – 7 March 1967) was a Swedish botanist and lichenologist. His research interests included plant ecology, lichen taxonomy, and biogeography. He was a leading figure in the Uppsala school of phytosociology ...
, along with Sten Ahlner,
Torsten Hasselrot Thorsten (Thorstein, Torstein, Torsten) is a Scandinavian given name. The Old Norse name was ''Þórsteinn''. It is a compound of the theonym ''Þór'' (''Thor'') and ''steinn'' "stone", which became ''Thor'' and ''sten'' in Old Danish and Old Swe ...
, and
Rolf Santesson Rolf Santesson (1916–2013) was a Swedish lichenologist and university lecturer. He was awarded the Acharius Medal in 1992 for his lifetime contributions to lichenology. Early life and education Santesson was born in 1916 in Trollhättan, Sw ...
– all of whom became notable lichenologists. During this time, Degelius travelled (often with Ahlner and Hasselrot, at least initially) to many locations in
Scandinavia Scandinavia is a subregion#Europe, subregion of northern Europe, with strong historical, cultural, and linguistic ties between its constituent peoples. ''Scandinavia'' most commonly refers to Denmark, Norway, and Sweden. It can sometimes also ...
to collect lichens. In 1939, he visited the United States and subsequently published two papers on the lichens of
Maine Maine ( ) is a U.S. state, state in the New England region of the United States, and the northeasternmost state in the Contiguous United States. It borders New Hampshire to the west, the Gulf of Maine to the southeast, and the Provinces and ...
, and the lichens of the
Great Smoky Mountains The Great Smoky Mountains (, ''Equa Dutsusdu Dodalv'') are a mountain range rising along the Tennessee–North Carolina border in the southeastern United States. They are a subrange of the Appalachian Mountains and form part of the Blue Ridg ...
of
Tennessee Tennessee (, ), officially the State of Tennessee, is a landlocked U.S. state, state in the Southeastern United States, Southeastern region of the United States. It borders Kentucky to the north, Virginia to the northeast, North Carolina t ...
, both of which were highly regarded by later researchers. He has been credited for having introduced the lichen terms and in a 1945 publication, referring to previously undescribed asexual
propagule In biology, a propagule is any material that functions in propagating an organism to the next stage in its life cycle, such as by dispersal. The propagule is usually distinct in form from the parent organism. Propagules are produced by organisms ...
s he had observed on the lichen '' Lempholemma cladodes''. Degelius returned to Gothenburg in 1955 to take a position at the
Gothenburg Botanical Garden The Gothenburg Botanical Garden () is located in Gothenburg, Sweden, and is one of the larger botanical gardens in Europe. History The Gothenburg Botanical Garden is situated in a formerly completely rural area, where earlier a great country e ...
. Degelius also lectured about the subject of systematic botany at the
University of Gothenburg The University of Gothenburg () is a List of universities in Sweden, university in Sweden's second largest city, Gothenburg. Founded in 1891, the university is the third-oldest of the current List of universities in Sweden#Public universities, S ...
until his retirement in 1969. After his retirement, Degelius travelled the world, visiting exotic locations on several continents to continue his lichen studies. He collected his last lichen in Estonia in 1991. His private
herbarium A herbarium (plural: herbaria) is a collection of preserved plant biological specimen, specimens and associated data used for scientific study. The specimens may be whole plants or plant parts; these will usually be in dried form mounted on a sh ...
, which had accumulated more than 50,000 specimens, is now housed at Uppsala University. He described 124 new
taxa In biology, a taxon (back-formation from ''taxonomy''; : taxa) is a group of one or more populations of an organism or organisms seen by taxonomists to form a unit. Although neither is required, a taxon is usually known by a particular name and ...
(mostly species), and published about 130 scientific papers. Although most of these were about lichens, he also published papers on
phanerogam A seed plant or spermatophyte (; New Latin ''spermat-'' and Greek ' (phytón), plant), also known as a phanerogam (taxon Phanerogamae) or a phaenogam (taxon Phaenogamae), is any plant that produces seeds. It is a category of embryophyte (i.e. la ...
s, mosses, and non-lichenized fungi. Degelius was also interested in studies on
floristics Floristics is the study of plants of geographical regions. It is a branch of phytogeography, which technically makes it a branch of botany, geography, and a subbranch of biogeography. Harvard University has a history of research with early contr ...
, publishing
flora Flora (: floras or florae) is all the plant life present in a particular region or time, generally the naturally occurring (indigenous (ecology), indigenous) native plant, native plants. The corresponding term for animals is ''fauna'', and for f ...
s for several Nordic locales as well as other more distant locations. Lichen biology and ecology was another interest, and he published research on the competition between rock-dwelling lichens, discovered a type of
diaspore Diaspore ()also called diasporite, empholite, kayserite, or tanatariteis an aluminium hydroxide oxide mineral, α-AlO(OH), crystallizing in the orthorhombic system and isomorphous with goethite. It occurs sometimes as flattened crystals, but ...
called a lichenized hormocyst, and studied the
ecological succession Ecological succession is the process of how species compositions change in an Community (ecology), ecological community over time. The two main categories of ecological succession are primary succession and secondary succession. Primary successi ...
of lichens growing on ''
Fraxinus ''Fraxinus'' (), commonly called ash, is a genus of plants in the olive and lilac family, Oleaceae, and comprises 45–65 species of usually medium-to-large trees, most of which are deciduous trees, although some Subtropics, subtropical specie ...
'' twigs. Degelius is best known for his work on the genus ''
Collema ''Collema'' (jelly lichen) is a genus of lichens in the family Collemataceae. The photobiont is the cyanobacterium genus ''Nostoc''.Dobson, F.S. (2000) Lichens, an illustrated guide to the British and Irish species. 4th edition. Richmond publish ...
''. His first publication on this group appeared in 1936. In 1954 he published a
monograph A monograph is generally a long-form work on one (usually scholarly) subject, or one aspect of a subject, typically created by a single author or artist (or, sometimes, by two or more authors). Traditionally it is in written form and published a ...
of the European species, and then 20 years later, a world monograph. Degelius won a
Linnean Medal The Linnean Medal of the Linnean Society of London was established in 1888, and is awarded annually to alternately a botanist or a zoologist or (as has been common since 1958) to one of each in the same year. The medal was of gold until 1976, and ...
for this work.


