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Gadiformes , also called the Anacanthini, are an
order Order, ORDER or Orders may refer to: * A socio-political or established or existing order, e.g. World order, Ancien Regime, Pax Britannica * Categorization, the process in which ideas and objects are recognized, differentiated, and understood ...
of
ray-finned fish Actinopterygii (; ), members of which are known as ray-finned fish or actinopterygians, is a class of bony fish that comprise over 50% of living vertebrate species. They are so called because of their lightly built fins made of webbings of sk ...
that include the
cod Cod (: cod) is the common name for the demersal fish genus ''Gadus'', belonging to the family (biology), family Gadidae. Cod is also used as part of the common name for a number of other fish species, and one species that belongs to genus ''Gad ...
, hakes,
pollock Pollock or pollack (pronounced ) is the common name used for either of the two species of North Atlantic ocean, marine fish in the genus ''Pollachius''. ''Pollachius pollachius'' is referred to as "pollock" in North America, Ireland and the Unit ...
,
haddock The haddock (''Melanogrammus aeglefinus'') is a saltwater ray-finned fish from the Family (biology), family Gadidae, the true cods. It is the only species in the Monotypy, monotypic genus ''Melanogrammus''. It is found in the North Atlantic Oce ...
,
burbot The burbot (''Lota lota''), also known as bubbot, mariah, loche, cusk, freshwater cod, freshwater ling, freshwater cusk, the lawyer, coney-fish, lingcod, or eelpout, is a species of coldwater ray-finned fish native to the subarctic regions of ...
, rocklings and moras, many of which are
food fish Many species of fish are caught by humans and consumed as food in virtually all regions around the world. Their meat has been an important dietary source of protein and other nutrients in the human diet. The English language does not have a s ...
of major commercial value. They are mostly marine fish found throughout the world and the vast majority are found in
temperate In geography, the temperate climates of Earth occur in the middle latitudes (approximately 23.5° to 66.5° N/S of the Equator), which span between the tropics and the polar regions of Earth. These zones generally have wider temperature ran ...
or colder regions (tropical species are typically deep-water) while a few species may enter
brackish Brackish water, sometimes termed brack water, is water occurring in a natural environment that has more salinity than freshwater, but not as much as seawater. It may result from mixing seawater (salt water) and fresh water together, as in estuari ...
estuaries An estuary is a partially enclosed coastal body of brackish water with one or more rivers or streams flowing into it, and with a free connection to the open sea. Estuaries form a transition zone between river environments and maritime environm ...
. Pacific tomcods, one of the two species that makes up the genus ''
Microgadus ''Microgadus'', the tomcods, is a genus of cods. Species There are currently two recognized species in this genus: * '' Microgadus proximus'' ( Girard, 1854) (Pacific tomcod) * '' Microgadus tomcod'' ( Walbaum, 1792) (Atlantic tomcod) Referenc ...
'', are able to enter freshwater, but there is no evidence that they breed there. Some populations of landlocked Atlantic tomcod on the other hand, complete their entire life cycle in freshwater. Yet only one species, the
burbot The burbot (''Lota lota''), also known as bubbot, mariah, loche, cusk, freshwater cod, freshwater ling, freshwater cusk, the lawyer, coney-fish, lingcod, or eelpout, is a species of coldwater ray-finned fish native to the subarctic regions of ...
(''Lota lota''), is a true
freshwater fish Freshwater fish are fish species that spend some or all of their lives in bodies of fresh water such as rivers, lakes, ponds and inland wetlands, where the salinity is less than 1.05%. These environments differ from marine habitats in many wa ...
