Gasosaurus Constructus Scale Diagram
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''Gasosaurus'' () is a
genus Genus (; : genera ) is a taxonomic rank above species and below family (taxonomy), family as used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In bino ...
of
tetanuran Tetanurae (/ˌtɛtəˈnjuːriː/ or "stiff tails") is a clade that includes most theropod dinosaurs, including megalosauroids, allosauroids, and coelurosaurs (which includes tyrannosauroids, ornithomimosaurs, compsognathids and maniraptoran ...
theropod Theropoda (; from ancient Greek , (''therion'') "wild beast"; , (''pous, podos'') "foot"">wiktionary:ποδός"> (''pous, podos'') "foot" is one of the three major groups (clades) of dinosaurs, alongside Ornithischia and Sauropodom ...
that lived approximately 171.6 to 161.2 
million years ago Million years ago, abbreviated as Mya, Myr (megayear) or Ma (megaannum), is a unit of time equal to (i.e. years), or approximately 31.6 teraseconds. Usage Myr is in common use in fields such as Earth science and cosmology. Myr is also used w ...
during the middle of the
Jurassic The Jurassic ( ) is a Geological period, geologic period and System (stratigraphy), stratigraphic system that spanned from the end of the Triassic Period million years ago (Mya) to the beginning of the Cretaceous Period, approximately 143.1 Mya. ...
Period Period may refer to: Common uses * Period (punctuation) * Era, a length or span of time *Menstruation, commonly referred to as a "period" Arts, entertainment, and media * Period (music), a concept in musical composition * Periodic sentence (o ...
. The name "''Gasosaurus''" is derived from the English "''
gasoline Gasoline ( North American English) or petrol ( Commonwealth English) is a petrochemical product characterized as a transparent, yellowish, and flammable liquid normally used as a fuel for spark-ignited internal combustion engines. When for ...
''" and the
Greek Greek may refer to: Anything of, from, or related to Greece, a country in Southern Europe: *Greeks, an ethnic group *Greek language, a branch of the Indo-European language family **Proto-Greek language, the assumed last common ancestor of all kno ...
() ("lizard / generic reptile"). Only one species is currently recognised, ''G. constructus'', from which the specific name honours the
gasoline Gasoline ( North American English) or petrol ( Commonwealth English) is a petrochemical product characterized as a transparent, yellowish, and flammable liquid normally used as a fuel for spark-ignited internal combustion engines. When for ...
company that found the Dashanpu fossil quarry in
Sichuan Sichuan is a province in Southwestern China, occupying the Sichuan Basin and Tibetan Plateau—between the Jinsha River to the west, the Daba Mountains to the north, and the Yunnan–Guizhou Plateau to the south. Its capital city is Cheng ...
Province,
China China, officially the People's Republic of China (PRC), is a country in East Asia. With population of China, a population exceeding 1.4 billion, it is the list of countries by population (United Nations), second-most populous country after ...
, now named as the
Lower Shaximiao Formation The Shaximiao Formation () is a Middle to Late Jurassic aged geological formation in Sichuan, China, most notable for the wealth of dinosaurs fossils that have been excavated from its strata. The Shaximiao Formation is exposed in and around the ...
.


Discovery and naming

The first and to date only
fossil A fossil (from Classical Latin , ) is any preserved remains, impression, or trace of any once-living thing from a past geological age. Examples include bones, shells, exoskeletons, stone imprints of animals or microbes, objects preserve ...
s, albeit
postcranial The postcranium ("behind the cranium"; plural: postcrania) or postcranial skeleton in zoology and vertebrate paleontology is the skeleton apart from the skull. The postcranium encompasses the axial skeleton, which includes the entirety of the verte ...
(missing the
skull The skull, or cranium, is typically a bony enclosure around the brain of a vertebrate. In some fish, and amphibians, the skull is of cartilage. The skull is at the head end of the vertebrate. In the human, the skull comprises two prominent ...
), were recovered in 1985 during the construction of a gas facility, which explains the dinosaur's unusual name. It consists of laterally compressed teeth, 4 cervical, 7 dorsal, 5 sacral, and 7 caudal vertebrae, both humeri, and highly abraded and reconstructed pelvic and hind limb material that includes the left ilium, left ischium, left pubis, left femur, left tibia and left fibula. The fossils were defined as the
type species In International_Code_of_Zoological_Nomenclature, zoological nomenclature, a type species (''species typica'') is the species name with which the name of a genus or subgenus is considered to be permanently taxonomically associated, i.e., the spe ...
''Gasosaurus constructus'' by the
paleontologist Paleontology, also spelled as palaeontology or palæontology, is the scientific study of the life of the past, mainly but not exclusively through the study of fossils. Paleontologists use fossils as a means to classify organisms, measure geolo ...
s
Dong Zhiming Dong Zhiming (Chinese language, Chinese: 董枝明, Pinyin: ''Dǒng Zhimíng''; January 1937 – 20 October 2024) was a Chinese vertebrate paleontologist formerly employed at the Institute of Vertebrate Paleontology and Paleoanthropology (IVPP) ...
and Tang Zilu.


