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''Geastrum'' (
orthographical variant In biology, within the science of scientific nomenclature, i.e. the naming of organisms, an orthographical variant (abbreviated orth. var.) in botany or an orthographic error in zoology, is a spelling mistake, typing mistake or writing mistake ...
''Geaster'') is a
genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of living and fossil organisms as well as viruses. In the hierarchy of biological classification, genus comes above species and below family. In binomial n ...
of puffball-like
mushroom A mushroom or toadstool is the fleshy, spore-bearing fruiting body of a fungus, typically produced above ground, on soil, or on its food source. ''Toadstool'' generally denotes one poisonous to humans. The standard for the name "mushroom" is ...
s in the family
Geastraceae Geastrales is an order of gasterocarpic basidiomycetes (fungi) that are related to Cantharellales. The order contains the single family Geastraceae, commonly known as "earthstars", which older classifications had placed in Lycoperdales, or ...
. Many species are known commonly as earthstars. The name, which comes from ''geo'' meaning ''earth'' and meaning ''star'', refers to the behavior of the outer peridium. At maturity, the outer layer of the fruiting body splits into segments which turn outward creating a star-like pattern on the ground. The inner peridium is called a spore sac. In some species, the outer peridium splits from a middle layer, causing the spore sac to arch off the ground. If the outer peridium opens when wet and closes when dry, it is described as hygroscopic. In some species, the inner peridium is borne on a stalk or pedicel. The columella is a column-like clump of sterile tissue found inside the inner peridium. The network of fertile tissue inside the inner peridium, the
capillitium Capillitium (pl. capillitia) is a mass of sterile fibers within a fruit body interspersed among spores. It is found in Mycetozoa (slime molds) and gasteroid fungi of the fungal subdivision Agaricomycotina The subdivision Agaricomycotina, also ...
, arises from the columella And is where
basidia A basidium () is a microscopic sporangium (a spore-producing structure) found on the hymenophore of fruiting bodies of basidiomycete fungi which are also called tertiary mycelium, developed from secondary mycelium. Tertiary mycelium is highly- ...
and
basidiospores A basidiospore is a reproductive spore produced by Basidiomycete fungi, a grouping that includes mushrooms, shelf fungi, rusts, and smuts. Basidiospores typically each contain one haploid nucleus that is the product of meiosis, and they are ...
are produced. The mouth in most species of "earth-stars" is quite prominent, often arising as a small cone at the apex of the inner peridium. It may be even or sulcate (grooved). They are generally not toxic but considered non-edible due to their fibrous texture in the mature stage at which they are generally found.