. Common characteristics include the positioning of the
pelvic fin Pelvic fins or ventral fins are paired fins located on the ventral (belly) surface of fish, and are the lower of the only two sets of paired fins (the other being the laterally positioned pectoral fins). The pelvic fins are homologous to the hi ...
s (if present), below or in front of the
pectoral fin Fins are moving appendages protruding from the body of fish that interact with water to generate thrust and help the fish aquatic locomotion, swim. Apart from the tail or caudal fin, fish fins have no direct connection with the vertebral column ...
s. Gadiformes are physoclists, which means their
swim bladder The swim bladder, gas bladder, fish maw, or air bladder is an internal gas-filled organ (anatomy), organ in bony fish that functions to modulate buoyancy, and thus allowing the fish to stay at desired water depth without having to maintain lift ...
s do not have a pneumatic duct. The fins are spineless. Gadiform fish range in size from the codlets, which may be as small as in adult length, to the
Atlantic cod The Atlantic cod (: cod; ''Gadus morhua'') is a fish of the family Gadidae, widely consumed by humans. It is also commercially known as '' cod'' or ''codling''.Palaeogadus weltoni'' from the
Maastrichtian The Maastrichtian ( ) is, in the International Commission on Stratigraphy (ICS) geologic timescale, the latest age (geology), age (uppermost stage (stratigraphy), stage) of the Late Cretaceous epoch (geology), Epoch or Upper Cretaceous series (s ...
of the United States and the undescribed, informally named "''Protocodus''" from the
Early Paleocene The Danian is the oldest age (geology), age or lowest stage (stratigraphy), stage of the Paleocene Epoch or series (stratigraphy), Series, of the Paleogene Period or system (stratigraphy), System, and of the Cenozoic Era or Erathem. The beginnin ...
of
Greenland Greenland is an autonomous territory in the Danish Realm, Kingdom of Denmark. It is by far the largest geographically of three constituent parts of the kingdom; the other two are metropolitan Denmark and the Faroe Islands. Citizens of Greenlan ...
.


Taxonomy

The following classification is based on ''
Eschmeyer's Catalog of Fishes Catalog of Fishes is a comprehensive on-line database and reference work on the scientific names of fish species and genera. It is global in its scope and is hosted by the California Academy of Sciences. It has been compiled and is continuously up ...
'': * Order Gadiformes ** Suborder Stylephoroidei *** Family Stylephoridae Swainson, 1839 (tube-eyes or threadtails) ** Suborder Bregmacerotoidei *** Family Bregmacerotidae
Gill A gill () is a respiration organ, respiratory organ that many aquatic ecosystem, aquatic organisms use to extract dissolved oxygen from water and to excrete carbon dioxide. The gills of some species, such as hermit crabs, have adapted to allow r ...
, 1872
(codlets) ** Suborder Gadoidei *** Family Phycidae Swainson, 1838 (phycid hakes) *** Family Gaidropsaridae
Jordan Jordan, officially the Hashemite Kingdom of Jordan, is a country in the Southern Levant region of West Asia. Jordan is bordered by Syria to the north, Iraq to the east, Saudi Arabia to the south, and Israel and the occupied Palestinian ter ...
& Evermann, 1898
(rocklings) *** Family
Lotidae The Lotidae are a family of cod-like fishes commonly known as lings or rocklings. They are found in the Arctic, Atlantic, and Pacific Oceans. Except for a few species of '' Gaidropsarus'', all are restricted to the Northern Hemisphere. All spe ...
Bonaparte, 1835 (hakes and burbots) *** Family
Gadidae The Gadidae are a family of marine fish, included in the order Gadiformes, known as the cods, codfishes, or true cods. It contains several commercially important fishes, including the cod, haddock, whiting, and pollock. Most gadid species ar ...
Rafinesque Constantine Samuel Rafinesque-Schmaltz (; 22 October 178318 September 1840) was a French early 19th-century polymath born near Constantinople in the Ottoman Empire and self-educated in France. He traveled as a young man in the United States, ult ...
, 1810
(cods and haddocks) ** Suborder Ranicipitoidei *** Family Ranicipitidae Bonaparte, 1835 (tadpole fishes) ** Suborder Merluccioidei *** Family Merlucciidae Rafinesque, 1815 (merlucciid hakes) ** Suborder Macrouroidei *** Family Euclichthyidae Cohen, 1984 (Eucla cods) *** Family Muraenolepididae Regan, 1903 (eel cods) *** Family Melanonidae Goode &
Bean A bean is the seed of some plants in the legume family (Fabaceae) used as a vegetable for human consumption or animal feed. The seeds are often preserved through drying (a ''pulse''), but fresh beans are also sold. Dried beans are traditi ...
