Geophagus
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''Geophagus'' is a
genus Genus (; : genera ) is a taxonomic rank above species and below family (taxonomy), family as used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In bino ...
of
cichlid Cichlids () are a large, diverse, and widespread family of percomorph fish in the family Cichlidae, order Cichliformes. At least 1,760 species have been scientifically described, making it one of the largest vertebrate families, with on ...
s that mainly live in
South America South America is a continent entirely in the Western Hemisphere and mostly in the Southern Hemisphere, with a considerably smaller portion in the Northern Hemisphere. It can also be described as the southern Subregion#Americas, subregion o ...
as far south as
Argentina Argentina, officially the Argentine Republic, is a country in the southern half of South America. It covers an area of , making it the List of South American countries by area, second-largest country in South America after Brazil, the fourt ...
and
Uruguay Uruguay, officially the Oriental Republic of Uruguay, is a country in South America. It shares borders with Argentina to its west and southwest and Brazil to its north and northeast, while bordering the Río de la Plata to the south and the A ...
, but a single species, ''G. crassilabris'' is from
Panama Panama, officially the Republic of Panama, is a country in Latin America at the southern end of Central America, bordering South America. It is bordered by Costa Rica to the west, Colombia to the southeast, the Caribbean Sea to the north, and ...
.Mattos, J.L.O., Costa, W.J.E.M. & Santos, A.C.A. (2015): ''Geophagus diamantinensis'', a new species of the ''G. brasiliensis'' species group from Chapada Diamantina, north-eastern Brazil (Cichlidae: Geophagini). ''Ichthyological Explorations of Freshwaters, 26 (3): 209-220.'' They are found in a wide range of freshwater habitats. They are part of a group popularly known as
eartheater Geophagini is a tribe of cichlids from the subfamily Cichlinae, the American cichlids. It is the sister taxon to the clade which includes the Cichlasomatini and Heroini. Fishes in the Geophagini are distributed from Panama south to Argentina, it ...
s and mostly feed by picking up mouthfuls of sediment to sift out food items such as invertebrates, plant material and
detritus In biology, detritus ( or ) is organic matter made up of the decomposition, decomposing remains of organisms and plants, and also of feces. Detritus usually hosts communities of microorganisms that colonize and decomposition, decompose (Reminera ...
. The largest species reach up to in
standard length Fish measurement is the measuring of individual fish and various parts of fish anatomy, their anatomies, for data used in many areas of ichthyology, including Taxonomy (biology), taxonomy and fishery biology. Overall length Standard length (SL) is ...
. They are mostly kept in aquariums.


As an invasive species

''Geophagus surinamensis'' was an invasive species in Malaysia, recently found in
Putrajaya Putrajaya (), officially the Federal Territory of Putrajaya (), is the administrative centre of Malaysia. The Seat of government, seat of the Government of Malaysia, federal government of Malaysia was moved in 1999 from Kuala Lumpur to Putrajay ...
, the populations are now controlled by
giant snakehead ''Channa micropeltes'', giant snakehead, giant mudfish or toman harimau, is among the largest species in the family Channidae, capable of growing to in length and a weight of . It is native to the fresh waters of Southeast Asia (South Indian pop ...
s.


