Gallirallus Australis
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The weka, also known as the Māori hen or woodhen (''Gallirallus australis'') is a
flightless bird Flightless birds are birds that cannot Bird flight, fly, as they have, through evolution, lost the ability to. There are over 60 extant species, including the well-known ratites (ostriches, emus, cassowary, cassowaries, Rhea (bird), rheas, an ...
species of the
rail Rail or rails may refer to: Rail transport *Rail transport and related matters *Railway track or railway lines, the running surface of a railway Arts and media Film * ''Rails'' (film), a 1929 Italian film by Mario Camerini * ''Rail'' (1967 fil ...
family Family (from ) is a Social group, group of people related either by consanguinity (by recognized birth) or Affinity (law), affinity (by marriage or other relationship). It forms the basis for social order. Ideally, families offer predictabili ...
. It is
endemic Endemism is the state of a species being found only in a single defined geographic location, such as an island, state, nation, country or other defined zone; organisms that are indigenous to a place are not endemic to it if they are also foun ...
to
New Zealand New Zealand () is an island country in the southwestern Pacific Ocean. It consists of two main landmasses—the North Island () and the South Island ()—and List of islands of New Zealand, over 600 smaller islands. It is the List of isla ...
. Some authorities consider it as the only extant member of the genus ''
Gallirallus ''Gallirallus'' is a genus of Rallidae, rails that live in the Australasian-Pacific region. The genus is characterised by an ability to colonise relatively small and isolated islands and thereafter to evolve flightless forms, many of which beca ...
''. Four
subspecies In Taxonomy (biology), biological classification, subspecies (: subspecies) is a rank below species, used for populations that live in different areas and vary in size, shape, or other physical characteristics (Morphology (biology), morpholog ...
are recognized but only two (northern/southern) are supported by genetic evidence. The weka are sturdy brown birds about the size of a chicken. As
omnivore An omnivore () is an animal that regularly consumes significant quantities of both plant and animal matter. Obtaining energy and nutrients from plant and animal matter, omnivores digest carbohydrates, protein, fat, and fiber, and metabolize t ...
s, they feed mainly on
invertebrate Invertebrates are animals that neither develop nor retain a vertebral column (commonly known as a ''spine'' or ''backbone''), which evolved from the notochord. It is a paraphyletic grouping including all animals excluding the chordata, chordate s ...
s and fruit. Weka usually lay
eggs An egg is an organic vessel in which an embryo begins to develop. Egg, EGG or eggs may also refer to: Biology * Egg cell, the female reproductive cell (gamete) in oogamous organisms Food * Eggs as food Places * Egg, Austria * Egg, Switzerland ...
between August and January; both sexes help to incubate.


Description

Weka are large rails. They are predominantly rich brown mottled with black and grey; the brown shade varies from pale to dark depending on subspecies. The male is the larger sex at in length and in weight. Females measure in length and weigh . The reduced wingspan ranges from . The relatively large, reddish-brown beak is about long, stout and tapered, and used as a weapon. The pointed tail is near-constantly being flicked, a sign of unease characteristic of the rail family. Weka have sturdy legs and reduced wings.


