Fish Fish are aquatic, craniate, gill-bearing animals that lack limbs with digits. Included in this definition are the living hagfish, lampreys, and cartilaginous and bony fish as well as various extinct related groups. Approximately 95% ...

are very diverse animals and can be categorised in many ways. This article is an overview of some of ways in which fish are categorised. Although most fish

species In biology, a species is the basic unit of Taxonomy (biology), classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of ...

have probably been discovered and described, about 250 new ones are still discovered every year. According to

FishBase FishBase is a global species database of fish species (specifically finfish). It is the largest and most extensively accessed online database on adult finfish on the web.

about 34,800 species of fish had been described as of February 2022, which is more than the combined total of all other

vertebrate Vertebrates () comprise all animal taxon, taxa within the subphylum Vertebrata () (chordates with vertebral column, backbones), including all mammals, birds, reptiles, amphibians, and fish. Vertebrates represent the overwhelming majority of the ...

species: mammals, amphibians, reptiles and

bird Birds are a group of warm-blooded vertebrates constituting the class Aves (), characterised by feathers, toothless beaked jaws, the laying of hard-shelled eggs, a high metabolic rate, a four-chambered heart, and a strong yet lightweig ...

s. Fish species diversity is roughly divided equally between

marine Marine is an adjective meaning of or pertaining to the sea or ocean. Marine or marines may refer to: Ocean * Maritime (disambiguation) * Marine art * Marine biology * Marine debris * Marine habitats * Marine life * Marine pollution Military * ...

(oceanic) and

freshwater Fresh water or freshwater is any naturally occurring liquid or frozen water containing low concentrations of dissolved salts and other total dissolved solids. Although the term specifically excludes seawater and brackish water, it does in ...

ecosystems.

Coral reef A coral reef is an underwater ecosystem characterized by reef-building corals. Reefs are formed of colonies of coral polyps held together by calcium carbonate. Most coral reefs are built from stony corals, whose polyps cluster in groups. C ...

s in the

Indo-Pacific The Indo-Pacific is a vast biogeographic region of Earth. In a narrow sense, sometimes known as the Indo-West Pacific or Indo-Pacific Asia, it comprises the tropical waters of the Indian Ocean, the western and central Pacific Ocean, and the ...

constitute the centre of diversity for marine fishes, whereas continental freshwater fishes are most diverse in large

river basin A drainage basin is an area of land where all flowing surface water converges to a single point, such as a river mouth, or flows into another body of water, such as a lake or ocean. A basin is separated from adjacent basins by a perimeter, the ...

s of

tropical rainforest Tropical rainforests are rainforests that occur in areas of tropical rainforest climate in which there is no dry season – all months have an average precipitation of at least 60 mm – and may also be referred to as ''lowland equatori ...

s, especially the

Amazon Amazon most often refers to: * Amazons, a tribe of female warriors in Greek mythology * Amazon rainforest, a rainforest covering most of the Amazon basin * Amazon River, in South America * Amazon (company), an American multinational technolog ...

, Congo, and

Mekong The Mekong or Mekong River is a trans-boundary river in East Asia and Southeast Asia. It is the world's twelfth longest river and the third longest in Asia. Its estimated length is , and it drains an area of , discharging of water annuall ...

basins. More than 5,600 fish species inhabit

Neotropic The Neotropical realm is one of the eight biogeographic realms constituting Earth's land surface. Physically, it includes the tropical terrestrial ecoregions of the Americas and the entire South American temperate zone. Definition In bi ...

al freshwaters alone, such that

Neotropical fishes The freshwater fish of tropical South and Central America represent one of the most diverse and extreme aquatic ecosystems on Earth, with more than 5,600 species, representing about 10% all living vertebrate species. The exceptional diversity of sp ...

represent about 10% of all

species on the Earth. Exceptionally rich sites in the

Amazon basin The Amazon basin is the part of South America drained by the Amazon River and its tributaries. The Amazon drainage basin covers an area of about , or about 35.5 percent of the South American continent. It is located in the countries of Boli ...

