The greater roadrunner (''Geococcyx californianus'') is a long-legged
bird
Birds are a group of warm-blooded vertebrates constituting the class (biology), class Aves (), characterised by feathers, toothless beaked jaws, the Oviparity, laying of Eggshell, hard-shelled eggs, a high Metabolism, metabolic rate, a fou ...
Southwestern United States
The Southwestern United States, also known as the American Southwest or simply the Southwest, is a geographic and cultural list of regions of the United States, region of the United States that includes Arizona and New Mexico, along with adjacen ...
and
Mexico
Mexico, officially the United Mexican States, is a country in North America. It is the northernmost country in Latin America, and borders the United States to the north, and Guatemala and Belize to the southeast; while having maritime boundar ...
. The
scientific name
In Taxonomy (biology), taxonomy, binomial nomenclature ("two-term naming system"), also called binary nomenclature, is a formal system of naming species of living things by giving each a name composed of two parts, both of which use Latin gramm ...
means "Californian earth-cuckoo". Along with the lesser roadrunner, it is one of two species in the genus '' Geococcyx''. This roadrunner is also known as the chaparral cock, ground cuckoo, and snake killer.
Taxonomy and systematics
Greater roadrunner fossils dating from the
Holocene
The Holocene () is the current geologic time scale, geological epoch, beginning approximately 11,700 years ago. It follows the Last Glacial Period, which concluded with the Holocene glacial retreat. The Holocene and the preceding Pleistocene to ...
and
Pleistocene
The Pleistocene ( ; referred to colloquially as the ''ice age, Ice Age'') is the geological epoch (geology), epoch that lasted from to 11,700 years ago, spanning the Earth's most recent period of repeated glaciations. Before a change was fin ...
have been found in
California
California () is a U.S. state, state in the Western United States that lies on the West Coast of the United States, Pacific Coast. It borders Oregon to the north, Nevada and Arizona to the east, and shares Mexico–United States border, an ...
,
Arizona
Arizona is a U.S. state, state in the Southwestern United States, Southwestern region of the United States, sharing the Four Corners region of the western United States with Colorado, New Mexico, and Utah. It also borders Nevada to the nort ...
,
New Mexico
New Mexico is a state in the Southwestern United States, Southwestern region of the United States. It is one of the Mountain States of the southern Rocky Mountains, sharing the Four Corners region with Utah, Colorado, and Arizona. It also ...
, and
Texas
Texas ( , ; or ) is the most populous U.S. state, state in the South Central United States, South Central region of the United States. It borders Louisiana to the east, Arkansas to the northeast, Oklahoma to the north, New Mexico to the we ...
, in the
United States
The United States of America (USA), also known as the United States (U.S.) or America, is a country primarily located in North America. It is a federal republic of 50 U.S. state, states and a federal capital district, Washington, D.C. The 48 ...
Nuevo León
Nuevo León, officially the Free and Sovereign State of Nuevo León, is a Administrative divisions of Mexico, state in northeastern Mexico. The state borders the Mexican states of Tamaulipas, Coahuila, Zacatecas, and San Luis Potosí, San Luis ...
. The oldest known fossil comes from a cave in New Mexico, estimated at an age of 33,500 years. In the
La Brea Tar Pits
La Brea Tar Pits comprise an active Paleontological site, paleontological research site in urban Los Angeles. Hancock Park was formed around a group of tar pits where natural Bitumen, asphalt (also called asphaltum, bitumen, or pitch; ''brea'' ...
, fragments from 25 greater roadrunner fossils have been found. Several other fossils are also known from Santa Barbara and Kern counties, as well as
Northern Mexico
Northern Mexico ( ), commonly referred as , is an informal term for the northern cultural and geographical area in Mexico. Depending on the source, it contains some or all of the states of Baja California, Baja California Sur, Chihuahua (state), ...
.
Prehistoric remains indicate that until 8,000 years ago, the greater roadrunner was found in sparse forests rather than scrubby deserts; only later did it adapt to arid environments. Due to this, along with human transformation of the landscape, it has recently started to move northeast of its normal distribution. Sparse forests can be found in these parts, in an environment similar to the prehistoric North American Southwest.