Taxonomy

Some cichlids previously included in this genus have been reallocated to ''
Biotodoma ''Biotodoma'' is a small genus of cichlids native to rivers in the Amazon, Orinoco and Essequibo basins in South America South America is a continent entirely in the Western Hemisphere and mostly in the Southern Hemisphere, with a cons ...
'', ''
Gymnogeophagus ''Gymnogeophagus'' is a genus of cichlid fishes from South America, where they are known from various river basins (notably Rio de la Plata and Lagoa dos Patos– Mirim) in southern Brazil, Paraguay, Uruguay and northern Argentina. They are part ...
'' or ''
Satanoperca ''Satanoperca'' is a small genus of cichlids from South America, where they are known from the Orinoco, Essequibo, Nickerie, Amazon and Paraná–Paraguay river basins. They are mainly found in areas with slow-moving waters, but some species h ...
''. Even with these as separate genera, ''Geophagus'' is currently
polyphyletic A polyphyletic group is an assemblage that includes organisms with mixed evolutionary origin but does not include their most recent common ancestor. The term is often applied to groups that share similar features known as Homoplasy, homoplasies ...
and in need of further
taxonomic 280px, Generalized scheme of taxonomy Taxonomy is a practice and science concerned with classification or categorization. Typically, there are two parts to it: the development of an underlying scheme of classes (a taxonomy) and the allocation ...
revision. There are three main groups: * ''Geophagus
sensu stricto ''Sensu'' is a Latin word meaning "in the sense of". It is used in a number of fields including biology, geology, linguistics, semiotics, and law. Commonly it refers to how strictly or loosely an expression is used in describing any particular c ...
'' are mostly relatively peaceful, often have long fin extensions and are native to the
Amazon Amazon most often refers to: * Amazon River, in South America * Amazon rainforest, a rainforest covering most of the Amazon basin * Amazon (company), an American multinational technology company * Amazons, a tribe of female warriors in Greek myth ...
,
Orinoco The Orinoco () is one of the longest rivers in South America at . Its drainage basin, sometimes known as the Orinoquia, covers approximately 1 million km2, with 65% of it in Venezuela and 35% in Colombia. It is the List of rivers by discharge, f ...
and
Parnaíba Parnaíba () is a city in the Brazilian state of Piauí. Having a population of over 169,000 inhabitants according to IBGE's 2024 estimates, it is the second most populous city in the state, after the capital Teresina. It is one of the four coast ...
basins, as well as rivers of the
Guianas The Guianas, also spelled Guyanas or Guayanas, are a geographical region in north-eastern South America. Strictly, the term refers to the three Guianas: Guyana, Suriname, and French Guiana, formerly British Guiana, British, Surinam (Dutch colo ...
. This group can be divided into two subgroups: The first is the ''G. surinamensis''
complex Complex commonly refers to: * Complexity, the behaviour of a system whose components interact in multiple ways so possible interactions are difficult to describe ** Complex system, a system composed of many components which may interact with each ...
, which includes most species (fish in the aquarium trade often are identified as ''G. surinamensis'' itself, but they are typically other members of this complex.) The second subgroup contains ''G. argyrostictus'', ''G. gottwaldi'', ''G. grammepareius'', ''G. harreri'' and ''G. taeniopareius'', which are somewhat less peaceful and can be separated from the ''G. surinamensis'' complex by their dark stripe below the eye (however, this feature is shared with the next group). * ''G. brasiliensis'' complex (including ''G. diamantinensis'', ''G. iporangensis'', ''G. itapicuruensis'', ''G. multiocellus'', ''G. obscurus'', ''G. rufomarginatus'' and ''G. santosi'') are more robust and aggressive species found in river basins of eastern and southeastern Brazil, Uruguay and northeastern Argentina. * ''G. steindachneri'' complex (including ''G. crassilabris'' and ''G. pellegrini'' and undescribed species entering the aquarium trade from Colombia) found west of the
Andes The Andes ( ), Andes Mountains or Andean Mountain Range (; ) are the List of longest mountain chains on Earth, longest continental mountain range in the world, forming a continuous highland along the western edge of South America. The range ...
in northern and western Colombia, northwestern Venezuela and Panama where adult males develop a distinct, bulbous red forehead.