, 1896
(arrowtails or pelagic cods) *** Family Trachyrincidae Goode & Bean, 1896 (armoured grenadiers) *** Family Moridae Moreau, 1881 (codlings or deepsea cods) *** Family Macruronidae Regan, 1903 (blue grenadiers) *** Family Lyconidae Günther, 1887 (Atlantic hakes) *** Family Bathygadidae Jordan & Evermann, 1898 (rattails) *** Family Steindachneriidae Parr, 1942 (luminous hakes) *** Family
Macrouridae Macrouridae is a family of deep sea fish, a diverse and ecologically important group, which are part of the order of cod-like fish, the Gadiformes. The species in the Macrouridae are characterised by their large heads which normally have a sing ...
Bonaparte, 1831 (grenadiers or rattails)Iwamoto, T., Nakayama, N., Shao, K.-T. & Ho, H.-C. (2015): Synopsis of the Grenadier Fishes (Gadiformes; Teleostei) of Taiwan. ''Proceedings of the California Academy of Sciences, (Series 4), 62 (3): 31–126.''


Timeline of genera

ImageSize = width:900px height:auto barincrement:15px PlotArea = left:10px bottom:50px top:10px right:50px Period = from:-145.5 till:10 TimeAxis = orientation:horizontal ScaleMajor = unit:year increment:10 start:-145.5 ScaleMinor = unit:year increment:1 start:-145.5 TimeAxis = orientation:hor AlignBars = justify Colors = #legends id:CAR value:claret id:ANK value:rgb(0.4,0.3,0.196) id:HER value:teal id:HAD value:green id:OMN value:blue id:black value:black id:white value:white id:cretaceous value:rgb(0.5,0.78,0.31) id:earlycretaceous value:rgb(0.63,0.78,0.65) id:latecretaceous value:rgb(0.74,0.82,0.37) id:cenozoic value:rgb(0.54,0.54,0.258) id:paleogene value:rgb(0.99,0.6,0.32) id:paleocene value:rgb(0.99,0.65,0.37) id:eocene value:rgb(0.99,0.71,0.42) id:oligocene value:rgb(0.99,0.75,0.48) id:neogene value:rgb(0.999999,0.9,0.1) id:miocene value:rgb(0.999999,0.999999,0) id:pliocene value:rgb(0.97,0.98,0.68) id:quaternary value:rgb(0.98,0.98,0.5) id:pleistocene value:rgb(0.999999,0.95,0.68) id:holocene value:rgb(0.999,0.95,0.88) BarData= bar:eratop bar:space bar:periodtop bar:space bar:NAM1 bar:NAM2 bar:NAM3 bar:NAM4 bar:NAM5 bar:NAM6 bar:NAM7 bar:NAM8 bar:NAM9 bar:NAM10 bar:NAM11 bar:NAM12 bar:NAM13 bar:NAM14 bar:NAM15 bar:NAM16 bar:NAM17 bar:NAM18 bar:NAM19 bar:NAM20 bar:NAM21 bar:NAM22 bar:NAM23 bar:NAM24 bar:NAM25 bar:NAM26 bar:NAM27 bar:NAM28 bar:NAM29 bar:NAM30 bar:NAM31 bar:NAM32 bar:NAM33 bar:NAM34 bar:NAM35 bar:NAM36 bar:NAM37 bar:NAM38 bar:NAM39 bar:NAM40 bar:NAM41 bar:NAM42 bar:NAM43 bar:NAM44 bar:NAM45 bar:NAM46 bar:NAM47 bar:NAM48 bar:NAM49 bar:NAM50 bar:NAM51 bar:NAM52 bar:space bar:period bar:space bar:era PlotData= align:center textcolor:black fontsize:M mark:(line,black) width:25 shift:(7,-4) bar:periodtop from: -145.5 till: -99.6 color:earlycretaceous text: Early from: -99.6 till: -65.5 color:latecretaceous text: Late from: -65.5 till: -55.8 color:paleocene text: Paleo. from: -55.8 till: -33.9 color:eocene text: Eo. from: -33.9 till: -23.03 color:oligocene text: Oligo. from: -23.03 till: -5.332 color:miocene text: Mio. from: -5.332 till: -2.588 color:pliocene text: Pl. from: -2.588 till: -0.0117 color:pleistocene text: Pl. from: -0.0117 till: 0 color:holocene text: H. bar:eratop from: -145.5 till: -65.5 color:cretaceous text:
Cretaceous The Cretaceous ( ) is a geological period that lasted from about 143.1 to 66 mya (unit), million years ago (Mya). It is the third and final period of the Mesozoic Era (geology), Era, as well as the longest. At around 77.1 million years, it is the ...
