The emerald tree monitor (''Varanus prasinus'') or green tree monitor, is a small to medium-sized
arboreal
Arboreal locomotion is the locomotion of animals in trees. In habitats in which trees are present, animals have evolved to move in them. Some animals may scale trees only occasionally, but others are exclusively arboreal. The habitats pose num ...
monitor lizard
Monitor lizards are lizards in the genus ''Varanus,'' the only extant genus in the family Varanidae. They are native to Africa, Asia, and Oceania, and one species is also found in the Americas as an invasive species. About 80 species are reco ...
. It is known for its unusual coloration, which consists of shades from green to
turquoise
Turquoise is an opaque, blue-to-green mineral that is a hydrated phosphate of copper and aluminium, with the chemical formula . It is rare and valuable in finer grades and has been prized as a gemstone and ornamental stone for thousands of y ...
, topped with dark,
transverse
Transverse may refer to:
*Transverse engine, an engine in which the crankshaft is oriented side-to-side relative to the wheels of the vehicle
*Transverse flute, a flute that is held horizontally
* Transverse force (or ''Euler force''), the tangen ...
dorsal
Dorsal (from Latin ''dorsum'' ‘back’) may refer to:
* Dorsal (anatomy), an anatomical term of location referring to the back or upper side of an organism or parts of an organism
* Dorsal, positioned on top of an aircraft's fuselage
* Dorsal co ...
banding. This coloration helps
camouflage
Camouflage is the use of any combination of materials, coloration, or illumination for concealment, either by making animals or objects hard to see, or by disguising them as something else. Examples include the leopard's spotted coat, the b ...
it in its arboreal habitat.
Its color also makes the emerald tree monitor highly prized in both the pet trade and zoos alike.
Names
It is known as wbl km in the
Kalam language
Kalam is a Kalam language of Papua New Guinea. It is closely related to Kobon, and shares many of the features of that language. Kalam is spoken in Middle Ramu District of Madang Province and in Mount Hagen District of Western Highlands Provi ...
of
Papua New Guinea
Papua New Guinea (abbreviated PNG; , ; tpi, Papua Niugini; ho, Papua Niu Gini), officially the Independent State of Papua New Guinea ( tpi, Independen Stet bilong Papua Niugini; ho, Independen Stet bilong Papua Niu Gini), is a country i ...
.
Taxonomy
''Varanus prasinus'' was first described as ''Monitor viridis'' by
John Edward Gray
John Edward Gray, Fellow of the Royal Society, FRS (12 February 1800 – 7 March 1875) was a British zoology, zoologist. He was the elder brother of zoologist George Robert Gray and son of the pharmacologist and botanist Samuel Frederick Gray ...
in 1831; however, Gray's original
holotype
A holotype is a single physical example (or illustration) of an organism, known to have been used when the species (or lower-ranked taxon) was formally described. It is either the single such physical example (or illustration) or one of seve ...
(RMNH 4812 in the
National Natural History Museum
National may refer to:
Common uses
* Nation or country
** Nationality – a ''national'' is a person who is subject to a nation, regardless of whether the person has full rights as a citizen
Places in the United States
* National, Maryland, ce ...
in
Leiden
Leiden (; in English and archaic Dutch also Leyden) is a city and municipality in the province of South Holland, Netherlands. The municipality of Leiden has a population of 119,713, but the city forms one densely connected agglomeration wit ...
) was lost and the species was redescribed by Schlegel eight years later as ''V. prasinus'' using the found specimen.
The
generic
Generic or generics may refer to:
In business
* Generic term, a common name used for a range or class of similar things not protected by trademark
* Generic brand, a brand for a product that does not have an associated brand or trademark, other ...
name ''Varanus'' is derived from the
Arabic
Arabic (, ' ; , ' or ) is a Semitic language spoken primarily across the Arab world.Semitic languages: an international handbook / edited by Stefan Weninger; in collaboration with Geoffrey Khan, Michael P. Streck, Janet C. E.Watson; Walte ...
word ''waral'' (ورل), which translates to English as "monitor".