Taxonomy and distribution

The common name, "weka" is a
Māori Māori or Maori can refer to: Relating to the Māori people * Māori people of New Zealand, or members of that group * Māori language, the language of the Māori people of New Zealand * Māori culture * Cook Islanders, the Māori people of the Co ...
word. The species was named ''Rallus australis'' by
Anders Erikson Sparrman Anders Sparrman (27 February 1748 – 9 August 1820) was a Swedish Natural history, naturalist, abolitionism, abolitionist and an Apostles of Linnaeus, apostle of Carl Linnaeus. Biography file:Anders Sparrman miniatyr.jpg, left, Portrait miniat ...
in 1789. Sparrman published the information in ''Museum Carlsonianum'', four fascicules based on specimens collected while voyaging with
Captain James Cook Captain James Cook (7 November 1728 – 14 February 1779) was a British Royal Navy officer, explorer, and cartographer famous for his three voyages of exploration to the Pacific and Southern Oceans, conducted between 1768 and 1779. He complet ...
between 1772 and 1775. ''Australis'' is Latin for "southern".
Johann Georg Wagler Johann Georg Wagler (28 March 1800 – 23 August 1832) was a German herpetologist and ornithologist. Wagler was assistant to Johann Baptist von Spix, and gave lectures in zoology at the Ludwig Maximilian University of Munich after it was moved t ...
's suggestion of the genus ''Ocydromus'' in 1830 to describe each weka as a species was generally adopted. However, weka were later classified as a single species in the genus ''
Gallirallus ''Gallirallus'' is a genus of Rallidae, rails that live in the Australasian-Pacific region. The genus is characterised by an ability to colonise relatively small and isolated islands and thereafter to evolve flightless forms, many of which beca ...
'' with four subspecies. The buff weka (''Gallirallus australis hectori'') formerly inhabited the eastern districts of the
South Island The South Island ( , 'the waters of Pounamu, Greenstone') is the largest of the three major islands of New Zealand by surface area, the others being the smaller but more populous North Island and Stewart Island. It is bordered to the north by ...
but is now confined to
Chatham Island Chatham Island ( ) ( Moriori: , 'Misty Sun'; ) is the largest island of the Chatham Islands group, in the south Pacific Ocean off the eastern coast of New Zealand's South Island. It is said to be "halfway between the equator and the pole, a ...
and
Pitt Island Pitt Island ( Moriori: , ) is the second largest island in New Zealand's Chatham Islands, with an area of . It lies about to the east of New Zealand's main islands, and about to the southeast of Chatham Island, from which it is separated by ...
, to which it was introduced in the early 1900s, and where they are widely hunted and eaten in the autumn (due to their status as non-native to the islands). Reintroduction into
Canterbury Canterbury (, ) is a City status in the United Kingdom, city and UNESCO World Heritage Site, in the county of Kent, England; it was a county borough until 1974. It lies on the River Stour, Kent, River Stour. The city has a mild oceanic climat ...
has been unsuccessful so far, but introductions to
Mou Waho Mou Waho is a 120-hectare island in Lake Wānaka, New Zealand. It is around the same size as the nearby Mou Tapu, these two islands being the largest in the lake. The island contains a small recursive lake, called Arethusa Pool: a glacial-sco ...
Island (in
Lake Wānaka Lake Wānaka is New Zealand's List of lakes in New Zealand#Largest lakes, fourth-largest lake and the seat of the town of Wānaka in the Otago region. The lake is 278 meters above sea level, covers , and is more than deep. "Wānaka" is the So ...
in 2004) and from there to
Pigeon Columbidae is a bird family consisting of doves and pigeons. It is the only family in the order Columbiformes. These are stout-bodied birds with small heads, relatively short necks and slender bills that in some species feature fleshy ceres. ...
and Pig Island (in
Lake Wakatipu Lake Wakatipu () is an inland lake (finger lake) in the South Island of New Zealand. It is in the southwest corner of the Otago region, near its boundary with Southland, New Zealand, Southland. ''Lake Wakatipu'' comes from the original Māori l ...
in 2005/2006) have been much more successful. It has a lighter overall colouring than the other subspecies. The North Island weka (''G. a. greyi'') is represented by original populations in Northland, and
Poverty Bay Poverty Bay (Māori language, Māori: ''Tūranganui-a-Kiwa''), officially named Tūranganui-a-Kiwa / Poverty Bay, is the largest of several small bays on the east coast of New Zealand's North Island to the north of Hawke Bay. It stretches for ...
, and by liberations elsewhere from that stock. This subspecies differs in its greyer underparts, and brown rather than reddish coloured legs.Penguin Pocket Guides: New Zealand's Native Birds of Bush and Countryside. Pages 76–77. 1996. Published by Penguin Books. Individual sightings around the
Dunedin Dunedin ( ; ) is the second-most populous city in the South Island of New Zealand (after Christchurch), and the principal city of the Otago region. Its name comes from ("fort of Edin"), the Scottish Gaelic name for Edinburgh, the capital of S ...
area suggest several individuals may exist in the southeastern South Island, either as surreptitious or inadvertent introductions, or possibly as remnants of a population previously thought extinct. The
nominate subspecies In biological classification, subspecies (: subspecies) is a rank below species, used for populations that live in different areas and vary in size, shape, or other physical characteristics ( morphology), but that can successfully interbreed. ...
, the western weka (''G. a. australis''), is found mainly in the northern and western regions of the South Island from
Nelson Nelson may refer to: Arts and entertainment * ''Nelson'' (1918 film), a historical film directed by Maurice Elvey * ''Nelson'' (1926 film), a historical film directed by Walter Summers * ''Nelson'' (opera), an opera by Lennox Berkeley to a lib ...
to
Fiordland Fiordland (, "The Pit of Tattooing", and also translated as "the Shadowlands"), is a non-administrative geographical region of New Zealand in the south-western corner of the South Island, comprising the western third of Southland. Most of F ...
. Distinguished by dark red-brown and black streaking on the breast, the western weka has two distinct colour phases, that of the southernmost range showing a greater degree of black. The Stewart Island weka (''G. a. scotti'') is smaller than the other subspecies and, like the western weka, has two colour phases; a chestnut form – similar to the chestnut-phase western weka – and a black phase which is not as dark as the black western weka. Reintroduced populations are confined to
Stewart Island / Rakiura Stewart Island (, 'Aurora, glowing skies', officially Stewart Island / Rakiura, formerly New Leinster) is New Zealand's third-largest island, located south of the South Island, across Foveaux Strait. It is a roughly triangular island wit ...
and outliers, and to
Kapiti Island Kapiti Island (), sometimes written as Kāpiti Island, is an island nature reserve located off the west coast of the lower North Island of New Zealand and within the Kāpiti Coast District. Parts of the island were previously farmed, but it is ...
to which it was introduced. Weka are host to two species of
feather lice A bird louse is any chewing louse (small, biting insects) of order Phthiraptera which parasitizes warm-blooded animals, especially birds. Bird lice may feed on feathers, skin, or blood. They have no wings, and their biting mouth parts distingui ...
(''Rallicola harrisoni'' and ''Pseudomenopon pilgrimi'') which show the same north–south population structure seen in their hosts.


