''Gryphoceratops'' is an extinct

genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of living and fossil organisms as well as viruses. In the hierarchy of biological classification, genus comes above species and below family. In binomial n ...

of leptoceratopsid

ceratopsian Ceratopsia or Ceratopia ( or ; Greek: "horned faces") is a group of herbivorous, beaked dinosaurs that thrived in what are now North America, Europe, and Asia, during the Cretaceous Period, although ancestral forms lived earlier, in the Jurassi ...

dinosaur Dinosaurs are a diverse group of reptiles of the clade Dinosauria. They first appeared during the Triassic period, between 243 and 233.23 million years ago (mya), although the exact origin and timing of the evolution of dinosaurs is t ...

known from the

Late Cretaceous The Late Cretaceous (100.5–66 Ma) is the younger of two epochs into which the Cretaceous Period is divided in the geologic time scale. Rock strata from this epoch form the Upper Cretaceous Series. The Cretaceous is named after ''creta'', ...

Alberta Alberta ( ) is one of the thirteen provinces and territories of Canada. It is part of Western Canada and is one of the three prairie provinces. Alberta is bordered by British Columbia to the west, Saskatchewan to the east, the Northwest Ter ...

, southern

Canada Canada is a country in North America. Its ten provinces and three territories extend from the Atlantic Ocean to the Pacific Ocean and northward into the Arctic Ocean, covering over , making it the world's second-largest country by tota ...

Discovery

''Gryphoceratops'' is known only from the

holotype A holotype is a single physical example (or illustration) of an organism, known to have been used when the species (or lower-ranked taxon) was formally described. It is either the single such physical example (or illustration) or one of seve ...

ROM Rom, or ROM may refer to: Biomechanics and medicine * Risk of mortality, a medical classification to estimate the likelihood of death for a patient * Rupture of membranes, a term used during pregnancy to describe a rupture of the amniotic sac * ...

56635, a partial right

dentary In anatomy, the mandible, lower jaw or jawbone is the largest, strongest and lowest bone in the human facial skeleton. It forms the lower jaw and holds the lower teeth in place. The mandible sits beneath the maxilla. It is the only movable bone ...

. The holotype was collected in the northwest corner of

Dinosaur Provincial Park Dinosaur Provincial Park is a UNESCO World Heritage Site situated a two hour drive east of Calgary, Alberta, Canada; or , about a half-hour drive northeast of Brooks. The park is situated in the Red Deer River valley, which is noted for its stri ...

, from bonebed 55 of the

Milk River Formation The Milk River Formation is a sandstone-dominated stratigraphic unit of the Western Canada Sedimentary Basin in southern Alberta, Canada. It was deposited in near-shore to coastal environments during Late Cretaceous (late Santonian to early Campani ...

, dating to the late

Santonian The Santonian is an age in the geologic timescale or a chronostratigraphic stage. It is a subdivision of the Late Cretaceous Epoch or Upper Cretaceous Series. It spans the time between 86.3 ± 0.7 mya (million years ago) and 83.6 ± 0.7 mya. ...

stage of the middle

period Period may refer to: Common uses * Era, a length or span of time * Full stop (or period), a punctuation mark Arts, entertainment, and media * Period (music), a concept in musical composition * Periodic sentence (or rhetorical period), a concept ...

, about 83.5

million years ago The abbreviation Myr, "million years", is a unit of a quantity of (i.e. ) years, or 31.556926 teraseconds. Usage Myr (million years) is in common use in fields such as Earth science and cosmology. Myr is also used with Mya (million years ago). ...

. Thus, ''Gryphoceratops'' represents the oldest known leptoceratopsid. However, a

cladistic Cladistics (; ) is an approach to biological classification in which organisms are categorized in groups (" clades") based on hypotheses of most recent common ancestry. The evidence for hypothesized relationships is typically shared derived ch ...

analysis found it to be one of the most advanced leptoceratopsids. Additionally, the holotype probably represents the smallest adult ceratopsian known from North America. The cladogram below represents the cladistic relationship of ''Gryphoceratops'' compared to other

leptoceratopsids Leptoceratopsidae is an extinct family of neoceratopsian dinosaurs from Asia, North America and Europe. Leptoceratopsids resembled, and were closely related to, other neoceratopsians, such as the families Protoceratopsidae and Ceratopsidae, but ...

Etymology

''Gryphoceratops'' was first named by Michael J. Ryan,

David C. Evans David Cannon Evans (February 24, 1924 – October 3, 1998) was the founder of the computer science department at the University of Utah and co-founder (with Ivan Sutherland) of Evans & Sutherland, a pioneering firm in computer graphics hardwa ...

Philip J. Currie Philip John Currie (born March 13, 1949) is a Canadian palaeontologist and museum curator who helped found the Royal Tyrrell Museum of Palaeontology in Drumheller, Alberta and is now a professor at the University of Alberta in Edmonton. In the ...

Caleb M. Brown Caleb (), sometimes transliterated as Kaleb ( he, כָּלֵב, ''Kalev'', ; Tiberian vocalization: Kālēḇ; Hebrew Academy: Kalev), is a figure who appears in the Hebrew Bible as a representative of the Tribe of Judah during the Israelit ...

and Don Brinkman in

2012 File:2012 Events Collage V3.png, From left, clockwise: The passenger cruise ship Costa Concordia lies capsized after the Costa Concordia disaster; Damage to Casino Pier in Seaside Heights, New Jersey as a result of Hurricane Sandy; People gather ...

and the

type species In zoological nomenclature, a type species (''species typica'') is the species name with which the name of a genus or subgenus is considered to be permanently taxonomically associated, i.e., the species that contains the biological type specimen( ...

is ''Gryphoceratops morrisoni''. The genus name means "

griffin The griffin, griffon, or gryphon (Ancient Greek: , ''gryps''; Classical Latin: ''grȳps'' or ''grȳpus''; Late Latin, Late and Medieval Latin: ''gryphes'', ''grypho'' etc.; Old French: ''griffon'') is a legendary creature with the body, tail ...

horned face", as a reference to the beaked head, and the species name honors Ian Morrison, who prepared the holotype specimen.

References

Leptoceratopsids Late Cretaceous dinosaurs of North America Fossil taxa described in 2012 Taxa named by Philip J. Currie Paleontology in Alberta Milk River Formation Campanian genus first appearances Campanian genus extinctions Ornithischian genera {{ceratopsian-stub

Discovery

Etymology

See also

References