Description
The greater roadrunner is about long, has a wingspan and weighs . It stands around tall and is the largest cuckoo of the Americas. The upper body is mostly brown with black streaks and sometimes pink spots. The neck and upper breast are white or pale brown with dark brown streaks, and the belly is white. A crest of brown feathers sticks up on the head, and a bare patch of orange and blue skin lies behind each eye; the blue is replaced by white in adult males (except the blue adjacent to the eye), and the orange (to the rear) is often hidden by feathers. Males and females have identical plumage. Females are slightly smaller, on average shorter and lighter than males. The long stout beak is grayish brown to gray and has a hooked tip. Roadrunners have four toes on each zygodactyl foot; two face forward, and two face backward. The toes are brown in color and have pale gold spots.
Although capable of limited flight, it spends most of its time on the ground, and can run at speeds up to . Cases where roadrunners have run as fast as have been reported. This is the fastest running speed clocked for a flying bird, but not nearly as fast as the of the completely flightless and much larger ostrich.
Vocalizations
The vocalizations of the greater roadrunner have seven distinct variants. The most frequent call is a slow and descending sequence of about six low, "cooing" noises, emitted by the male and which is heard at . This call is usually made early in the morning, from a high perch such as a fence post, dead tree or cactus. Females give off a number of up to twenty-two short, low-frequency shrills, resembling coyote squeals, which can be heard away. Both male and female roadrunners emit a series of five or six chatters accompanied by groaning, loud enough to be heard away. This sound is the roadrunner's most common vocalization during the incubation period and the rearing of chicks.
Distribution and habitat
The greater roadrunner is found in the Aridoamerica ecoregion, within the
southwestern United States
The Southwestern United States, also known as the American Southwest or simply the Southwest, is a geographic and cultural list of regions of the United States, region of the United States that includes Arizona and New Mexico, along with adjacen ...
and northern
Mexico
Mexico, officially the United Mexican States, is a country in North America. It is the northernmost country in Latin America, and borders the United States to the north, and Guatemala and Belize to the southeast; while having maritime boundar ...
. It can be seen regularly in the US states of California, Texas, New Mexico, Arizona,
Nevada
Nevada ( ; ) is a landlocked state in the Western United States. It borders Oregon to the northwest, Idaho to the northeast, California to the west, Arizona to the southeast, and Utah to the east. Nevada is the seventh-most extensive, th ...
,
Oklahoma
Oklahoma ( ; Choctaw language, Choctaw: , ) is a landlocked U.S. state, state in the South Central United States, South Central region of the United States. It borders Texas to the south and west, Kansas to the north, Missouri to the northea ...
,
Utah
Utah is a landlocked state in the Mountain states, Mountain West subregion of the Western United States. It is one of the Four Corners states, sharing a border with Arizona, Colorado, and New Mexico. It also borders Wyoming to the northea ...
and a tiny bit of
Colorado
Colorado is a U.S. state, state in the Western United States. It is one of the Mountain states, sharing the Four Corners region with Arizona, New Mexico, and Utah. It is also bordered by Wyoming to the north, Nebraska to the northeast, Kansas ...
, and less frequently in
Kansas
Kansas ( ) is a landlocked U.S. state, state in the Midwestern United States, Midwestern region of the United States. It borders Nebraska to the north; Missouri to the east; Oklahoma to the south; and Colorado to the west. Kansas is named a ...
,
Louisiana
Louisiana ( ; ; ) is a state in the Deep South and South Central regions of the United States. It borders Texas to the west, Arkansas to the north, and Mississippi to the east. Of the 50 U.S. states, it ranks 31st in area and 25 ...
,
Arkansas
Arkansas ( ) is a landlocked state in the West South Central region of the Southern United States. It borders Missouri to the north, Tennessee and Mississippi to the east, Louisiana to the south, Texas to the southwest, and Oklahoma ...
, and
Missouri
Missouri (''see #Etymology and pronunciation, pronunciation'') is a U.S. state, state in the Midwestern United States, Midwestern region of the United States. Ranking List of U.S. states and territories by area, 21st in land area, it border ...
, as well as the Mexican states of
Baja California
Baja California, officially the Free and Sovereign State of Baja California, is a state in Mexico. It is the northwesternmost of the 32 federal entities of Mexico. Before becoming a state in 1952, the area was known as the North Territory of B ...
Sonora
Sonora (), officially Estado Libre y Soberano de Sonora (), is one of the 31 states which, along with Mexico City, comprise the Administrative divisions of Mexico, Federal Entities of Mexico. The state is divided into Municipalities of Sonora, 72 ...
,
Sinaloa
Sinaloa (), officially the (), is one of the 31 states which, along with Mexico City, compose the Federal Entities of Mexico. It is divided into 18 municipalities, and its capital city is Culiacán Rosales.
