The Formicinae are a
subfamily
In biological classification, a subfamily (Latin: ', plural ') is an auxiliary (intermediate) taxonomic rank, next below family but more inclusive than genus. Standard nomenclature rules end subfamily botanical names with "-oideae", and zoologi ...
within the
Formicidae containing
ants of moderate evolutionary development.
Formicines retain some primitive features, such as the presence of
cocoons around
pupae, the presence of
ocelli in workers, and little tendency toward reduction of
palp or
antenna
Antenna ( antennas or antennae) may refer to:
Science and engineering
* Antenna (radio), also known as an aerial, a transducer designed to transmit or receive electromagnetic (e.g., TV or radio) waves
* Antennae Galaxies, the name of two collid ...
l segmentation in most species, except
subterranean
Subterranean(s) or The Subterranean(s) may refer to:
* Subterranea (geography), underground structures, both natural and man-made
Literature
* ''Subterranean'' (novel), a 1998 novel by James Rollins
* ''Subterranean Magazine'', an American fa ...
groups. Extreme modification of
mandibles is rare, except in the genera ''
Myrmoteras'' and ''
Polyergus''. However, some members show considerable evolutionary advancement in behaviors such as
slave-making and symbiosis with root-feeding
hemiptera
Hemiptera (; ) is an order (biology), order of insects, commonly called true bugs, comprising over 80,000 species within groups such as the cicadas, aphids, planthoppers, leafhoppers, Reduviidae, assassin bugs, Cimex, bed bugs, and shield bugs. ...
ns. Finally, all formicines have very reduced
sting
Sting may refer to:
* Stinger or sting, a structure of an animal to inject venom, or the injury produced by a stinger
* Irritating hairs or prickles of a stinging plant, or the plant itself
Fictional characters and entities
* Sting (Middle-eart ...
s and enlarged
venom reservoirs, with the venom gland, specialized (uniquely among ants) for the production of
formic acid
Formic acid (), systematically named methanoic acid, is the simplest carboxylic acid, and has the chemical formula HCOOH and structure . It is an important intermediate in chemical synthesis and occurs naturally, most notably in some ants. Es ...
.
All members of the Formicinae "have a one-segmented
petiole in the form of a vertical scale".
Identification
Formicine ants have a single node-like or scale-like
petiole (postpetiole entirely lacking) and the apex of the abdomen has a circular or U-shaped opening (the
acidopore), usually fringed with hairs. A functional sting is absent, and defense is provided by the ejection of formic acid through the acidopore. If the acidopore is concealed by the pygidium and difficult to discern, then the antennal sockets are located well behind the posterior margin of the
clypeus (cf.
Dolichoderinae). In most formicines, the eyes are well developed (ocelli may also be present), the antennal insertions are not concealed by the frontal carinae, and the promesonotal suture is present and flexible.
Tribes and genera
The
tribal structure of the Formicinae is not completely understood. This list follows the scheme at AntCat,
but other schemes and names are used.
*
Camponotini Forel, 1878
** ''
Calomyrmex''
Emery, 1895
** ''
Camponotus''
Mayr, 1861 – carpenter ants (global)
** †''
Chimaeromyrma''
Dlussky, 1988
** ''
Colobopsis
''Colobopsis'' is a genus of ant in the subfamily Formicinae. This genus was first described in 1861 by Mayr and contains 95 species. The type species is '' Colobopsis truncata''.
Description
As part of Camponotini, ''Colobopsis'' workers have ...
''
Mayr, 1861
** ''
Dinomyrmex''
Ashmead, 1905
** ''
Echinopla''
Smith, 1857
** ''
Opisthopsis''
Dalla Torre, 1893
** ''
Overbeckia
''Overbeckia'' is a genus of ants in the subfamily Formicinae originally containing only the single species ''Overbeckia subclavata''. Although the genus has been known over 100 years only from Singapore
Singapore (), officially the Re ...
''
Viehmeyer, 1916
** ''
Polyrhachis''
Smith, 1857 (Asian, African tropics)
** †''
Pseudocamponotus
''Pseudocamponotus'' is an extinct, monotypic genus of ant. This genus was first described by F.M. Carpenter in 1930. The type species is ''Pseudocamponotus elkoanus'', whose fossil was found in Nevada. Species
*''Pseudocamponotus elkoanus''
Ref ...