Species

There are currently 32 recognized species in this genus. * ''
Geophagus abalios ''Geophagus'' is a genus of cichlids that mainly live in South America as far south as Argentina and Uruguay, but a single species, ''G. crassilabris'' is from Panama.Mattos, J.L.O., Costa, W.J.E.M. & Santos, A.C.A. (2015): ''Geophagus diamantin ...
'' López-Fernández & Taphorn, 2004 * ''
Geophagus altifrons ''Geophagus altifrons'' are a freshwater eartheater cichlid, native to the Amazon River Basin, Brazil. It is also commonly known as the threadfin eartheater or the surinam eartheater. Introduced to Singapore and considered an invasive species, i ...
'' Heckel, 1840 * '' Geophagus argyrostictus'' S. O. Kullander, 1991 * '' Geophagus brachybranchus'' S. O. Kullander & Nijssen, 1989 * ''
Geophagus brasiliensis The pearl cichlid (''Geophagus brasiliensis'') is a species of fish in the North American cichlid family. It is native to southeastern Brazil (north to Bahia), Paraguay, Uruguay and far northeastern Argentina,Mattos, J.L.O., Costa, W.J.E.M. & Sa ...
'' ( Quoy &
Gaimard Joseph Paul Gaimard (31 January 1793 – 10 December 1858) was a French naval surgeon and naturalist. Biography Gaimard was born at Saint-Zacharie on January 31, 1793. He studied medicine at the naval medical school in Toulon, subsequent ...
, 1824)
(Pearl cichlid) * '' Geophagus brokopondo'' S. O. Kullander & Nijssen, 1989 * '' Geophagus camopiensis'' Pellegrin, 1903 (Oyapock eartheater) * '' Geophagus crassilabris''
Steindachner Franz Steindachner (11 November 1834 in Vienna – 10 December 1919 in Vienna) was an Austrian zoologist, ichthyologist, and herpetologist. He published over 200 papers on fishes and over 50 papers on reptiles and amphibians. Steindachner descri ...
, 1876
(Panamanian eartheater) * '' Geophagus crocatus''
Hauser Hauser is a German-language surname. Notable people with the surname include: * Arnold Hauser (shortstop), Arnold George Hauser (1888–1966), American baseball player * Arnold Hauser (art historian) (1892–1978), Hungarian art historian * Bodo ...
& López-Fernández, 2013
* '' Geophagus diamantinensis'' Mattos, W. J. E. M. Costa & A. C. A. Santos, 2015 * '' Geophagus dicrozoster'' López-Fernández & Taphorn, 2004 * ''
Geophagus gottwaldi ''Geophagus'' is a genus of cichlids that mainly live in South America as far south as Argentina and Uruguay, but a single species, ''G. crassilabris'' is from Panama.Mattos, J.L.O., Costa, W.J.E.M. & Santos, A.C.A. (2015): ''Geophagus diamantin ...
'' I. Schindler & Staeck, 2006 * '' Geophagus grammepareius'' S. O. Kullander & Taphorn, 1992 * ''
Geophagus harreri ''Geophagus'' is a genus of cichlids that mainly live in South America as far south as Argentina and Uruguay, but a single species, ''G. crassilabris'' is from Panama.Mattos, J.L.O., Costa, W.J.E.M. & Santos, A.C.A. (2015): ''Geophagus diamantin ...
'' J. P. Gosse, 1976 (Maroni eartheater) * '' Geophagus iporangensis'' Haseman, 1911 * '' Geophagus itapicuruensis'' Haseman, 1911 * '' Geophagus megasema'' Heckel, 1840 * '' Geophagus mirabilis'' Deprá, S. O. Kullander, Pavanelli & da Graça, 2014 * '' Geophagus multiocellus'' Mattos & W. J. E. M. Costa, 2018 * '' Geophagus neambi'' P. H. L. Lucinda, C. A. S. de Lucena &
Assis Assis is a city and a municipality in the southwestern part of the state of São Paulo in Brazil. The population is 101,409 (2022 est.) in an area of 460.61 km. The town was founded on July 5, 1905, and became a municipality in 1917, when it ...
, 2010
* '' Geophagus obscurus'' ( Castelnau, 1855) * '' Geophagus parnaibae'' Staeck & I. Schindler, 2006 * '' Geophagus pellegrini'' Regan, 1912 (Yellowhump eartheater) * '' Geophagus proximus'' ( Castelnau, 1855) * ''
Geophagus pyrocephalus ''Geophagus'' is a genus of cichlids that mainly live in South America as far south as Argentina and Uruguay, but a single species, ''G. crassilabris'' is from Panama.Mattos, J.L.O., Costa, W.J.E.M. & Santos, A.C.A. (2015): ''Geophagus diamanti ...
'' Chuctaya et al., 2022 * '' Geophagus rufomarginatus'' Mattos & W. J. E. M. Costa, 2018 * '' Geophagus santosi'' Mattos & W. J. E. M. Costa, 2018 * ''
Geophagus steindachneri The redhump eartheater (''Geophagus steindachneri'') is a species of eartheater cichlid from freshwater habitats in northwestern South America.Geophagus steindachneri'. FishBase. Agustin, L., Kullander, S., & Torres, A. (2001-2008). Distributi ...
'' C. H. Eigenmann &
Hildebrand Hildebrand is a character from Germanic heroic legend. ''Hildebrand'' is the modern German form of the name: in Old High German it is ''Hiltibrant'' and in Old Norse ''Hildibrandr''. The word ''hild'' means "battle" and ''brand'' means "sword" ...
, 1922
(Redhump eartheater) * '' Geophagus surinamensis'' (
Bloch Bloch is a surname of German origin. Notable people with this surname include: A *Adele Bloch-Bauer (1881–1925), Austrian entrepreneur *Albert Bloch (1882–1961), American painter *Alexandre Bloch (1857–1919), French painter *Alfred Bloch ( ...
, 1791)
(Red-striped eartheater) * '' Geophagus sveni'' P. H. F. Lucinda, C. A. S. de Lucena &
Assis Assis is a city and a municipality in the southwestern part of the state of São Paulo in Brazil. The population is 101,409 (2022 est.) in an area of 460.61 km. The town was founded on July 5, 1905, and became a municipality in 1917, when it ...
, 2010
* '' Geophagus taeniopareius'' S. O. Kullander & Royero-L., 1992 * '' Geophagus winemilleri'' López-Fernández & Taphorn, 2004


References

{{Authority control Cichlid genera Taxa named by Johann Jakob Heckel