It is located in northwest Mexic ...
Durango
Durango, officially the Free and Sovereign State of Durango, is one of the 31 states which make up the Political divisions of Mexico, 32 Federal Entities of Mexico, situated in the northwest portion of the country. With a population of 1,832,650 ...
,
Jalisco
Jalisco, officially the Free and Sovereign State of Jalisco, is one of the 31 states which, along with Mexico City, comprise the 32 Political divisions of Mexico, Federal Entities of Mexico. It is located in western Mexico and is bordered by s ...
Zacatecas
Zacatecas, officially the Free and Sovereign State of Zacatecas, is one of the Political divisions of Mexico, 31 states of Mexico. It is divided into Municipalities of Zacatecas, 58 municipalities and its capital city is Zacatecas City, Zacatec ...
,
Aguascalientes
Aguascalientes, officially the Free and Sovereign State of Aguascalientes, is one of the 32 states which comprise the Political divisions of Mexico, Federal Entities of Mexico. At 22°N and with an average altitude of above sea level it is pre ...
,
Guanajuato
Guanajuato, officially the Free and Sovereign State of Guanajuato, is one of the 32 states that make up the Political divisions of Mexico, Federal Entities of Mexico. It is divided into Municipalities of Guanajuato, 46 municipalities and its cap ...
,
Michoacán
Michoacán, formally Michoacán de Ocampo, officially the Free and Sovereign State of Michoacán de Ocampo, is one of the 31 states which, together with Mexico City, compose the Political divisions of Mexico, Federal Entities of Mexico. The stat ...
,
Querétaro
Querétaro, officially the Free and Sovereign State of Querétaro, is one of the Political divisions of Mexico, 32 federal entities of Mexico. It is divided into Municipalities of Querétaro, 18 municipalities. Its capital city is Querétaro Cit ...
,
México
Mexico, officially the United Mexican States, is a country in North America. It is the northernmost country in Latin America, and borders the United States to the north, and Guatemala and Belize to the southeast; while having maritime boundar ...
,
Puebla
Puebla, officially the Free and Sovereign State of Puebla, is one of the 31 states that, along with Mexico City, comprise the Federal Entities of Mexico. It is divided into 217 municipalities and its capital is Puebla City. Part of east-centr ...
, Nuevo León,
Tamaulipas
Tamaulipas, officially the Free and Sovereign State of Tamaulipas, is a state in Mexico; one of the 31 states which, along with Mexico City, comprise the 32 federal entities of Mexico. It is divided into 43 municipalities.
It is located in nor ...
, and
San Luis Potosí
San Luis Potosí, officially the Free and Sovereign State of San Luis Potosí, is one of the 32 states which compose the Federal Entities of Mexico. It is divided in 59 municipalities and is named after its capital city, San Luis Potosí.
It ...
. The species is not migratory.
The greater roadrunner can be found from below sea level to , but rarely above . It occupies arid and semiarid
scrubland
Shrubland, scrubland, scrub, brush, or bush is a plant community characterized by vegetation dominance (ecology), dominated by shrubs, often also including grasses, herbaceous plant, herbs, and geophytes. Shrubland may either occur naturally o ...
, with scattered vegetation (typically less than 50% cover) with a height that doesn't exceed .
Behavior
Breeding and nesting
Until a mate is found, the greater roadrunner typically lives a solitary life. The greater roadrunner is monogamous, forming long-term
pair bond
In biology, a pair bond is the strong affinity that develops in some species between a mating pair, often leading to the production and rearing of young and potentially a lifelong bond. Pair-bonding is a term coined in the 1940s that is frequently ...
s. Breeding season begins from mid-March to early September. Courtship behaviors between greater roadrunners are a lengthy process including a combination of chasing, tail wagging, play-fighting, and acoustic signals. Once the chasing stage has passed, male greater roadrunners will present nesting materials or food to the females.
Often males will offer food to females during the act of copulation. A unique feature of the greater roadrunner is that mated couples will continue their copulation rituals long after the need for egg fertilization. This factor is believed to contribute to the couple's pair bond maintenance. Greater roadrunner couples defend a territory of about in size. The male is more territorial, calling out to warn competitors, and does not hesitate to physically push the intruders out of his territory. Some couples defend the same territory all year long.