Its
specific
Specific may refer to:
* Specificity (disambiguation)
* Specific, a cure or therapy for a specific illness
Law
* Specific deterrence, focussed on an individual
* Specific finding, intermediate verdict used by a jury in determining the final ...
name, ''prasinus'', is
Latin
Latin (, or , ) is a classical language belonging to the Italic branch of the Indo-European languages. Latin was originally a dialect spoken in the lower Tiber area (then known as Latium) around present-day Rome, but through the power ...
for the color
green
Green is the color between cyan and yellow on the visible spectrum. It is evoked by light which has a dominant wavelength of roughly 495570 Nanometre, nm. In subtractive color systems, used in painting and color printing, it is created by ...
.
''Varanus prasinus'' is a member of the subgenus ''Euprepiosaurus''. It is closely related to several other
arboreal
Arboreal locomotion is the locomotion of animals in trees. In habitats in which trees are present, animals have evolved to move in them. Some animals may scale trees only occasionally, but others are exclusively arboreal. The habitats pose num ...
species; when combined, these are often referred to as the ''V. prasinus''
species group
In biology, a species complex is a group of closely related organisms that are so similar in appearance and other features that the boundaries between them are often unclear. The taxa in the complex may be able to hybridize readily with each oth ...
or species complex. In addition to ''V. prasinus'', this species group, whose members are all
allopatric
Allopatric speciation () – also referred to as geographic speciation, vicariant speciation, or its earlier name the dumbbell model – is a mode of speciation that occurs when biological populations become geographically isolated from ...
, includes ''
V. beccarii'' (
Aru Islands
The Aru Islands Regency ( id, Kabupaten Kepulauan Aru) is a group of about 95 low-lying islands in the Maluku Islands of eastern Indonesia. It also forms a regency of Maluku Province, with a land area of . At the 2011 Census the Regency had a p ...
), ''
V. boehmei'' (
Waigeo Island
Waigeo is an island in Southwest Papua province of eastern Indonesia. The island is also known as Amberi, or Waigiu. It is the largest of the four main islands in the Raja Ampat Islands archipelago, between Halmahera and about to the north-wes ...
), ''
V. bogerti'' (
D'Entrecasteaux Archipelago
D'Entrecasteaux Islands () are situated near the eastern tip of New Guinea in the Solomon Sea in Milne Bay Province of Papua New Guinea. The group spans a distance of , has a total land area of approximately and is separated from the Papua N ...
), ''
V. keithhornei'' (
Cape York Peninsula
Cape York Peninsula is a large peninsula located in Far North Queensland, Australia. It is the largest unspoiled wilderness in northern Australia.Mittermeier, R.E. et al. (2002). Wilderness: Earth’s last wild places. Mexico City: Agrupación ...
), ''
V. kordensis'' (
Biak Island
Biak is an island located in Cenderawasih Bay near the northern coast of Papua, an Indonesian province, and is just northwest of New Guinea. Biak is the largest island in its small archipelago, and has many atolls, reefs, and corals.
The large ...
), ''
V. macraei'' (
Batanta Island), ''
V. reisingeri'' (
Misool Island
Misool, formerly spelled Mysol (Dutch: Misoöl) or Misol, is one of the four major islands in the Raja Ampat Islands in Southwest Papua, Indonesia. Its area is 2,034 km2. The highest point is 561 m and the main towns are Waigama, located ...
) and ''
V. telenesetes'' (
Rossel Island
Rossel Island (named after de Rossel, a senior officer on the French expedition of d'Entrecasteaux, 1791-1793; also known as Yela) is the easternmost island of the Louisiade Archipelago, within the Milne Bay Province of Papua New Guinea. Tree Isl ...
).
Evolutionary history
The family
Varanidae
The Varanidae are a family of lizards in the superfamily Varanoidea within the Anguimorpha group. The family, a group of carnivorous and frugivorous lizards, includes the living genus '' Varanus'' and a number of extinct genera more closely r ...
probably originated in
Asia
Asia (, ) is one of the world's most notable geographical regions, which is either considered a continent in its own right or a subcontinent of Eurasia, which shares the continental landmass of Afro-Eurasia with Africa. Asia covers an ...
about 65 million years ago, although some estimates are as early as the late
Mesozoic
The Mesozoic Era ( ), also called the Age of Reptiles, the Age of Conifers, and colloquially as the Age of the Dinosaurs is the second-to-last era of Earth's geological history, lasting from about , comprising the Triassic, Jurassic and Creta ...
