Euphrasia

''Euphrasia'', or eyebright, is a genus of about 450 species of herbaceous flowering plants in the family Orobanchaceae (formerly included in the Scrophulariaceae), with a cosmopolitan distribution. They are semi-parasitic on grasses and other plants. The common name refers to the plant's use in treating eye infections.

Many species are found in alpine or sub-alpine meadows where snow is common. Flowers usually are borne terminally, are zygomorphic, and have a lower petal shaped like a lip. The most common flower colours are purple, blue-white, and violet. Some species have yellow markings on the lower petal to act as a guide to pollinating insects.

Alternative names, mainly in herbalism, are ''Augentrostkraut, Euphrasiae herba, Herba Euphrasiae'' and ''Herbe d'Euphraise''.

Use in herbalism and medicine

The plant was known to classical herbalists, but then was not referred to until mentioned again in 1305. Nicholas Culpeper assigned it to the Zodiac sign Leo, claiming that it strengthened the brain. It was also used to treat bad memory and vertigo.

In the Elizabethan era, the plant was used in ales, and Gervase Markham's ''Countrie Farm'' (1616) said that one should "Drinke everie morning a small draught of Eyebright wine."

Herbalists use eyebright as a poultice with or without concurrent administration of a tea for the redness, swelling, and visual disturbances caused by blepharitis, and conjunctivitis. The herb is also used for eyestrain and to relieve inflammation caused by colds, coughs, sinus infections, sore throats, and hay fever.

Parts used include the leaf, the stem, and small pieces of the flowers. Typical preparations include a warm compress, or tea. Eyebright preparations are also available as an extract or capsule.

A 2010 report from the European Medicines Agency on the efficacy of Euphrasia remedies states:

From the presence of secondary metabolites, an astringent and anti-inflammatory activity can be hypothesized for Euphrasia preparations. The ocular use of Euphrasia is based upon tradition. However, since the efficacy of the claimed ocular uses is undocumented and external eye application is not hygienic, therapeutic use cannot be recommended.

Phylogeny

The phylogeny of the genera of Rhinantheae has been explored using molecular characters. ''Euphrasia'' belongs to the core Rhinantheae. ''Euphrasia'' is the sister genus to '' Odontites'', '' Bellardia'', ''Tozzia'', and '' Hedbergia''. In turn, these five genera share phylogenetic affinities with ''Bartsia''.

Taxonomy and identification

The genus ''Euphrasia'' is taxonomically complicated due to many species being interfertile and prone to hybridisation. Despite there having been a number of taxonomic revisions the appropriate rank of many taxa, as well as the relationships between them, remains unclear.

Selected species

* '' Euphrasia × aequalis''
* '' Euphrasia alpina''
* '' Euphrasia alsa'' F.Muell.
* '' Euphrasia anglica'' Pugsley
* ''Euphrasia arctica'' Lange ex Rostrup
**''Euphrasia arctica'' subsp. ''arctica'' (''E.borealis'' auct. non (F.Towns.) Wettsd.)
**''Euphrasia arctica'' sybsp. ''borealis'' (F.Towns) Yeo
* ''Euphrasia arguta'' – believed extinct, rediscovered 2008
* '' Euphrasia azorica''
* '' Euphrasia brevipila''
* '' Euphrasia calida''
* '' Euphrasia cambrica'' Pugsley
* '' Euphrasia campbelliae'' Pugsley
* '' Euphrasia collina'' R.Br. – purple eyebright
** ''Euphrasia collina'' ssp. ''muelleri'' – Mueller's eyebright
** ''Euphrasia collina'' ssp. ''osbornii'' – Osborn's eyebright
* '' Euphrasia confusa'' Pugsley
* '' Euphrasia coreana'' W.Becker – Korean eyebright
* '' Euphrasia cuneata'' – North Island eyebright
* ''Euphrasia crassiuscula'' Gand
* '' Euphrasia fabula''
* '' Euphrasia fennica''
* '' Euphrasia foulaensis'' F.Towns. ex Wettst
* '' Euphrasia fragosa'' – shy eyebright, Southport eyebright
* '' Euphrasia frigida'' Pugsley – cold-weather eyebright
* '' Euphrasia gibbsiae''
** ''Euphrasia gibbsiae'' subsp. ''psilantherea''
* '' Euphrasia glabrescens''
* '' Euprasia grandiflora''
* '' Euphrasia heslop-harrisonii'' Pugsley
* ''Euphrasia hirtella''
* ''Euphrasia hudsoniana'' – Hudson's eyebright
* '' Euphrasia insignis'' Wettst
* '' Euphrasia kingii''
* '' Euphrasia lasianthera'' – hairy eyebright
* '' Euphrasia marshallii'' Pugsley
* '' Euphrasia micrantha'' Rchb.
* '' Euphrasia minima''
* '' Euphrasia nemorosa'' (Pers.) Wallr. – common eyebright
* '' Euphrasia officinalis coll.'' – doctor's eyebright, or medical eyebright
* '' Euphrasia officinalis L'' – ''see'' ''Euphrasia rostkoviana''
* '' Euphrasia oakesii'' – Oakes' eyebright
* '' Euphrasia ostenfeldii'' (Pugsley) Yeo
* '' Euphrasia parviflora''
* '' Euphrasia pseudokerneri'' Pugsley – chalk eyebright
* '' Euphrasia randii'' – small eyebright
* ''Euphrasia rivularis'' Pugsley
* ''Euphrasia rostkoviana'' Hayne – red eyebright, " figwort"
**''Euphrasia rostkoviana'' subsp. ''rostkoviana''
**''Euphrasia rostkoviana'' subsp. ''montana'' (Jord.) Wettst.
* '' Euphrasia rotundifolia'' Pugsley
* ''Euphrasia ruptura'' – extinct
* '' Euphrasia salisburgensis'' Funk.
* '' Euphrasia scabra'' R.Br. – rough eyebright
* '' Euphrasia scottica'' Wettst.
* '' Euphrasia semipicta'' – peninsula eyebright
* '' Euphrasia striata'' R.Br.
* ''Euphrasia stricta'' D.Wolff ex J.F.Lehm.
* '' Euphrasia subarctica'' – arctic eyebright
* '' Euphrasia suborbicularis'' – roundleaf eyebright
* '' Euphrasia tatrae''
* '' Euphrasia tetraquetra'' (Bréb.) .
* '' Euphrasia vernalis''
* '' Euphrasia × vestita''
* '' Euphrasia vigursii'' Davey
* '' Euphrasia × villosa''
* '' Euphrasia vinacea'' – glacier eyebright
* ''Euphrasia zelandica''
* ''Euphrasia'' sp. 'Bivouac Bay' – Bivouac Bay eyebright

Footnotes

References

{{Authority control

Orobanchaceae genera
Taxa named by Carl Linnaeus
Parasitic plants