Both birds build the nest, with the male collecting the material and the female constructing the nest. The nests are compact platforms of thorny branches lined with grasses, feathers, snakeskin, roots, and other fine material. They are built low in a cactus, bush, or man-made structure; 3–10 feet above ground. Greater roadrunners lay three to six eggs, which hatch in 20 days. The chicks
fledge
Fledging is the stage in a flying animal's life between egg, hatching or birth and becoming capable of flight.
This term is most frequently applied to birds, but is also used for bats. For altricial birds, those that spend more time in vulnera ...
in another 18 days. Pairs may occasionally rear a second brood when there is an abundance of food in rainy summers. A young fledge will typically remain with its parents until it is at least 50 days old.
Similarly to some other cuckoos, greater roadrunners occasionally lay their eggs in the nests of other birds, such as the common raven and northern mockingbird.
Predators
The primary predators of this species include ground predators ( coyotes,
bobcat
The bobcat (''Lynx rufus''), also known as the wildcat, bay lynx, or red lynx, is one of the four extant species within the medium-sized wild cat genus '' Lynx''. Native to North America, it ranges from southern Canada through most of the c ...
s,
lynx
A lynx ( ; : lynx or lynxes) is any of the four wikt:extant, extant species (the Canada lynx, Iberian lynx, Eurasian lynx and the bobcat) within the medium-sized wild Felidae, cat genus ''Lynx''. The name originated in Middle Engl ...
es and
cougar
The cougar (''Puma concolor'') (, ''Help:Pronunciation respelling key, KOO-gər''), also called puma, mountain lion, catamount and panther is a large small cat native to the Americas. It inhabits North America, North, Central America, Cent ...
The greater roadrunner is omnivorous and uses its speed to outrun and catch prey. It feeds mainly on small animals, such as insects, spiders (including black widows and
tarantula
Tarantulas comprise a group of large and often hairy spiders of the family Theraphosidae. , 1,100 species have been identified, with 166 genera. The term "tarantula" is usually used to describe members of the family Theraphosidae, although ...
s),
centipede
Centipedes (from Neo-Latin , "hundred", and Latin , "foot") are predatory arthropods belonging to the class Chilopoda (Ancient Greek , ''kheilos'', "lip", and Neo-Latin suffix , "foot", describing the forcipules) of the subphylum Myriapoda, ...
s,
scorpion
Scorpions are predatory arachnids of the Order (biology), order Scorpiones. They have eight legs and are easily recognized by a pair of Chela (organ), grasping pincers and a narrow, segmented tail, often carried in a characteristic forward cur ...
s,
mice
A mouse (: mice) is a small rodent. Characteristically, mice are known to have a pointed snout, small rounded ears, a body-length scaly tail, and a high breeding rate. The best known mouse species is the common house mouse (''Mus musculus' ...
, small birds (including
hummingbird
Hummingbirds are birds native to the Americas and comprise the Family (biology), biological family Trochilidae. With approximately 366 species and 113 genus, genera, they occur from Alaska to Tierra del Fuego, but most species are found in Cen ...
s), lizards and young rattlesnakes, and some plants. Some instances of the greater roadrunner feeding on the dead carcasses of larger mammals (including bats, ground
squirrel
Squirrels are members of the family Sciuridae (), a family that includes small or medium-sized rodents. The squirrel family includes tree squirrels, ground squirrels (including chipmunks and prairie dogs, among others), and flying squirrel ...
s, and one juvenile desert cottontail) have been observed. They are opportunistic and are known to feed on eggs and young of other birds as well as carrion. Early pioneer accounts report that when the roadrunner "sees a rattlesnake, it will gather pieces of cactus and put them around the snake, in such manner that escape is impossible".
Thermoregulation
Because of the greater roadrunner's diurnal nature and arid habitat, it has various biological and behavioral
adaptation
In biology, adaptation has three related meanings. Firstly, it is the dynamic evolutionary process of natural selection that fits organisms to their environment, enhancing their evolutionary fitness. Secondly, it is a state reached by the p ...
s, known as
thermoregulation
Thermoregulation is the ability of an organism to keep its body temperature within certain boundaries, even when the surrounding temperature is very different. A thermoconforming organism, by contrast, simply adopts the surrounding temperature ...
, to reduce dehydration and overheating. During the hot season, it is active mostly from sunrise to mid-morning, and late afternoon to evening. It rests in the shade during the hottest part of the day.