from: -65.5 till: -23.03 color:paleogene text:
Paleogene The Paleogene Period ( ; also spelled Palaeogene or Palæogene) is a geologic period and system that spans 43 million years from the end of the Cretaceous Period Ma (million years ago) to the beginning of the Neogene Period Ma. It is the fir ...
from: -23.03 till: -2.588 color:neogene text:
Neogene The Neogene ( ,) is a geologic period and system that spans 20.45 million years from the end of the Paleogene Period million years ago ( Mya) to the beginning of the present Quaternary Period million years ago. It is the second period of th ...
from: -2.588 till: 0 color:quaternary text: Q. PlotData= align:left fontsize:M mark:(line,white) width:5 anchor:till align:left color:latecretaceous bar:NAM1 from: -85.8 till: -70.6 text: Rankinian color:latecretaceous bar:NAM2 from: -83.5 till: -65.5 text: Paratichthys color:eocene bar:NAM3 from: -55.8 till: -48.6 text: Rhinocephalus color:eocene bar:NAM4 from: -55.8 till: -37.2 text: Trichurides color:eocene bar:NAM5 from: -55.8 till: -11.608 text: Palaeogadus color:eocene bar:NAM6 from: -55.8 till: 0 text: Urophycis color:eocene bar:NAM7 from: -55.8 till: 0 text: Raniceps color:eocene bar:NAM8 from: -55.8 till: 0 text: Nezumia color:eocene bar:NAM9 from: -55.8 till: 0 text: Merluccius color:eocene bar:NAM10 from: -55.8 till: 0 text: Melanonus color:eocene bar:NAM11 from: -48.6 till: 0 text: Bregmaceros color:eocene bar:NAM12 from: -37.2 till: 0 text: Tripterophycis color:eocene bar:NAM13 from: -37.2 till: 0 text: Macruronus color:eocene bar:NAM14 from: -37.2 till: 0 text: Eulichthys color:eocene bar:NAM15 from: -37.2 till: 0 text: Coelorhynchus color:eocene bar:NAM16 from: -37.2 till: 0 text: Bathygadus color:oligocene bar:NAM17 from: -33.9 till: -28.4 text: Eophycis color:oligocene bar:NAM18 from: -33.9 till: 0 text: Trachyrhinchus color:oligocene bar:NAM19 from: -33.9 till: 0 text: Squalogadus color:oligocene bar:NAM20 from: -33.9 till: 0 text: Phycis color:oligocene bar:NAM21 from: -33.9 till: 0 text: Lota color:oligocene bar:NAM22 from: -33.9 till: 0 text: Hymenocephalus color:oligocene bar:NAM23 from: -33.9 till: 0 text: Gaidropsarus color:oligocene bar:NAM24 from: -33.9 till: 0 text: Gadiculus color:oligocene bar:NAM25 from: -28.4 till: -15.97 text: Pseudoraniceps color:oligocene bar:NAM26 from: -28.4 till: -11.608 text: Palaeomolva color:oligocene bar:NAM27 from: -28.4 till: 0 text: Trisopterus color:oligocene bar:NAM28 from: -28.4 till: 0 text: Micromesistius color:oligocene bar:NAM29 from: -28.4 till: 0 text: Coryphaenoides color:oligocene bar:NAM30 from: -28.4 till: 0 text: Brosmius color:miocene bar:NAM31 from: -23.03 till: -15.97 text: Bregmacerinia color:miocene bar:NAM32 from: -23.03 till: -5.332 text: Trichiurichthys color:miocene bar:NAM33 from: -23.03 till: 0 text: Ventrifossa color:miocene bar:NAM34 from: -23.03 till: 0 text:
Physiculus ''Physiculus'' is a genus of morid cods. Species The 42 currently recognized species in this genus are: * ''Physiculus andriashevi'' Yuri Nikolaevich Shcherbachev, Shcherbachev, 1993 * ''Physiculus argyropastus'' Alfred William Alcock, Alcock, 1 ...