Behaviour


Habitat and diet

Weka occupy areas such as forests,
sub-alpine Montane ecosystems are found on the slopes of mountains. The alpine climate in these regions strongly affects the ecosystem because temperatures fall as elevation increases, causing the ecosystem to stratify. This stratification is a crucial f ...
grassland,
sand dune A dune is a landform composed of wind- or water-driven sand. It typically takes the form of a mound, ridge, or hill. An area with dunes is called a dune system or a dune complex. A large dune complex is called a dune field, while broad, flat ...
s,
rocky shore A rocky shore is an intertidal area of seacoasts where solid rock predominates. Rocky shores are biologically rich environments, and are a useful "natural laboratory" for studying intertidal ecology and other biological processes. Due to th ...
s and modified semi-urban environments. They are omnivorous, with a diet comprising 30% animal foods and 70% plant foods. Animal foods include
earthworm An earthworm is a soil-dwelling terrestrial invertebrate that belongs to the phylum Annelida. The term is the common name for the largest members of the class (or subclass, depending on the author) Oligochaeta. In classical systems, they we ...
s,
larva A larva (; : larvae ) is a distinct juvenile form many animals undergo before metamorphosis into their next life stage. Animals with indirect development such as insects, some arachnids, amphibians, or cnidarians typically have a larval phase ...
e, beetles,
wētā Wētā (also spelled weta in English) is the common name for a group of about 100 insect species in the families Anostostomatidae and Rhaphidophoridae endemism, endemic to New Zealand. They are giant wingless insect, flightless cricket (insect ...
, ants, grass grubs, slugs, snails, insect eggs,
slater A slater, or slate mason, is a tradesperson who covers buildings with slate. Tools used The various tools of the slater's trade are all drop-forged. The slater's hammer is forged in one single piece, from crucible-cast steel, and has a lea ...
s, frogs, spiders, rats, mice, and small birds. Stewart Island weka (''G. a. scotti'') have been observed preying on
sooty shearwater The sooty shearwater (''Ardenna grisea'') is a medium-large shearwater in the seabird family Procellariidae. In New Zealand, it is also known by its Māori language, Māori name , and is harvested by Māori people for muttonbirding, muttonbird, l ...
(''Puffinus griseus'') eggs and chicks. Plant foods include leaves, grass, berries and seeds. Weka are important in the bush as seed dispersers, distributing seeds too large for smaller berry-eating birds. Where the weka is relatively common, their furtive curiosity leads them to search around houses and camps for food scraps, or anything unfamiliar and transportable. They have been known to take shiny objects in particular.


Breeding and Nesting

The breeding season varies, but when food is plentiful, weka can raise up to four broods throughout the whole year.
Nest A nest is a structure built for certain animals to hold Egg (biology), eggs or young. Although nests are most closely associated with birds, members of all classes of vertebrates and some invertebrates construct nests. They may be composed of ...
s are made on the ground under the cover of thick vegetation, and built by making grass (or similar material) into a bowl to hold about four eggs. On average, female weka lay three creamy or pinkish eggs blotched with brown and mauve. Both sexes incubate. The chicks hatch after a month, and are fed by both parents until fully grown between six and ten weeks.