, such as

Cantão State Park The Cantão State Park ( pt, Parque Estadual do Cantão) is a state park in the state of Tocantins, Brazil. It protects the Cantão wetlands, an area of igapó flooded forest in the ecotone where the Amazon rainforest merges into the cerrado. ...

, can contain more freshwater fish species than occur in all of Europe.

By taxonomy

Fish

systematics Biological systematics is the study of the diversification of living forms, both past and present, and the relationships among living things through time. Relationships are visualized as evolutionary trees (synonyms: cladograms, phylogenetic t ...

is the formal description and organisation of fish

taxa In biology, a taxon ( back-formation from '' taxonomy''; plural taxa) is a group of one or more populations of an organism or organisms seen by taxonomists to form a unit. Although neither is required, a taxon is usually known by a particular n ...

into systems. It is complex and still evolving. Controversies over "arcane, but important, details of classification are still quietly raging". The term "fish" describes any non-

tetrapod Tetrapods (; ) are four-limb (anatomy), limbed vertebrate animals constituting the superclass Tetrapoda (). It includes extant taxon, extant and extinct amphibians, sauropsids (reptiles, including dinosaurs and therefore birds) and synapsids (p ...

chordate A chordate () is an animal of the phylum Chordata (). All chordates possess, at some point during their larval or adult stages, five synapomorphies, or primary physical characteristics, that distinguish them from all the other taxa. These five ...

, (i.e., an animal with a backbone), that has

gill A gill () is a respiratory organ that many aquatic organisms use to extract dissolved oxygen from water and to excrete carbon dioxide. The gills of some species, such as hermit crabs, have adapted to allow respiration on land provided they ar ...

s throughout life and has limbs, if any, in the shape of fins. Unlike groupings such as

s or mammals, fish are not a single clade but a

paraphyletic In taxonomy (general), taxonomy, a group is paraphyletic if it consists of the group's most recent common ancestor, last common ancestor and most of its descendants, excluding a few Monophyly, monophyletic subgroups. The group is said to be pa ...

collection of

, including jawless, cartilaginous and skeletal types. Tree of life web project
Chordates

Jawless fish

Jawless fish Agnatha (, Ancient Greek 'without jaws') is an infraphylum of jawless fish in the phylum Chordata, subphylum Vertebrata, consisting of both present (Cyclostomata, cyclostomes) and extinct (conodonts and ostracoderms) species. Among recent animal ...

were the earliest fish to evolve. There is current debate over whether these are really fish at all. They have no jaw, no scales, no paired

fin A fin is a thin component or appendage attached to a larger body or structure. Fins typically function as foils that produce lift or thrust, or provide the ability to steer or stabilize motion while traveling in water, air, or other fluids. Fin ...

s, and no bony skeleton. Their skin is smooth and soft to the touch, and they are very flexible. Instead of a jaw, they possess an oral sucker. They use this to fasten onto other fish, and then use their rasp-like teeth to grind through their host's skin into the

viscera In biology, an organ is a collection of tissues joined in a structural unit to serve a common function. In the hierarchy of life, an organ lies between tissue and an organ system. Tissues are formed from same type cells to act together in a ...

. Jawless fish inhabit both fresh and salt water environments. Some are

anadromous Fish migration is mass relocation by fish from one area or body of water to another. Many types of fish migrate on a regular basis, on time scales ranging from daily to annually or longer, and over distances ranging from a few metres to thousan ...

, moving between both fresh and salt water habitats. Extant jawless fish are either

lamprey Lampreys (sometimes inaccurately called lamprey eels) are an ancient extant lineage of jawless fish of the order Petromyzontiformes , placed in the superclass Cyclostomata. The adult lamprey may be characterized by a toothed, funnel-like s ...

hagfish Hagfish, of the class Myxini (also known as Hyperotreti) and order Myxiniformes , are eel-shaped, slime-producing marine fish (occasionally called slime eels). They are the only known living animals that have a skull but no vertebral column, ...