Personal life

In her obituary of Degelius, Polish lichenologist Jolanta Miądlikowska described him as belonging "to the group of scholars who are appreciated not only for their knowledge and scientific achievements, but also for 'humanity'". Degelius was described by former student
Lars Arvidsson Lars is a common male name in Scandinavian countries. Origin ''Lars'' means "from the city of Laurentum". Lars is derived from the Latin name Laurentius, which means "from Laurentum" or "crowned with laurel", and is therefore related to the name ...
as generous and helpful with his students and colleagues, who often visited his home in
Askim Askim () is a town and a former municipality in (from January 1, 2020) Indre Østfold municipality in the county of Østfold, Norway. The administrative centre of the Askim municipality was the town of Askim. Askim was established as a municip ...
(a district of Gothenburg) and made use of his expansive library. He was frequently consulted for assistance with
nomenclatural Nomenclature (, ) is a system of names or terms, or the rules for forming these terms in a particular field of arts or sciences. (The theoretical field studying nomenclature is sometimes referred to as ''onymology'' or ''taxonymy'' ). The principl ...
issues or with translating
Latin Latin ( or ) is a classical language belonging to the Italic languages, Italic branch of the Indo-European languages. Latin was originally spoken by the Latins (Italic tribe), Latins in Latium (now known as Lazio), the lower Tiber area aroun ...
diagnoses. Degelius prided himself on accuracy, and complained about the carelessness of the Latin written by younger colleagues. A former student,
Ingvar Kärnefelt Jan Eric Ingvar Kärnefelt (born 1944) is a Swedish lichenologist. Early life and education Kärnefelt was born in Gothenburg, Sweden in 1944. His initial goal in his higher-level studies at University of Cologne in 1966–1967 was to become a ...
, recalled his typical professorial appearance: "Gunnar usually appeared in his slightly old-fashioned "uniform", i.e. a simple blue club blazer, grey trousers, brown and less well matching shoes. He also generally wore
Kissinger Henry Alfred Kissinger (May 27, 1923 – November 29, 2023) was an American diplomat and political scientist who served as the 56th United States secretary of state from 1973 to 1977 and the 7th national security advisor from 1969 to 1975, se ...
-style spectacles." For the final 30 years of his life, Degelius lived in a
villa A villa is a type of house that was originally an ancient Roman upper class country house that provided an escape from urban life. Since its origins in the Roman villa, the idea and function of a villa have evolved considerably. After the f ...
in Askim, with his sister and brother-in-law. He had a lichenological library comprising more than of shelf space, including original classic works by
Carl Linnaeus Carl Linnaeus (23 May 1707 – 10 January 1778), also known after ennoblement in 1761 as Carl von Linné,#Blunt, Blunt (2004), p. 171. was a Swedish biologist and physician who formalised binomial nomenclature, the modern system of naming o ...
,
Erik Acharius Erik Acharius (10 October 1757 – 14 August 1819) was a Swedish botanist who pioneered the Taxonomy (biology), taxonomy of lichens and is known as the "father of lichenology". Acharius was famously the last pupil of Carl Linnaeus. Life Ac ...
,
Elias Magnus Fries Elias Magnus Fries (15 August 1794 – 8 February 1878) was a Swedish mycologist and botanist. He is sometimes called the Mycology, "Linnaeus of Mycology". In his works he described and assigned botanical names to hundreds of fungus and li ...
, and others. This collection is now kept at the Department of Systematic Botany at Uppsala University. The New Zealander botanist David Galloway, who attended a class of Degelius' in the 1960s, recalled the atmosphere at one of the frequent gatherings there: "Very many will remember the warmly convivial discussion in the spacious book-lined sitting room with candles lit, a bottle of port or punch open on the table, and the cigars circulating. It was a warmly welcoming and special ambience which was very much part of Gunnar ... the celebrated photograph albums of past and present lichenologists would be produced, recent publications would be discussed, minutes would fly past into hours and by midnight when the guests were wilting, Gunnar would just be getting into his stride." Degelius died in Askim on 22 July 1993. At his funeral, a specimen of ''
Degelia atlantica ''Degelia'' is a genus of lichen-forming fungi in the family Pannariaceae. The genus is named after Swedish lichenologist Gunnar Degelius. The genus was circumscribed by Lars Arvidsson and David John Galloway in 1981. Species *'' Degelia atla ...
'' was placed on his coffin by the Norwegian lichenologist
Per Magnus Jørgensen Per Magnus Jørgensen (born 1944) is a Norwegian botanist and lichenologist, and Professor Emeritus of systematic botany at the University of Bergen. He is known for his work on the lichen families Pannariaceae and Collemataceae. Jørgensen wa ...
.