color:miocene bar:NAM35 from: -23.03 till: 0 text: Austrophycis color:miocene bar:NAM36 from: -15.97 till: 0 text: Molva color:miocene bar:NAM37 from: -15.97 till: 0 text:
Merlangius ''Merlangius merlangus'', commonly known as whiting or merling, is an important food fish in the eastern North Atlantic Ocean and the northern Mediterranean, western Baltic, and Black Sea. In Anglophonic countries outside the whiting's natura ...
color:miocene bar:NAM38 from: -15.97 till: 0 text: Lepidion color:miocene bar:NAM39 from: -15.97 till: 0 text: Laemonema color:miocene bar:NAM40 from: -11.608 till: -5.332 text: Eclipes color:miocene bar:NAM41 from: -11.608 till: -5.332 text: Bolbocara color:miocene bar:NAM42 from: -11.608 till: 0 text: Paratrisopterus color:miocene bar:NAM43 from: -11.608 till: 0 text: Gadus color:miocene bar:NAM44 from: -11.608 till: 0 text: Gadomus color:miocene bar:NAM45 from: -11.608 till: 0 text: Brosme color:pliocene bar:NAM46 from: -5.332 till: -2.588 text:
Pseudophycis ''Pseudophycis'' is a genus of codlings of the family Moridae found around New Zealand New Zealand () is an island country in the southwestern Pacific Ocean. It consists of two main landmasses—the North Island () and the South Islan ...
color:pliocene bar:NAM47 from: -5.332 till: -2.588 text: Gadella color:pliocene bar:NAM48 from: -5.332 till: 0 text: Theragra color:pliocene bar:NAM49 from: -5.332 till: 0 text:
Microgadus ''Microgadus'', the tomcods, is a genus of cods. Species There are currently two recognized species in this genus: * '' Microgadus proximus'' ( Girard, 1854) (Pacific tomcod) * '' Microgadus tomcod'' ( Walbaum, 1792) (Atlantic tomcod) Referenc ...
color:pliocene bar:NAM50 from: -5.332 till: 0 text: Melanogrammus color:pliocene bar:NAM51 from: -5.332 till: 0 text: Lycodopsis color:pleistocene bar:NAM52 from: -2.588 till: -0.0117 text: Pollachius PlotData= align:center textcolor:black fontsize:M mark:(line,black) width:25 bar:period from: -145.5 till: -99.6 color:earlycretaceous text: Early from: -99.6 till: -65.5 color:latecretaceous text: Late from: -65.5 till: -55.8 color:paleocene text: Paleo. from: -55.8 till: -33.9 color:eocene text: Eo. from: -33.9 till: -23.03 color:oligocene text: Oligo. from: -23.03 till: -5.332 color:miocene text: Mio. from: -5.332 till: -2.588 color:pliocene text: Pl. from: -2.588 till: -0.0117 color:pleistocene text: Pl. from: -0.0117 till: 0 color:holocene text: H. bar:era from: -145.5 till: -65.5 color:cretaceous text:
Cretaceous The Cretaceous ( ) is a geological period that lasted from about 143.1 to 66 mya (unit), million years ago (Mya). It is the third and final period of the Mesozoic Era (geology), Era, as well as the longest. At around 77.1 million years, it is the ...
from: -65.5 till: -23.03 color:paleogene text:
Paleogene The Paleogene Period ( ; also spelled Palaeogene or Palæogene) is a geologic period and system that spans 43 million years from the end of the Cretaceous Period Ma (million years ago) to the beginning of the Neogene Period Ma. It is the fir ...
from: -23.03 till: -2.588 color:neogene text:
Neogene The Neogene ( ,) is a geologic period and system that spans 20.45 million years from the end of the Paleogene Period million years ago ( Mya) to the beginning of the present Quaternary Period million years ago. It is the second period of th ...
from: -2.588 till: 0 color:quaternary text: Q.


References

{{Authority control Taxa named by Edwin Stephen Goodrich Ray-finned fish orders Extant Maastrichtian first appearances