. Juvenile

feed by sucking up mud containing micro-organisms and organic debris. The lamprey has well-developed eyes, while the hagfish has only primitive eyespots. The hagfish coats itself and carcasses it finds with noxious slime to deter predators, and periodically ties itself into a knot to scrape the slime off. It is the only

invertebrate Invertebrates are a paraphyletic group of animals that neither possess nor develop a vertebral column (commonly known as a ''backbone'' or ''spine''), derived from the notochord. This is a grouping including all animals apart from the chordate ...

fish and the only animal which has a

skull The skull is a bone protective cavity for the brain. The skull is composed of four types of bone i.e., cranial bones, facial bones, ear ossicles and hyoid bone. However two parts are more prominent: the cranium and the mandible. In humans, t ...

but no

vertebral column The vertebral column, also known as the backbone or spine, is part of the axial skeleton. The vertebral column is the defining characteristic of a vertebrate in which the notochord (a flexible rod of uniform composition) found in all chordate ...

. Image:Sea Lamprey fish.jpg, Lampreys attached to a

lake trout The lake trout (''Salvelinus namaycush'') is a freshwater char living mainly in lakes in northern North America. Other names for it include mackinaw, namaycush, lake char (or charr), touladi, togue, and grey trout. In Lake Superior, it can als ...

Image:Boca de lamprea.1 - Aquarium Finisterrae.JPG, Mouth of a

sea lamprey The sea lamprey (''Petromyzon marinus'') is a parasitic lamprey native to the Northern Hemisphere. It is sometimes referred to as the "vampire fish". Description The sea lamprey has an eel-like body without paired fins. Its mouth is jawless, ...

File:Pacific_hagfish_Myxine.jpg,

Pacific hagfish The Pacific hagfish (''Eptatretus stoutii'') is a species of hagfish. It lives in the mesopelagic to abyssal Pacific ocean, near the ocean floor. It is a jawless fish and has a body plan that resembles early paleozoic fish. They are able to excre ...

resting on bottom at 280 m File:Korean cuisine-Kkomjangeo bokkeum-01.jpg, Stir-fried hagfish, from Korean cuisine

Cartilaginous fish

Cartilaginous fish Chondrichthyes (; ) is a class that contains the cartilaginous fishes that have skeletons primarily composed of cartilage. They can be contrasted with the Osteichthyes or ''bony fishes'', which have skeletons primarily composed of bone tissue. ...

have a

cartilaginous Cartilage is a resilient and smooth type of connective tissue. In tetrapods, it covers and protects the ends of long bones at the joints as articular cartilage, and is a structural component of many body parts including the rib cage, the neck a ...

skeleton. However, their ancestors were bony animals, and were the first fish to develop paired fins. Cartilaginous fish don't have

swim bladder The swim bladder, gas bladder, fish maw, or air bladder is an internal gas-filled organ that contributes to the ability of many bony fish (but not cartilaginous fish) to control their buoyancy, and thus to stay at their current water depth w ...

s. Their skin is covered in

placoid scale A fish scale is a small rigid plate that grows out of the skin of a fish. The skin of most jawed fishes is covered with these protective scales, which can also provide effective camouflage through the use of reflection and colouration, as we ...

s (dermal denticles) that are as rough as

sandpaper upright=1.35, Sheets of sandpaper with different grit sizes (40 (coarse), 80, 150, 240, 600 (fine)). Sandpaper and glasspaper are names used for a type of coated abrasive that consists of sheets of paper or cloth with abrasive material glued t ...

. Because cartilaginous fish do not have bone marrow, the

spleen The spleen is an organ found in almost all vertebrates. Similar in structure to a large lymph node, it acts primarily as a blood filter. The word spleen comes .

and special tissue around the

gonad A gonad, sex gland, or reproductive gland is a mixed gland that produces the gametes and sex hormones of an organism. Female reproductive cells are egg cells, and male reproductive cells are sperm. The male gonad, the testicle, produces ...

s produces

red blood cell Red blood cells (RBCs), also referred to as red cells, red blood corpuscles (in humans or other animals not having nucleus in red blood cells), haematids, erythroid cells or erythrocytes (from Greek ''erythros'' for "red" and ''kytos'' for "holl ...

s. Their tails can be asymmetric, with the upper lobe longer than the lower lobe. Some cartilaginous fishes possess an organ called a