Recognition

Degelius was among the first recipients of the
Acharius Medal __NOTOC__ The Acharius Medal is awarded by the International Association for Lichenology (IAL) for lifetime achievement in lichenology. The organization resolved at its 1990 meeting that it would simultaneously honor professional achievement and ...
in 1992, awarded for his lifetime achievements in lichenology. He was elected an honorary member of the
British Lichen Society The British Lichen Society (BLS) was founded in 1958 with the objective of promoting the study and conservation of lichen. Although the society was founded in London, UK, it is of relevance to lichens worldwide. It has been a registered charity ...
in 1976. An issue of the scientific journal ''The Lichenologist'' was dedicated to him in 1983 on the occasion of his 80th birthday, and an issue of the journal ''Graphis Scripta'' (named for a species of lichen) was dedicated to him for his 90th birthday in 1993.


Eponymy

Three genera have been named to honour Gunnar Degelius: ''
Degelia ''Degelia'' is a genus of lichen-forming fungi in the family Pannariaceae. The genus is named after Swedish lichenologist Gunnar Degelius. The genus was circumscription (taxonomy), circumscribed by Lars Arvidsson and David John Galloway in 1981 ...
'' ; '' Degeliella'' ; and '' Gudelia'' . Degelius has also had several species named after him, including: '' Arthopyrenia degelii'' ; '' Carbonea degelii'' ; '' Chaenotheca degelii'' ; '' Lecanora degelii'' ; '' Lecanora gunnarii'' ; '' Lecidea degeliana'' ; '' Lempholemma degelianum'' ; '' Leptogium degelii'' ; '' Parmelia degelii'' ; '' Pertusaria degelii'' ; '' Pseudocyphellaria degelii'' ; '' Rinodina degeliana'' ; '' Squamarina degelii'' ; '' Stigmidium degelii'' ; and '' Verrucaria degelii'' .


Selected works

A complete listing of Degelius's scientific publications is given in Arvidsson and Galloway's 1999 bibliography. Some of his major works include the following: * * * * * * *


See also

* :Taxa named by Gunnar Degelius


References

{{DEFAULTSORT:Degelius, Gunnar 1903 births 1993 deaths Swedish lichenologists 20th-century Swedish people Scientists from Uppsala Swedish taxonomists Uppsala University alumni Academic staff of the University of Gothenburg Acharius Medal recipients