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''Ginkgo dissecta'' is an extinct ginkgo species in the family Ginkgoaceae described from a series of isolated
fossil A fossil (from Classical Latin , ) is any preserved remains, impression, or trace of any once-living thing from a past geological age. Examples include bones, shells, exoskeletons, stone imprints of animals or microbes, objects preserved ...
leaves A leaf (plural, : leaves) is any of the principal appendages of a vascular plant plant stem, stem, usually borne laterally aboveground and specialized for photosynthesis. Leaves are collectively called foliage, as in "autumn foliage", wh ...
. The species is known from
Early Eocene In the geologic timescale the Ypresian is the oldest age or lowest stratigraphic stage of the Eocene. It spans the time between , is preceded by the Thanetian Age (part of the Paleocene) and is followed by the Eocene Lutetian Age. The Ypresian i ...
sediments exposed in the province of
British Columbia British Columbia (commonly abbreviated as BC) is the westernmost province of Canada, situated between the Pacific Ocean and the Rocky Mountains. It has a diverse geography, with rugged landscapes that include rocky coastlines, sandy beaches, ...
, Canada, and
Washington Washington commonly refers to: * Washington (state), United States * Washington, D.C., the capital of the United States ** A metonym for the federal government of the United States ** Washington metropolitan area, the metropolitan area centered on ...
, USA. It is one of two ''Ginkgo'' species found at the Washington and British Columbia sites.


History and classification

''Ginkgo dissecta'' is represented by a group of fossil specimens from four different geologic formations. The type locality for the species is at the
Ypresian In the geologic timescale the Ypresian is the oldest age (geology), age or lowest stage (stratigraphy), stratigraphic stage of the Eocene. It spans the time between , is preceded by the Thanetian Age (part of the Paleocene) and is followed by th ...
McAbee Fossil Beds The McAbee Fossil Beds is a Heritage Site that protects an Eocene Epoch fossil locality east of Cache Creek, British Columbia, Canada, just north of and visible from Provincial Highway 97 / the Trans-Canada Highway ( Highway 1). The McAbee Fos ...
, near
Cache Creek, British Columbia Cache Creek is a historic transportation junction and incorporated village northeast of Vancouver in British Columbia, Canada. It is on the Trans-Canada Highway in the province of British Columbia at a junction with Highway 97. The same int ...
, in the Tranquille Formation belonging to the
Kamloops Group Kamloops ( ) is a city in south-central British Columbia, Canada, at the confluence of the South flowing North Thompson River and the West flowing Thompson River, east of Kamloops Lake. It is located in the Thompson-Nicola Regional District, w ...
. ''G. dissecta'' is also known from the similarly aged sites of the
Klondike Mountain Formation The Klondike Mountain Formation is an Early Eocene (Ypresian) geological formation located in the northeast central area of Washington state. The formation, named for the type location designated in 1962, Klondike Mountain north of Republic, Wash ...
, which crop out around the town of
Republic A republic () is a "state in which power rests with the people or their representatives; specifically a state without a monarchy" and also a "government, or system of government, of such a state." Previously, especially in the 17th and 18th c ...
, Ferry County, Washington. At least one specimen has been recovered from the fossil sites around the town of
Princeton, British Columbia Princeton (originally Vermilion Forks) is a town in the Similkameen region of southern British Columbia, Canada."The rich history of Princeton or how Vermilion Forks made it on the map...", Princeton 2008 Visitors Guide, p. 4. It lies just east o ...
, and a fourth occurrence for the species was reported from the "Falkland site" near the town of
Falkland, British Columbia Falkland is an unincorporated community located in the Columbia-Shuswap Regional District of British Columbia, Canada between Vernon, British Columbia, Vernon and Kamloops. Falkland is recognized for one of Canada's largest Canadian flags, whic ...
, also part of the Tranquille Formation. The
type specimen In biology, a type is a particular wiktionary:en:specimen, specimen (or in some cases a group of specimens) of an organism to which the scientific name of that organism is formally attached. In other words, a type is an example that serves to a ...
s for ''G. dissecta'' include two leaf fossils, a
holotype A holotype is a single physical example (or illustration) of an organism, known to have been used when the species (or lower-ranked taxon) was formally described. It is either the single such physical example (or illustration) or one of several ...
and a paratype, both from the McAbee Fossil Beds. The holotype leaf is numbered number WWU-GK-020 and the paratype is number WWU-GK-008, both of which are currently preserved in the
paleobotanical Paleobotany, which is also spelled as palaeobotany, is the branch of botany dealing with the recovery and identification of plant remains from geological contexts, and their use for the biological reconstruction of past environments (paleogeogr ...
collections of Western Washington University in Bellingham, Washington. In addition to the two type specimens, the Western Washington University collections have 39 other fossils from the McAbee site and one from the Princeton fossil sites. For the species description one fossil, SR 96-09-01, was borrowed from the Stonerose Interpretive Center in Republic, Washington. The specimens were studied by paleobotanist George Mustoe of the Western Washington University Geology Department. Mustoe published his 2002
type description A species description is a formal description of a newly discovered species, usually in the form of a scientific paper. Its purpose is to give a clear description of a new species of organism and explain how it differs from species that have be ...
for ''G. dissecta'' in the '' Canadian Journal of Botany''. The
etymology Etymology ()The New Oxford Dictionary of English (1998) – p. 633 "Etymology /ˌɛtɪˈmɒlədʒi/ the study of the class in words and the way their meanings have changed throughout time". is the study of the history of the Phonological chan ...
of the chosen
specific name Specific name may refer to: * in Database management systems, a system-assigned name that is unique within a particular database In taxonomy, either of these two meanings, each with its own set of rules: * Specific name (botany), the two-part (bino ...
''dissecta'' was not identified by Mustoe in the type description, but he noted it is a formalization of the name which had been first used in 1974 in an unpublished thesis by Verschoor.


Description

Leaf widths of ''G. dissecta'' range between and petioles reaching up to in length. In contrast to the two-lobed structure of modern ''
Ginkgo biloba ''Ginkgo biloba'', commonly known as ginkgo or gingko ( ), also known as the maidenhair tree, is a species of tree native to China. It is the last living species in the order Ginkgoales, which first appeared over 290 million years ago. Fossils ...
'' leaves, the morphology of ''G. dissecta'' is four-lobed. These lobes are separated by deep notches which almost reach the leaf base, while additional smaller notches may be present in the upper portions of the lobes. The vein structure of ''G. dissecta'' is composed of numerous forking veins which start at the base of the leaf. This forking is different from the vein structure of ''G. biloba'' in which all the veins fork from a single vein along the edge of the blade. The overall vein density in ''G. dissecta'' is 12–14 veins per centimeter, which is lower than the reported average of 16–17 veins per centimeter in ''G. biloba''. ''G. dissecta'' is superficially similar to the older species '' G. digitata'' and '' G. huttonii'' known from Jurassic fossils. However, both ''G. digitata'' and ''G. huttoni'' have leaves which are typically divided into six lobes rather than four and both of them have distinct trichomes, hairs, on the underside of the leaves.


References

{{Taxonbar, from=Q5563145 dissecta Plants described in 2002 Fossil taxa described in 2002 Ypresian plants of North America Flora of British Columbia Flora of Washington (state) Extinct flora of North America
A dagger, obelisk, or obelus is a typographical mark that usually indicates a footnote if an asterisk has already been used. The symbol is also used to indicate death (of people) or extinction (of species). It is one of the modern descendan ...
Klondike Mountain Formation Prehistoric trees Tranquille Formation