Conservation status

Weka are classed as a
vulnerable species A vulnerable species is a species which has been Conservation status, categorized by the International Union for Conservation of Nature as being threatened species, threatened with extinction unless the circumstances that are threatened species, ...
. The Department of Conservation's weka recovery plan, approved in 1999, aims to improve the conservation status of threatened weka, clarify the status of data deficient weka, maintain the non-threatened status of other weka, and eventually restore all weka to their traditional ranges as a significant component of the ecosystems. Weka are problematic in conservation; some subspecies are threatened, but have been a problem to other threatened wildlife on offshore islands, especially when introduced to an island that they would not naturally inhabit. Weka are unable to withstand the current pressures faced in both the
North Island The North Island ( , 'the fish of Māui', historically New Ulster) is one of the two main islands of New Zealand, islands of New Zealand, separated from the larger but less populous South Island by Cook Strait. With an area of , it is the List ...
and
South Island The South Island ( , 'the waters of Pounamu, Greenstone') is the largest of the three major islands of New Zealand by surface area, the others being the smaller but more populous North Island and Stewart Island. It is bordered to the north by ...
. However, they can be very productive in good conditions and high food availability. Year-round breeding has been recorded at several sites with up to 14 young produced in a year. Weka populations can persist in highly modified habitats, but they have disappeared from huge areas of their former range, suggesting that they can adapt to a wide range of conditions but are particularly vulnerable to threats.


Threats

The Department of Conservation identifies eight main threats to weka.
Predation Predation is a biological interaction in which one organism, the predator, kills and eats another organism, its prey. It is one of a family of common List of feeding behaviours, feeding behaviours that includes parasitism and micropredation ...
by
ferret The ferret (''Mustela furo'') is a small, domesticated species belonging to the family Mustelidae. The ferret is most likely a domesticated form of the wild European polecat (''Mustela putorius''), as evidenced by the ferret's ability to inter ...
s, cats, and dogs are a threat to adult weka;
stoat The stoat (''Mustela erminea''), also known as the Eurasian ermine or ermine, is a species of mustelid native to Eurasia and the northern regions of North America. Because of its wide circumpolar distribution, it is listed as Least Concern on th ...
s and ferrets are a threat to chicks; stoats and rats are a threat to eggs. It faces competition with introduced species for fruits and invertebrates, and suffers from the impact browsers have on forest composition and regeneration. Habitat depletion is caused by the modification and degradation of forests and wetlands. Diseases and
parasite Parasitism is a Symbiosis, close relationship between species, where one organism, the parasite, lives (at least some of the time) on or inside another organism, the Host (biology), host, causing it some harm, and is Adaptation, adapted str ...
s have been associated with population declines, although little is known. Drought has been implicated in the disappearance of weka from some areas. In some regions, motor vehicles cause a significant amount of
roadkill Roadkill is a wild animal that has been killed by collision with motor vehicles. Wildlife-vehicle collisions (WVC) have increasingly been the topic of academic research to understand the causes, and how they can be mitigated. History Essenti ...
death. Pest control operations sometimes kill weka, as they have ground foraging habits vulnerable to poison baits, and traps are laid in a way that weka can reach.
Genetic diversity Genetic diversity is the total number of genetic characteristics in the genetic makeup of a species. It ranges widely, from the number of species to differences within species, and can be correlated to the span of survival for a species. It is d ...
can be lost during the transmission of genes through generations, affecting isolated populations.


Human interaction

Weka are significant to some
Māori Māori or Maori can refer to: Relating to the Māori people * Māori people of New Zealand, or members of that group * Māori language, the language of the Māori people of New Zealand * Māori culture * Cook Islanders, the Māori people of the Co ...
iwi Iwi () are the largest social units in New Zealand Māori society. In Māori, roughly means or , and is often translated as "tribe". The word is both singular and plural in the Māori language, and is typically pluralised as such in English. ...
(tribes) who admire their curiosity and feisty, bold personality traits which have led to them being relatively easy to catch. Weka were used by the Māori as a source of food, perfume, oil to treat
inflammation Inflammation (from ) is part of the biological response of body tissues to harmful stimuli, such as pathogens, damaged cells, or irritants. The five cardinal signs are heat, pain, redness, swelling, and loss of function (Latin ''calor'', '' ...
s, feathers in clothing and lures to catch dogs. Early European explorers and settlers frequently encountered and utilised weka; they were called "bush hens".


Incidents

In October 2023 a two-person team competing in a reality show were disqualified after one killed and ate a weka in what he claimed was an act of desperation and hunger, despite knowing it was a protected species. They were also issued with a letter of reprimand from the Department of Conservation.Rawling, C
US reality contestant kills and eats protected bird on NZ-filmed TV show
''
ABC News ABC News most commonly refers to: * ABC News (Australia), a national news service of the Australian Broadcasting Corporation * ABC News (United States), a news-gathering and broadcasting division of the American Broadcasting Company ABC News may a ...
'', 24 July 2024. Retrieved 24 July 2024.


References


External links


BirdLife Species Factsheet.
* {{Authority control Endemic birds of New Zealand Flightless birds Gallirallus Birds described in 1786 Taxa named by Anders Sparrman