Species

Although the ''Dictionary of the Fungi'' (2008) estimated roughly 50 species in ''Geastrum'', a more recent estimate (2014) suggests that there may be up to 120 species. ''Geastrum coronatum'' Pers. has been proposed as the conserved type for the genus. Some similar species that are otherwise difficult to differentiate using classical morphological features (such as ''G. triplex'', ''G. saccatum'', and ''G. lageniforme'') can be identified using chemical spot tests that detect
phenoloxidase Polyphenol oxidase (PPO; also polyphenol oxidase i, chloroplastic), an enzyme involved in Browning (food process), fruit browning, is a tetramer that contains four atoms of copper per molecule. PPO may accept monophenol , monophenols and/or ...
enzymatic activity, a differences in the crystal structure of
calcium oxalate Calcium oxalate (in archaic terminology, oxalate of lime) is a calcium salt of oxalic acid with the chemical formula . It forms hydrates , where ''n'' varies from 1 to 3. Anhydrous and all hydrated forms are colorless or white. The monohydrate ...
deposits or DNA sequences. Species include: * ''
Geastrum aculeatum ''Geastrum aculeatum'' is a species of mushroom belonging to the genus '' Geastrum'', or earthstar fungi. It is found in eastern Brazil. References aculeatum Fungi described in 2013 {{geastrales-stub ...
'' * ''
Geastrum albonigrum ''Geastrum albonigrum'' is an inedible species of mushroom belonging to the genus '' Geastrum'', or earthstar genus. It is found in Costa Rica and Mexico. ''G. albonigrum'' can be distinguished from superficially similar species '' G. coronatum'' ...
'' * '' Geastrum andrewsii'' * '' Geastrum argentatum'' * '' Geastrum argentinum'' * '' Geastrum australe'' * '' Geastrum austrominimum'' * '' Geastrum benitoi'' * '' Geastrum britannicum'' * '' Geastrum berkeleyi'' * '' Geastrum bushnellii'' * '' Geastrum campestre'' * '' Geastrum clelandii'' * '' Geastrum congolense'' * '' Geastrum corollinum'' * '' Geastrum coronatum'' * '' Geastrum dissimile'' * '' Geastrum drummondii'' * ''
Geastrum dubowskii ''Geastrum'' (orthographical variant ''Geaster'') is a genus of puffball-like mushrooms in the family Geastraceae. Many species are known commonly as earthstars. The name, which comes from ''geo'' meaning ''earth'' and meaning ''star'', refers ...
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Geastrum echinulatum ''Geastrum'' (orthographical variant ''Geaster'') is a genus of puffball-like mushrooms in the family Geastraceae. Many species are known commonly as earthstars. The name, which comes from ''geo'' meaning ''earth'' and meaning ''star'', refers ...
'' * ''
Geastrum elegans ''Geastrum elegans'' is an inedible species of mushroom A mushroom or toadstool is the fleshy, spore-bearing fruiting body of a fungus, typically produced above ground, on soil, or on its food source. ''Toadstool'' generally denotes one pois ...
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Geastrum elliptice ''Geastrum'' (orthographical variant ''Geaster'') is a genus of puffball-like mushrooms in the family Geastraceae. Many species are known commonly as earthstars. The name, which comes from ''geo'' meaning ''earth'' and meaning ''star'', refers ...
'' * ''
Geastrum entomophilum ''Geastrum'' (orthographical variant ''Geaster'') is a genus of puffball-like mushrooms in the family Geastraceae. Many species are known commonly as earthstars. The name, which comes from ''geo'' meaning ''earth'' and meaning ''star'', refers ...
'' * '' Geastrum episcopale'' * ''
Geastrum fimbriatum ''Geastrum fimbriatum'', commonly known as the fringed earthstar or the sessile earthstar, is an inedible species of mushroom belonging to the genus ''Geastrum'', or earthstar fungi. First described in 1829, the species has a widespread distributi ...
'' * '' Geastrum flexuosum'' * ''
Geastrum floriforme ''Geastrum'' (orthographical variant ''Geaster'') is a genus of puffball-like mushrooms in the family Geastraceae. Many species are known commonly as earthstars. The name, which comes from ''geo'' meaning ''earth'' and meaning ''star'', refers ...
'' * '' Geastrum fornicatum'' * '' Geastrum fuscogleba'' * ''
Geastrum hieronymi ''Geastrum'' (orthographical variant ''Geaster'') is a genus of puffball-like mushrooms in the family Geastraceae. Many species are known commonly as earthstars. The name, which comes from ''geo'' meaning ''earth'' and meaning ''star'', refers ...
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Geastrum hirsutum ''Geastrum'' (orthographical variant ''Geaster'') is a genus of puffball-like mushrooms in the family Geastraceae. Many species are known commonly as earthstars. The name, which comes from ''geo'' meaning ''earth'' and meaning ''star'', refers ...
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Geastrum huneckii ''Geastrum huneckii'' is a species of earthstar fungus in the family Geastraceae. Found in Mongolia, it was formally described as a new species in 1981 by German mycologist Heinrich Dörfelt. The species epithet honours his colleague Siegfried ...
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Geastrum hungaricum ''Geastrum'' (orthographical variant ''Geaster'') is a genus of puffball-like mushrooms in the family Geastraceae. Many species are known commonly as earthstars. The name, which comes from ''geo'' meaning ''earth'' and meaning ''star'', refe ...