Leydig's organ Leydig's organ (named after the German histologist Franz Leydig who first described it in 1857) is a unique structure found only in some, but not all, elasmobranchs (sharks and rays). Nestled along the top and bottom of the esophagus, it produces r ...

which also produces red blood cells. There are over 980 species of cartilaginous fish. They include

shark Sharks are a group of elasmobranch fish characterized by a cartilaginous skeleton, five to seven gill slits on the sides of the head, and pectoral fins that are not fused to the head. Modern sharks are classified within the clade Selachi ...

ray Ray may refer to: Fish * Ray (fish), any cartilaginous fish of the superorder Batoidea * Ray (fish fin anatomy), a bony or horny spine on a fin Science and mathematics * Ray (geometry), half of a line proceeding from an initial point * Ray (gra ...

s and

chimaera Chimaeras are cartilaginous fish in the order Chimaeriformes , known informally as ghost sharks, rat fish, spookfish, or rabbit fish; the last three names are not to be confused with rattails, Opisthoproctidae, or Siganidae, respectively. A ...

. File:Tiger shark.png,

Tiger shark The tiger shark (''Galeocerdo cuvier'') is a species of requiem shark and the last extant member of the family Galeocerdonidae. It is a large macropredator, capable of attaining a length over . Populations are found in many tropical and temper ...

File:Whale shark tofo mozambique 2007.jpg,

Whale shark The whale shark (''Rhincodon typus'') is a slow-moving, filter-feeding carpet shark and the largest known extant fish species. The largest confirmed individual had a length of .McClain CR, Balk MA, Benfield MC, Branch TA, Chen C, Cosgrove J, ...

File:Pastinachus_sephen_Day.jpg,

Stingray Stingrays are a group of sea rays, which are cartilaginous fish related to sharks. They are classified in the suborder Myliobatoidei of the order Myliobatiformes and consist of eight families: Hexatrygonidae (sixgill stingray), Plesiobatid ...

File:Callorhinchus callorhynchus.JPG, This elephant fish is a

Bony fish

Bony fish Osteichthyes (), popularly referred to as the bony fish, is a diverse superclass of fish that have skeletons primarily composed of bone tissue. They can be contrasted with the Chondrichthyes, which have skeletons primarily composed of cartila ...

include the

lobe-finned fish Sarcopterygii (; ) — sometimes considered synonymous with Crossopterygii () — is a taxon (traditionally a class or subclass) of the bony fishes known as the lobe-finned fishes. The group Tetrapoda, a mostly terrestrial superclass includi ...

and the

ray finned fish Actinopterygii (; ), members of which are known as ray-finned fishes, is a class of bony fish. They comprise over 50% of living vertebrate species. The ray-finned fishes are so called because their fins are webs of skin supported by bony or ho ...

. The lobe-finned fish is the

class Class or The Class may refer to: Common uses not otherwise categorized * Class (biology), a taxonomic rank * Class (knowledge representation), a collection of individuals or objects * Class (philosophy), an analytical concept used differently ...

of fleshy finned fishes, consisting of

lungfish Lungfish are freshwater vertebrates belonging to the order Dipnoi. Lungfish are best known for retaining ancestral characteristics within the Osteichthyes, including the ability to breathe air, and ancestral structures within Sarcopterygii, i ...

and

coelacanth The coelacanths ( ) are fish belonging to the order Actinistia that includes two extant species in the genus '' Latimeria'': the West Indian Ocean coelacanth (''Latimeria chalumnae''), primarily found near the Comoro Islands off the east c ...

s. They are bony fish with fleshy, lobed paired fins, which are joined to the body by a single bone. These fins evolved into the legs of the first

land

vertebrates Vertebrates () comprise all animal taxa within the subphylum Vertebrata () ( chordates with backbones), including all mammals, birds, reptiles, amphibians, and fish. Vertebrates represent the overwhelming majority of the phylum Chordata, wi ...