''
Carpenter, 1930
*
Formicini Latreille, 1809
** ''
Alloformica''
Dlussky, 1969
** ''
Bajcaridris''
Agosti, 1994
** ''
Cataglyphis
''Cataglyphis'' is a genus of ant, desert ants, in the subfamily Formicinae. Its most famous species is ''C. bicolor'', the Sahara Desert ant, which runs on hot sand to find insects that died of heat exhaustion, and can, like other several oth ...
''
Foerster, 1850
** †''
Cataglyphoides''
Dlussky, 2008
** †''
Conoformica''
Dlussky, 2008
** ''
Formica''
Linnaeus, 1758
** ''
Iberoformica''
Tinaut, 1990
** ''
Polyergus''
Latreille, 1804 – Amazon ants
** ''
Proformica
''Proformica'' is a genus of ants in the subfamily Formicinae. The genus is known from the Palearctic realm, from Mongolia through Central Asia to Spain. Colonies are small, generally containing a few hundred individuals, with a single ...
''
Ruzsky, 1902
** †''
Protoformica''
Dlussky, 1967
** ''
Rossomyrmex''
Arnol'di, 1928
*
Gesomyrmecini Ashmead, 1905
** ''
Gesomyrmex
''Gesomyrmex'' is a genus of ants in the subfamily Formicinae. The genus contains six extant species, known from the Indomalayan realm, and nine fossil species. Of the extant species, four are known only from workers (''G. chaperi'', ''G. howar ...
''
Mayr, 1868
** †''
Prodimorphomyrmex''
Wheeler, 1915
** ''
Santschiella
''Santschiella'' is an Afrotropical genus of ants in the subfamily
In biological classification, a subfamily (Latin: ', plural ') is an auxiliary (intermediate) taxonomic rank, next below family but more inclusive than genus. Standard no ...
''
Forel, 1916
** †''
Sicilomyrmex''
Wheeler, 1915
*
Gigantiopini Ashmead, 1905
** ''
Gigantiops
''Gigantiops'' is a South American genus of jumping ants in the subfamily Formicinae. The genus contains the single species ''Gigantiops destructor'', which is also the sole member of the tribe Gigantiopini. They have the largest eyes among all ...
''
Roger, 1863 (Neotropical)
*
Lasiini Ashmead, 1905
** ''
Acropyga
''Acropyga'' is a genus of small formicine ants. Some species can be indirect pests. '' A. acutiventris'', which is found from India to Australia, tends subterranean, root-feeding mealybugs of the species '' Xenococcus annandalei''. Living, grav ...
''
Roger, 1862
** ''
Anoplolepis''
Santschi, 1914
** ''
Cladomyrma''
Wheeler, 1920
** †''
Glaphyromyrmex''
Wheeler, 1915
** ''
Lasiophanes
''Lasiophanes'' is a small genus of ant with six described species endemic to Patagonia (Argentina and Chile). The genus is related to '' Lasius'' of the Northern Hemisphere and '' Melophorus'' of Australia and New Zealand.
Little is known ab ...
''
Emery, 1895
** ''
Lasius''
Fabricius, 1804
** ''
Myrmecocystus''
Wesmael, 1838
** ''
Prolasius''
Forel, 1892
** ''
Stigmacros''
Forel, 1905
** ''
Teratomyrmex
''Teratomyrmex'' is a genus of ants in the subfamily Formicinae. The genus is known only from forested areas on the east coast of Australia.
Species
* ''Teratomyrmex greavesi'' McAreavey, 1957
* ''Teratomyrmex substrictus'' Shattuck & O'Reilly ...
''
McAreavey, 1957
*
Melophorini
Melophorini is a tribe of ants in the family Formicidae. There are about 10 genera in Melophorini.
Genera
These genera belong to the tribe Melophorini:
*'' Lasiophanes'' Emery, 1895
*'' Melophorus'' Lubbock, 1883
*''Myrmecorhynchus'' André, 1 ...