Description

''Gasosaurus'' is a
carnivorous A carnivore , or meat-eater (Latin, ''caro'', genitive ''carnis'', meaning meat or "flesh" and ''vorare'' meaning "to devour"), is an animal or plant whose nutrition and energy requirements are met by consumption of animal tissues (mainly mu ...
theropod Theropoda (; from ancient Greek , (''therion'') "wild beast"; , (''pous, podos'') "foot"">wiktionary:ποδός"> (''pous, podos'') "foot" is one of the three major groups (clades) of dinosaurs, alongside Ornithischia and Sauropodom ...
with strong legs but short arms. It measured between in length, with a weight of around . However, some estimates put its weight as high as , as very little is known about it. Many potentially informative features of the holotype skeleton (IVPP V7264) are difficult to assess based on published descriptions and images. The taxon represents a tetanuran based on the presence of a pubic peduncle of the ilium that is substantially larger than the ischial peduncle. Examination of casts reveals that the lesser trochanter does not rise above the level of the femoral head, and instead, the proximal portion of the femoral head is broken and the lesser trochanter reaches approximately midlevel of the head as in non-coelurosaurian tetanurans. Furthermore, it is recognized by an amphiplatyan cervical centra with incipient weak ventral keel, dorsals with amphiplatyan centra, low neural arches and plate-like neural spines, dorsal neural spines lacking expanded bulks at tips, five sacral centra and arches firmly fused while neural spines not fused, presence of a humeral foramen, low ilium with a less developed anterior process, and distal ends of pubis and ischium expanded but lacking foot-like processes.


Classification

Traditionally thought to be a
megalosauroid Megalosauroidea (meaning 'great/big lizard forms') is a Taxonomic rank, superfamily (or clade) of Tetanurae, tetanuran theropod dinosaurs that lived from the Middle Jurassic to the Late Cretaceous period. The group is defined as ''Megalosaurus, M ...
, Holtz (2000) found it to be a basal
coelurosauria Coelurosauria (; from Greek, meaning "hollow-tailed lizards") is the clade containing all theropod dinosaurs more closely related to birds than to carnosaurs. Coelurosauria is a subgroup of theropod dinosaurs that includes compsognathids, tyra ...
n, although later Holtz ''et al.'' (2004) suggested it was a basal
carnosaur Carnosauria is an extinct group of carnivorous theropod dinosaurs that lived during the Jurassic and Cretaceous periods. While Carnosauria was historically considered largely synonymous with Allosauroidea, some recent studies have revived Carn ...
(possibly a
metriacanthosaurid Metriacanthosauridae (Greek for "moderately-spined lizards") is an extinct family of allosauroid theropod dinosaurs that lived in Europe and Asia from the Middle Jurassic to the Early Cretaceous. The family is split into two subgroups: Metriacant ...
) on the basis of data from undescribed specimens.Holtz TR Jr, Molnar RE, Currie PJ. 2004. Basal Tetanurae. In: Weishampel DB, Dodson P, Osmólska H, eds. The dinosauria, 2nd edn. Berkeley, CA: University of California Press, 71–110. It may in fact be the most basal coelurosaurian yet known, or may even be close to the
common ancestor Common descent is a concept in evolutionary biology applicable when one species is the ancestor of two or more species later in time. According to modern evolutionary biology, all living beings could be descendants of a unique ancestor commonl ...
of the two groups; in any case, it represents one of the oldest definitive
tetanuran Tetanurae (/ˌtɛtəˈnjuːriː/ or "stiff tails") is a clade that includes most theropod dinosaurs, including megalosauroids, allosauroids, and coelurosaurs (which includes tyrannosauroids, ornithomimosaurs, compsognathids and maniraptoran ...
theropods. Some paleontologists have speculated that ''Gasosaurus'' and ''
Kaijiangosaurus ''Kaijiangosaurus'' (meaning "Kaijiang lizard") is a genus of carnivorous tetanuran theropod dinosaur from the Middle Jurassic of China. In 1984 He Xinlu named and described the type species ''Kaijiangosaurus lini''. The generic name refers to t ...
'' may be one and the same species. Because of the fragmentary nature of the known ''Gasosaurus'' fossils, it has an uncertain position within
Tetanurae Tetanurae (/ˌtɛtəˈnjuːriː/ or "stiff tails") is a clade that includes most Theropoda, theropod dinosaurs, including Megalosauroidea, megalosauroids, Allosauroidea, allosauroids, and Coelurosauria, coelurosaurs (which includes Tyrannosauroi ...
, but probably lays outside
Coelurosauria Coelurosauria (; from Greek, meaning "hollow-tailed lizards") is the clade containing all theropod dinosaurs more closely related to birds than to carnosaurs. Coelurosauria is a subgroup of theropod dinosaurs that includes compsognathids, tyra ...
. A detailed restudy of the holotype is underway that may change its classification.