, amphibians. Ray finned fishes are so-called because they possess

lepidotrichia Fins are distinctive anatomical features composed of bony spine (zoology), spines or Ray (fish fin anatomy), rays protruding from the body of a fish. They are covered with skin and joined together either in a webbed fashion, as seen in most bon ...

or "fin rays", their fins being webs of skin supported by bony or horny spines ("rays"). There are three types of ray finned fishes: the chondrosteans,

holosteans Holostei is a group of ray-finned bony fish. It is divided into two major clades, the Halecomorphi, represented by a single living species, the bowfin ('' Amia calva''), as well as the Ginglymodi, the sole living representatives being the gars ( ...

, and

teleosts Teleostei (; Greek ''teleios'' "complete" + ''osteon'' "bone"), members of which are known as teleosts ), is, by far, the largest infraclass in the class Actinopterygii, the ray-finned fishes, containing 96% of all extant species of fish. Teleo ...

. The chondrosteans and holosteans are among the earlier fish to evolve, and share characteristics with both teleosts and sharks. In comparison with the other chondrosteans, the holosteans are closer to the teleosts and further from sharks. File:Barramunda.jpg,

Lungfish Lungfish are freshwater vertebrates belonging to the order Dipnoi. Lungfish are best known for retaining ancestral characteristics within the Osteichthyes, including the ability to breathe air, and ancestral structures within Sarcopterygii, i ...

can breathe in air as well as water File:Coelacanth1.JPG, Model of a

, thought until 1938 to be extinct. They are deep blue. File:Sturgeon2.jpg, This

Atlantic sturgeon The Atlantic sturgeon (''Acipenser oxyrinchus oxyrinchus'') is a member of the family Acipenseridae and along with other sturgeon it is sometimes considered a living fossil. The Atlantic sturgeon is one of two subspecies of ''Acipenser oxyrinchu ...

is a chondrostean File:Amia calva1.jpg, This

bowfin The bowfin (''Amia calva'') is a bony fish, native to North America. Common names include mudfish, mud pike, dogfish, grindle, grinnel, swamp trout, and choupique. It is regarded as a relict, being the sole surviving species of the Halecomorp ...

is a holostean

Teleosts

Teleost Teleostei (; Greek ''teleios'' "complete" + ''osteon'' "bone"), members of which are known as teleosts ), is, by far, the largest infraclass in the class Actinopterygii, the ray-finned fishes, containing 96% of all extant species of fish. Tel ...

s are the most advanced or "modern" fishes. They are overwhelmingly the dominant class of fishes (or for that matter,

s) with nearly 30,000 species, covering about 96 percent of all extant fish species. They are ubiquitous throughout

fresh water Fresh water or freshwater is any naturally occurring liquid or frozen water containing low concentrations of dissolved salts and other total dissolved solids. Although the term specifically excludes seawater and brackish water, it does incl ...

and

environments from the deep sea to the highest mountain streams. Included are nearly all the important

commercial Commercial may refer to: * a dose of advertising conveyed through media (such as - for example - radio or television) ** Radio advertisement ** Television advertisement * (adjective for:) commerce, a system of voluntary exchange of products and s ...

and recreational fishes. Teleosts have a movable

maxilla The maxilla (plural: ''maxillae'' ) in vertebrates is the upper fixed (not fixed in Neopterygii) bone of the jaw formed from the fusion of two maxillary bones. In humans, the upper jaw includes the hard palate in the front of the mouth. The ...

and

premaxilla The premaxilla (or praemaxilla) is one of a pair of small cranial bones at the very tip of the upper jaw of many animals, usually, but not always, bearing teeth. In humans, they are fused with the maxilla. The "premaxilla" of therian mammal has ...

and corresponding modifications in the jaw musculature. These modifications make it possible for teleosts to protrude their jaws outwards from the mouth. The

caudal fin Fins are distinctive anatomical features composed of bony spines or rays protruding from the body of a fish. They are covered with skin and joined together either in a webbed fashion, as seen in most bony fish, or similar to a flipper, as s ...

is homocercal, meaning the upper and lower lobes are about equal in size. The spine ends at the

caudal peduncle Fins are distinctive anatomical features composed of bony spines or rays protruding from the body of a fish. They are covered with skin and joined together either in a webbed fashion, as seen in most bony fish, or similar to a flipper, as se ...