Forel, 1912
** ''
Melophorus''
Lubbock, 1883 (Australian)
*
Myrmecorhynchini Wheeler, 1917
** ''
Myrmecorhynchus
''Myrmecorhynchus'' is a genus of ants in the subfamily Formicinae. The genus is endemic to Australia
Australia, officially the Commonwealth of Australia, is a Sovereign state, sovereign country comprising the mainland of the Australia ...
''
André, 1896
** ''
Notoncus''
Emery, 1895
** ''
Pseudonotoncus
''Pseudonotoncus'' is a genus of ants in the subfamily Formicinae. The genus is known only from forested areas on the east coast of Australia
Australia, officially the Commonwealth of Australia, is a Sovereign state, sovereign countr ...
''
Clark, 1934
*
Myrmelachistini
Myrmelachistini is a tribe of ants in the family Formicidae. There are at least 2 genera and 50 described species in Myrmelachistini.
In 2016, the tribe Myrmelachistini was resurrected and the genera ''Brachymyrmex'' and ''Myrmelachista'' were ...
** ''
Brachymyrmex
''Brachymyrmex'' is a genus in the ants subfamily Formicinae. The genus can be recognized by the combination of having nine antennal segments (fewer than most ants) and the petiole concealed by the gaster in dorsal view. They are sometimes called ...
''
Mayr, 1868
** ''
Myrmelachista
''Myrmelachista'' is a Neotropical genus of ants in the subfamily Formicinae. The genus is found exclusively in the Neotropical realm. Little is known regarding their biology.
Habitat and distribution
The genus is restricted to the Neotropical r ...
''
Roger, 1863
*
Myrmoteratini Emery, 1895
** ''
Myrmoteras''
Forel, 1893
*
Notostigmatini Bolton, 2003
** ''
Notostigma''
Emery, 1920
*
Oecophyllini Emery, 1895
** ''
Oecophylla''
Smith, 1860 – weaver ants
*
Plagiolepidini Forel, 1886
** ''
Agraulomyrmex
''Agraulomyrmex'' is a genus of ants in the subfamily Formicinae. The genus is known from southern Africa.
Species
* '' Agraulomyrmex meridionalis'' Prins, 1983 – South Africa
* '' Agraulomyrmex wilsoni'' Prins, 1983 – Zimbabwe
...
''
Prins, 1983
** ''
Aphomomyrmex''
Emery, 1899
** ''
Bregmatomyrma''
Wheeler, 1929
** ''
Euprenolepis''
Emery, 1906
** ''
Lepisiota''
Santschi, 1926
** ''
Nylanderia''
Emery, 1906
** ''
Paraparatrechina''
Donisthorpe, 1947
** ''
Paratrechina''
Motschoulsky, 1863 – crazy ants
** ''
Petalomyrmex''
Snelling, 1979
** ''
Plagiolepis''
Mayr, 1861
** ''
Prenolepis''
Mayr, 1861
** ''
Pseudolasius''
Emery, 1887
** ''
Tapinolepis''
Emery, 1925
** ''
Zatania''
LaPolla, Kallal & Brady, 2012
* ''
incertae sedis''
** †''
Attopsis
''Attopsis'' is an extinct genus of ants in the formicid subfamily Formicinae. While formerly containing a number of species, the genus is currently monotypic; the type species, ''Attopsis longipennis'', is known from a single Early Miocene fo ...
''
Heer, 1850
** †''
Leucotaphus''
Donisthorpe, 1920
** †''
Liaoformica''
Hong, 2002
** †''
Longiformica''
Hong, 2002
** †''
Magnogasterites''
Hong, 2002
** †''
Orbicapitia''
Hong, 2002
** †''
Ovalicapito''
Hong, 2002
** †''
Ovaligastrula''
Hong, 2002
** †''
Protrechina
''Protrechina'' is an extinct, monotypic genus of ant, first described by Wilson (1985) from a Middle Eocene fossil found near Malvern, Arkansas. The genus contains a single described species ''Protrechina carpenteri'' known from a solitary worke ...
''
Wilson, 1985
** †''
Sinoformica''
Hong, 2002
** †''
Sinotenuicapito''
Hong, 2002
** †''
Wilsonia''
Hong, 2002
References
External links
*
*
{{Taxonbar, from=Q119691
Ant subfamilies
Taxa named by Amédée Louis Michel le Peletier