(112 million years ago). Monitor lizards probably migrated to Australia and the Indonesian archipelago between 39 and 26 million years ago. All members of the ''V. prasinus'' species group (sometimes referred to as subgenus ''Euprepiosaurus'') are found east of
Lydekker’s line in the Indo-Australian Archipelago. This group probably originated in the
Moluccas
The Maluku Islands (; Indonesian: ''Kepulauan Maluku'') or the Moluccas () are an archipelago in the east of Indonesia. Tectonically they are located on the Halmahera Plate within the Molucca Sea Collision Zone. Geographically they are located ...
, although dating their origins is complicated by the paucity of the fossil record. The absence of large placental predatory mammals east of
Wallace's line
The Wallace Line or Wallace's Line is a faunal boundary line drawn in 1859 by the British naturalist Alfred Russel Wallace and named by English biologist Thomas Henry Huxley that separates the biogeographical realms of Asia and Wallacea, a tran ...
may be partially responsible for the radiation of the ''V. prasinus'' group in the Australo-Papuan region.
Distribution
Emerald tree monitors and their close relatives can be found on the island of
New Guinea
New Guinea (; Hiri Motu: ''Niu Gini''; id, Papua, or , historically ) is the world's second-largest island with an area of . Located in Oceania in the southwestern Pacific Ocean, the island is separated from Australia by the wide Torr ...
(split between
Indonesia
Indonesia, officially the Republic of Indonesia, is a country in Southeast Asia and Oceania between the Indian and Pacific oceans. It consists of over 17,000 islands, including Sumatra, Java, Sulawesi, and parts of Borneo and New Guine ...
and
Papua New Guinea
Papua New Guinea (abbreviated PNG; , ; tpi, Papua Niugini; ho, Papua Niu Gini), officially the Independent State of Papua New Guinea ( tpi, Independen Stet bilong Papua Niugini; ho, Independen Stet bilong Papua Niu Gini), is a country i ...
) as well as several adjacent islands,
and the northern
Torres Strait Islands
The Torres Strait Islands are a group of at least 274 small islands in the Torres Strait, a waterway separating far northern continental Australia's Cape York Peninsula and the island of New Guinea. They span an area of , but their total land ...
.
The green tree monitor is reported to thrive in lowland environments, including tropical evergreen forests, palm swamps and
cocoa
Cocoa may refer to:
Chocolate
* Chocolate
* ''Theobroma cacao'', the cocoa tree
* Cocoa bean, seed of ''Theobroma cacao''
* Chocolate liquor, or cocoa liquor, pure, liquid chocolate extracted from the cocoa bean, including both cocoa butter and ...
plantations.
[
]
Description
The emerald tree monitor is about long with a slender body that helps it support itself on narrow branches. It uses its prehensile
Prehensility is the quality of an appendage or organ that has adapted for grasping or holding. The word is derived from the Latin term ''prehendere'', meaning "to grasp". The ability to grasp is likely derived from a number of different origi ...
tail and long claws to grip branches. Unlike other varanids, this monitor defends its tail rather than lashing with it for defense when threatened. The soles of the feet of the emerald tree monitor have enlarged scales which aid the lizard when climbing.
Ecology
When threatened, the emerald tree monitor will flee through vegetation or bite if cornered.
It is one of the few social monitors, living in small groups made up of a dominant male, several females, and a few other males and juveniles.
Diet
The emerald tree monitor's diet consists of large tree-dwelling arthropod
Arthropods (, (gen. ποδός)) are invertebrate animals with an exoskeleton, a Segmentation (biology), segmented body, and paired jointed appendages. Arthropods form the phylum Arthropoda. They are distinguished by their jointed limbs and Arth ...
s, such as katydid
Insects in the family Tettigoniidae are commonly called katydids (especially in North America), or bush crickets. They have previously been known as "long-horned grasshoppers". More than 8,000 species are known. Part of the suborder Ensifera, ...
s, grasshoppers, other orthopteran
Orthoptera () is an order of insects that comprises the grasshoppers, locusts, and crickets, including closely related insects, such as the bush crickets or katydids and wētā. The order is subdivided into two suborders: Caelifera – grasshop ...