Paleoecology

''Gasosarurus'' lived during the mid-
Jurassic The Jurassic ( ) is a Geological period, geologic period and System (stratigraphy), stratigraphic system that spanned from the end of the Triassic Period million years ago (Mya) to the beginning of the Cretaceous Period, approximately 143.1 Mya. ...
period (
Bathonian In the geologic timescale the Bathonian is an age (geology), age and stage (stratigraphy), stage of the Middle Jurassic. It lasted from approximately 168.2 ±1.2 annum, Ma to around 165.3 ±1.1 Ma (million years ago). The Bathonian Age succeeds ...
and/ or
Callovian In the geologic timescale, the Callovian is an age and stage in the Middle Jurassic, lasting between 165.3 ± 1.1 Ma (million years ago) and 161.5 ± 1.0 Ma. It is the last stage of the Middle Jurassic, following the Bathonian and preceding the ...
stages), around 164 million years ago. It coexisted with other mid-range theropods such as ''
Chuandongocoelurus ''Chuandongocoelurus'' ( ) is a genus of carnivorous tetanuran theropod dinosaur from the Jurassic of China. Discovery and naming The type species ''Chuandongocoelurus primitivus'' was first described and named by Chinese paleontologist He ...
'' and ''
Kaijiangosaurus ''Kaijiangosaurus'' (meaning "Kaijiang lizard") is a genus of carnivorous tetanuran theropod dinosaur from the Middle Jurassic of China. In 1984 He Xinlu named and described the type species ''Kaijiangosaurus lini''. The generic name refers to t ...
'', two other dinosaurs found from the
Lower Shaximiao Formation The Shaximiao Formation () is a Middle to Late Jurassic aged geological formation in Sichuan, China, most notable for the wealth of dinosaurs fossils that have been excavated from its strata. The Shaximiao Formation is exposed in and around the ...
. These three taxa pertain to medium to large bodied theropods and are known from associated prostcranial skeletons. China, boasts the highest taxic diversity of Middle Jurassic theropods of all body sizes from anywhere in the world. Most Middle
Jurassic The Jurassic ( ) is a Geological period, geologic period and System (stratigraphy), stratigraphic system that spanned from the end of the Triassic Period million years ago (Mya) to the beginning of the Cretaceous Period, approximately 143.1 Mya. ...
theropods from China are 'medium-sized', as are the majority of Middle
Jurassic The Jurassic ( ) is a Geological period, geologic period and System (stratigraphy), stratigraphic system that spanned from the end of the Triassic Period million years ago (Mya) to the beginning of the Cretaceous Period, approximately 143.1 Mya. ...
theropod Theropoda (; from ancient Greek , (''therion'') "wild beast"; , (''pous, podos'') "foot"">wiktionary:ποδός"> (''pous, podos'') "foot" is one of the three major groups (clades) of dinosaurs, alongside Ornithischia and Sauropodom ...
s globally. These similar dinosaurs show an intriguing combination of derived tetanuran synapomorphies and primitive features shared with non-tetanurans, which suggest they occupy a basal position within
Tetanurae Tetanurae (/ˌtɛtəˈnjuːriː/ or "stiff tails") is a clade that includes most Theropoda, theropod dinosaurs, including Megalosauroidea, megalosauroids, Allosauroidea, allosauroids, and Coelurosauria, coelurosaurs (which includes Tyrannosauroi ...
. Understanding the anatomy of Chinese Jurassic taxa may reveal further details of primitive tetanuran anatomy and help resolve the early evolution of this successful theropod clade. Future exploration of the Chinese Middle Jurassic theropod record may bring new insights into the diversification of Tetanurae, the origin of larger body sizes among more derived theropods, and Middle Jurassic paleobiogeography.


References


External links

{{Taxonbar, from=Q131771 Carnosauria Dinosaur genera Callovian dinosaurs Shaximiao Formation Fossil taxa described in 1985 Taxa named by Dong Zhiming Dinosaurs of China