'' * '' Geastrum inpaense'' – Brazil * ''
Geastrum kotlabae ''Geastrum'' (orthographical variant ''Geaster'') is a genus of puffball-like mushrooms in the family Geastraceae. Many species are known commonly as earthstars. The name, which comes from ''geo'' meaning ''earth'' and meaning ''star'', refers ...
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Geastrum kuharii ''Geastrum'' (orthographical variant ''Geaster'') is a genus of puffball-like mushrooms in the family Geastraceae. Many species are known commonly as earthstars. The name, which comes from ''geo'' meaning ''earth'' and meaning ''star'', refers ...
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Geastrum lageniforme ''Geastrum lageniforme '' is a species of fungus A fungus ( : fungi or funguses) is any member of the group of eukaryotic organisms that includes microorganisms such as yeasts and molds, as well as the more familiar mushrooms. These organ ...
'' * '' Geastrum leptospermum'' * '' Geastrum lilloi'' * '' Geastrum litchiforme'' * '' Geastrum lloydianum'' * ''
Geastrum magnosporum ''Geastrum'' (orthographical variant ''Geaster'') is a genus of puffball-like mushrooms in the family Geastraceae. Many species are known commonly as earthstars. The name, which comes from ''geo'' meaning ''earth'' and meaning ''star'', refers ...
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Geastrum melanocephalum ''Geastrum melanocephalum'' is a species of fungus A fungus ( : fungi or funguses) is any member of the group of eukaryotic organisms that includes microorganisms such as yeasts and molds, as well as the more familiar mushrooms. These org ...
'' * '' Geastrum meridionale'' * '' Geastrum minimum'' * ''
Geastrum mirabile ''Geastrum'' (orthographical variant ''Geaster'') is a genus of puffball-like mushrooms in the family Geastraceae. Many species are known commonly as earthstars. The name, which comes from ''geo'' meaning ''earth'' and meaning ''star'', refe ...
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Geastrum morganii ''Geastrum'' (orthographical variant ''Geaster'') is a genus of puffball-like mushrooms in the family Geastraceae. Many species are known commonly as earthstars. The name, which comes from ''geo'' meaning ''earth'' and meaning ''star'', refers ...
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Geastrum ovalisporum ''Geastrum'' (orthographical variant ''Geaster'') is a genus of puffball-like mushrooms in the family Geastraceae. Many species are known commonly as earthstars. The name, which comes from ''geo'' meaning ''earth'' and meaning ''star'', refers ...
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Geastrum oxylobum ''Geastrum'' (orthographical variant ''Geaster'') is a genus of puffball-like mushrooms in the family Geastraceae. Many species are known commonly as earthstars. The name, which comes from ''geo'' meaning ''earth'' and meaning ''star'', refers ...
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Geastrum papinuttii ''Geastrum'' (orthographical variant ''Geaster'') is a genus of puffball-like mushrooms in the family Geastraceae. Many species are known commonly as earthstars. The name, which comes from ''geo'' meaning ''earth'' and meaning ''star'', refers ...
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Geastrum parvisporum ''Geastrum'' (orthographical variant ''Geaster'') is a genus of puffball-like mushrooms in the family Geastraceae. Many species are known commonly as earthstars. The name, which comes from ''geo'' meaning ''earth'' and meaning ''star'', refers ...
'' * '' Geastrum parvistriatum'' * ''
Geastrum pectinatum ''Geastrum pectinatum'' is an inedible species of mushroom belonging to the Geastraceae, earthstar family of fungi. Although young specimens are spherical, basidiocarp, fruit body development involves the outer layer of tissue splitting open l ...
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Geastrum pleosporus ''Geastrum'' (orthographical variant ''Geaster'') is a genus of puffball-like mushrooms in the family Geastraceae. Many species are known commonly as earthstars. The name, which comes from ''geo'' meaning ''earth'' and meaning ''star'', refers ...
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Geastrum pouzarii ''Geastrum'' (orthographical variant ''Geaster'') is a genus of puffball-like mushrooms in the family Geastraceae. Many species are known commonly as earthstars. The name, which comes from ''geo'' meaning ''earth'' and meaning ''star'', refers ...
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Geastrum pseudolimbatum ''Geastrum'' (orthographical variant ''Geaster'') is a genus of puffball-like mushrooms in the family Geastraceae. Many species are known commonly as earthstars. The name, which comes from ''geo'' meaning ''earth'' and meaning ''star'', refers ...
'' * '' Geastrum quadrifidum'' * '' Geastrum reticulatum'' * ''
Geastrum rhizophorum ''Geastrum'' (orthographical variant ''Geaster'') is a genus of puffball-like mushrooms in the family Geastraceae. Many species are known commonly as earthstars. The name, which comes from ''geo'' meaning ''earth'' and meaning ''star'', refe ...