Body water
In physiology, body water is the water content of an animal body that is contained in the tissues, the blood, the bones and elsewhere. The percentages of body water contained in various fluid compartments add up to total body water (TBW). This ...
may be retained via liquid reabsorption, by the mucous membranes in the
cloaca
A cloaca ( ), : cloacae ( or ), or vent, is the rear orifice that serves as the only opening for the digestive (rectum), reproductive, and urinary tracts (if present) of many vertebrate animals. All amphibians, reptiles, birds, cartilagin ...
,
rectum
The rectum (: rectums or recta) is the final straight portion of the large intestine in humans and some other mammals, and the gut in others. Before expulsion through the anus or cloaca, the rectum stores the feces temporarily. The adult ...
and
caecum
The cecum ( caecum, ; plural ceca or caeca, ) is a pouch within the peritoneum that is considered to be the beginning of the large intestine. It is typically located on the right side of the body (the same side of the body as the appendix, ...
. The roadrunner's nasal glands eliminate excess body salts.
The greater roadrunner reduces excess heat by the formation of
water vapor
Water vapor, water vapour, or aqueous vapor is the gaseous phase of Properties of water, water. It is one Phase (matter), state of water within the hydrosphere. Water vapor can be produced from the evaporation or boiling of liquid water or from th ...
, released by breathing or through the skin. It sometimes pants in heavy heat to accelerate this action. At night, it reduces its energy expenditure by more than 30 percent, lowering its body temperature from . In the morning, it accelerates heat recovery by sunbathing. In winter, it takes refuge in dense vegetation or among rocks to shelter from cold winds.
The roadrunner frequently sunbathes for warmth. It turns perpendicular to the ground with its back turned towards the sun. Wings apart, the roadrunner ruffles the black feathers on its back and head, exposing its black skin, allowing both skin and feathers to absorb the heat of the sun's rays. Early in the morning, it can stay in this posture for two or three hours. In winter, when the temperatures are around , roadrunners may warm themselves in the sun several times during the day for more than half an hour at a time.
Locomotion
The greater roadrunner can maintain a speed of over long distances. While running, it places its head and tail parallel to the ground and uses its tail as a rudder to help change its direction. It prefers to run in open areas, such as roads, packed trails and dry riverbeds rather than dense vegetation.
The roadrunner less often engages in flight. It hovers from a perch, such as a tree or a human construction. More rarely, it flies short distances of between potential roosts.
Relationship to humans
Some
Pueblo
Pueblo refers to the settlements of the Pueblo peoples, Native American tribes in the Southwestern United States, currently in New Mexico, Arizona, and Texas. The permanent communities, including some of the oldest continually occupied settlement ...
Hopi
The Hopi are Native Americans who primarily live in northeastern Arizona. The majority are enrolled in the Hopi Tribe of Arizona and live on the Hopi Reservation in northeastern Arizona; however, some Hopi people are enrolled in the Colorado ...
, believed the roadrunner provided protection against evil spirits. In Mexico, some said it brought babies, as the white stork was said to in Europe. Some Anglo frontier people believed roadrunners led lost people to trails.Wile E. Coyote and the Road Runner are the two main characters and protagonists of a long-running (since 1949) Warner Bros. animated series.
The greater roadrunner is the state bird of New Mexico and, as such, appeared in a 1982 sheet of 20-cent United States stamps showing 50 state birds and flowers.
It is also the mascot of numerous high schools and colleges in the United States, including California State University, Bakersfield and the
University of Texas at San Antonio
The University of Texas at San Antonio (UTSA or UT San Antonio) is a Public university, public research university in San Antonio, Texas, United States. Established in 1969, The
College of DuPage
College of DuPage is a Public college, public community colleges in the United States, community college with its main campus in Glen Ellyn, Illinois. The college also owns and operates satellite campuses in Addison, Illinois, Addison, Carol St ...
mascot takes the bird's alternate name, Chapparal, inspired by students driving between various temporary classroom locations before the main campus was fully constructed. The roadrunner is also the mascot of the Tucson Roadrunners, a professional hockey team in
Tucson, Arizona
Tucson (; ; ) is a city in Pima County, Arizona, United States, and its county seat. It is the second-most populous city in Arizona, behind Phoenix, Arizona, Phoenix, with a population of 542,630 in the 2020 United States census. The Tucson ...
.
References
Sources
*
Further reading
Montalvo, A. E., Ransom, D., & Lopez, R. R. (2014). Modeling Greater Roadrunners’ (Geococcyx californianus) Habitat Use in West Texas. The Wilson Journal of Ornithology, 126(2), 359–366. http://www.jstor.org/stable/26455981