, distinguishing this group from those in which the spine extends into the upper lobe of the

. File:Swordfish (PSF).png,

Swordfish Swordfish (''Xiphias gladius''), also known as broadbills in some countries, are large, highly migratory predatory fish characterized by a long, flat, pointed bill. They are a popular sport fish of the billfish category, though elusive. Swordfi ...

are teleosts File:Rose fish.jpg,

Rose fish ''Sebastes norvegicus'', the rose fish, ocean perch, Atlantic redfish, Norway haddock, golden redfish or pinkbelly rosefish, is a species of marine ray-finned fish belonging to the subfamily Sebastinae, the rockfishes, part of the family Scor ...

are also teleosts File:Anguilla japonica 1856.jpg,

Eel Eels are ray-finned fish belonging to the order Anguilliformes (), which consists of eight suborders, 19 families, 111 genera, and about 800 species. Eels undergo considerable development from the early larval stage to the eventual adult stage ...

s are teleosts too Image:Hippocampus.jpg, So are

seahorse A seahorse (also written ''sea-horse'' and ''sea horse'') is any of 46 species of small marine fish in the genus ''Hippocampus''. "Hippocampus" comes from the Ancient Greek (), itself from () meaning "horse" and () meaning "sea monster" or " ...

By habitat

By life span

Some of the shortest-lived species are

gobies Gobiidae or gobies is a family of bony fish in the order Gobiiformes, one of the largest fish families comprising more than 2,000 species in more than 200 genera. Most of gobiid fish are relatively small, typically less than in length, and th ...

, which are small

coral reef A coral reef is an underwater ecosystem characterized by reef-building corals. Reefs are formed of colonies of coral polyps held together by calcium carbonate. Most coral reefs are built from stony corals, whose polyps cluster in groups. C ...

–dwelling fish. Some of the longest-lived are

rockfish Rockfish is a common term for several species of fish, referring to their tendency to hide among rocks. The name rockfish is used for many kinds of fish used for food. This common name belongs to several groups that are not closely related, and ca ...

By size

By breeding behavior

By brooding behavior

By feeding behaviour

By vision

By shape

Boxfish Ostraciidae is a family of squared, bony fish belonging to the order Tetraodontiformes, closely related to the pufferfishes and filefishes. Fish in the family are known variously as boxfishes, cofferfishes, cowfishes and trunkfishes. It contains ...

es have heavily armoured plate-like scales fused into a solid, triangular, boxlike

carapace A carapace is a dorsal (upper) section of the exoskeleton or shell in a number of animal groups, including arthropods, such as crustaceans and arachnids, as well as vertebrates, such as turtles and tortoises. In turtles and tortoises, the und ...

, from which the fins, tail, eyes and mouth protrude. Because of this heavy armour, boxfish move slowly, but few other fish are able to eat the adults. File:Humpback Turretfish - Tetrosomus gibbosus 2.jpg, The

humpback turretfish ''Tetrosomus gibbosus'', commonly called camel cowfish because of the hump on its dorsal keel, is one of 22 species in the boxfish family, Ostraciidae. It is a ray finned fish. Other common names include helmet cowfish, humpback turretfish and th ...

is a

boxfish Ostraciidae is a family of squared, bony fish belonging to the order Tetraodontiformes, closely related to the pufferfishes and filefishes. Fish in the family are known variously as boxfishes, cofferfishes, cowfishes and trunkfishes. It contains ...

with an armoured triangular shaped body File:Leafy Sea Dragon SA.jpg, The leafy sea dragon is camouflaged to look like floating seaweed

By locomotion

By toxicity

By human use

By vulnerability

resilience Resilience, resilient, resiliency, or ''variation'', may refer to: Science Ecology * Ecological resilience, the capacity of an ecosystem to recover from perturbations ** Climate resilience, the ability of systems to recover from climate change * ...

Other

Notes

References

* * * * Weis, Judith S (2011
'Do Fish Sleep?: Fascinating Answers to Questions about Fishes''
Rutgers University Press. .

External links

Oceans for Youth Foundation.
The 20 weirdest fish in the ocean
''Christian Science Monitor''. 22 February 22, 2010. {{DEFAULTSORT:Diversity Of Fish Ichthyology