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Geastrum rufescens ''Geastrum rufescens'', commonly known as the rosy earthstar, is a species of fungus in the family Geastraceae. It was first described scientifically by Christian Hendrik Persoon in 1801. It has a pale pinkish-buff to pinkish exoperidium and ra ...
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Geastrum rugulosum ''Geastrum'' (orthographical variant ''Geaster'') is a genus of puffball-like mushrooms in the family Geastraceae. Many species are known commonly as earthstars. The name, which comes from ''geo'' meaning ''earth'' and meaning ''star'', refers ...
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Geastrum rusticum ''Geastrum'' (orthographical variant ''Geaster'') is a genus of puffball-like mushrooms in the family Geastraceae. Many species are known commonly as earthstars. The name, which comes from ''geo'' meaning ''earth'' and meaning ''star'', refers ...
'' – Brazil * ''
Geastrum saccatum ''Geastrum saccatum'', commonly known as the rounded earthstar, is a species of mushroom belonging to the genus ''Geastrum''. It has a worldwide distribution and is found growing on rotting wood. It is considered inedible by mushroomers because ...
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Geastrum schmidelii ''Geastrum'' (orthographical variant ''Geaster'') is a genus of puffball-like mushrooms in the family Geastraceae. Many species are known commonly as earthstars. The name, which comes from ''geo'' meaning ''earth'' and meaning ''star'', refers ...
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Geastrum schweinitzii ''Geastrum'' (orthographical variant ''Geaster'') is a genus of puffball-like mushrooms in the family Geastraceae. Many species are known commonly as earthstars. The name, which comes from ''geo'' meaning ''earth'' and meaning ''star'', refers ...
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Geastrum senoretiae ''Geastrum'' (orthographical variant ''Geaster'') is a genus of puffball-like mushrooms in the family Geastraceae. Many species are known commonly as earthstars. The name, which comes from ''geo'' meaning ''earth'' and meaning ''star'', refers ...
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Geastrum setiferum ''Geastrum'' (orthographical variant ''Geaster'') is a genus of puffball-like mushrooms in the family Geastraceae. Many species are known commonly as earthstars. The name, which comes from ''geo'' meaning ''earth'' and meaning ''star'', refers ...
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Geastrum smardae ''Geastrum'' (orthographical variant ''Geaster'') is a genus of puffball-like mushrooms in the family Geastraceae. Many species are known commonly as earthstars. The name, which comes from ''geo'' meaning ''earth'' and meaning ''star'', refers ...
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Geastrum smithii ''Geastrum'' (orthographical variant ''Geaster'') is a genus of puffball-like mushrooms in the family Geastraceae. Many species are known commonly as earthstars. The name, which comes from ''geo'' meaning ''earth'' and meaning ''star'', refers ...
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Geastrum stiptatum ''Geastrum'' (orthographical variant ''Geaster'') is a genus of puffball-like mushrooms in the family Geastraceae. Many species are known commonly as earthstars. The name, which comes from ''geo'' meaning ''earth'' and meaning ''star'', refers ...
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Geastrum striatum ''Geastrum'' (orthographical variant ''Geaster'') is a genus of puffball-like mushrooms in the family Geastraceae. Many species are known commonly as earthstars. The name, which comes from ''geo'' meaning ''earth'' and meaning ''star'', refers ...
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Geastrum subiculosum ''Geastrum subiculosum'' is an inedible species of fungus belonging to the genus '' Geastrum'', or earthstar fungi. The fungus was first described scientifically by Cooke and Massee in 1887 (as ''Geaster subiculosus''; ''Geaster'' is an older nam ...
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Geastrum tichifer ''Geastrum'' (orthographical variant ''Geaster'') is a genus of puffball-like mushrooms in the family Geastraceae. Many species are known commonly as earthstars. The name, which comes from ''geo'' meaning ''earth'' and meaning ''star'', refers ...
'' * '' Geastrum triplex'' * ''
Geastrum velutinum ''Geastrum velutinum'' is a species of fungus in the family Geastraceae. Found in North America, it was first species description, described scientifically by Andrew Price Morgan in 1895. References External links

* Geastrum, velutinum ...
'' * '' Geastrum verrucoramulosum'' * ''
Geastrum welwitschii ''Geastrum welwitschii'' is a species of fungus in the earthstar family. When young and unopened, the fruit bodies resemble small spheres lying in the soil. As the mushroom matures, the thick leathery outer layer of tissue (the peridium) spli ...
'' * '' Geastrum xerophilum'' * '' Geastrum xylogenum''


References


Sources


New Zealand Species

British Species

Czech Species and Photos
* *
Ponce de Leon, Patricio, A Revision of the Family Geastraceae, Field Museum of Natural History 1968
* * {{Taxonbar, from=Q687050 Agaricomycetes genera