s, stick insect
The Phasmatodea (also known as Phasmida, Phasmatoptera or Spectra) are an order of insects whose members are variously known as stick insects, stick-bugs, walking sticks, stick animals, or bug sticks. They are also occasionally referred to as D ...
s, cockroach
Cockroaches (or roaches) are a Paraphyly, paraphyletic group of insects belonging to Blattodea, containing all members of the group except termites. About 30 cockroach species out of 4,600 are associated with human habitats. Some species are we ...
es, beetle
Beetles are insects that form the order Coleoptera (), in the superorder Endopterygota. Their front pair of wings are hardened into wing-cases, elytra, distinguishing them from most other insects. The Coleoptera, with about 400,000 describ ...
s, centipede
Centipedes (from New Latin , "hundred", and Latin , " foot") are predatory arthropods belonging to the class Chilopoda (Ancient Greek , ''kheilos'', lip, and New Latin suffix , "foot", describing the forcipules) of the subphylum Myriapoda, an ...
s, spider
Spiders (order Araneae) are air-breathing arthropods that have eight legs, chelicerae with fangs generally able to inject venom, and spinnerets that extrude silk. They are the largest order of arachnids and rank seventh in total species d ...
s and crabs, as well as bird
Birds are a group of warm-blooded vertebrates constituting the class Aves (), characterised by feathers, toothless beaked jaws, the laying of hard-shelled eggs, a high metabolic rate, a four-chambered heart, and a strong yet lightweig ...
s and small mammals (such as the Moncton's mosaic-tailed rat
Moncton's mosaic-tailed rat (''Paramelomys moncktoni'', previously known as ''Melomys moncktoni'') is a species of rodent in the family Muridae.
It is found only in Papua New Guinea
Papua New Guinea (abbreviated PNG; , ; tpi, Papua Niu ...
). Before swallowing stick insects, the lizards tear off the legs.[ ]Captive
Captive or Captives may refer to:
Arts, entertainment, and media Films
* ''Captive'' (1980 film), a sci-fi film, starring Cameron Mitchell and David Ladd
* ''Captive'' (1986 film), a British-French film starring Oliver Reed
* ''Captive'' (1991 ...
specimens tear off the limbs of rodents prior to eating them; as a result, they are capable of swallowing mammals of a considerable size: A 135-g lizard was documented as eating a 40-g rodent, almost one-third its size. Paleontologist and Temple University
Temple University (Temple or TU) is a public university, public Commonwealth System of Higher Education, state-related research university in Philadelphia, Pennsylvania. It was founded in 1884 by the Baptists, Baptist minister Russell Conwell an ...
professor Michael Balsai has observed ''V. prasinus'' eating fruit (bananas) in captivity as has herpetologist and author Robert G. Sprackland.
Reproduction
Clutches consist of up to five eggs
Humans and human ancestors have scavenged and eaten animal eggs for millions of years. Humans in Southeast Asia had domesticated chickens and harvested their eggs for food by 1,500 BCE. The most widely consumed eggs are those of fowl, especial ...
, each weighing and measuring about . As many as three clutch
A clutch is a mechanical device that engages and disengages power transmission, especially from a drive shaft to a driven shaft. In the simplest application, clutches connect and disconnect two rotating shafts (drive shafts or line shafts ...
es are laid throughout the year; clutches have been laid by captives in January, March, April, November, and December. The female emerald tree monitor lays her eggs in arboreal termite
Termites are small insects that live in colonies and have distinct castes ( eusocial) and feed on wood or other dead plant matter. Termites comprise the infraorder Isoptera, or alternatively the epifamily Termitoidae, within the order Blat ...
nests. The eggs hatch between 160 and 190 days later, typically from June to November, after which the young eat the termites and the termite's eggs within minutes of hatching. Sexual maturity is reached in about one year.
References
External links
Photos at Biolib.cz
Mampam.com
{{Taxonbar, from=Q852763
Varanus
Monitor lizards of Australia
Reptiles of Indonesia
Monitor lizards of New Guinea
Reptiles of Papua New Guinea
Reptiles described in 1839
Taxa named by Hermann Schlegel