The Eudaminae are a subfamily of

skipper butterflies Skippers are a family of the Lepidoptera (moths and butterflies) named the Hesperiidae. Being diurnal, they are generally called butterflies. They were previously placed in a separate superfamily, Hesperioidea; however, the most recent taxonomy ...

(

family Family (from la, familia) is a group of people related either by consanguinity (by recognized birth) or affinity (by marriage or other relationship). The purpose of the family is to maintain the well-being of its members and of society. Idea ...

Hesperiidae). Their original

type genus In biological taxonomy, the type genus is the genus which defines a biological family and the root of the family name. Zoological nomenclature According to the International Code of Zoological Nomenclature, "The name-bearing type of a nomina ...

''Eudamus'' is today a junior synonym of ''

Urbanus Urbain Servranckx (born 7 June 1949), better known as Urbanus, is a Belgian comedian, actor, singer and comic book writer. Although he is most famous as comedian, some of his songs became hits, such as ''Bakske vol met stro'' (1979), ''Madammen ...

''. They are largely found in the

Neotropics The Neotropical realm is one of the eight biogeographic realms constituting Earth's land surface. Physically, it includes the tropical terrestrial ecoregions of the Americas and the entire South American temperate zone. Definition In bi ...

, with some extending into temperate North America, and one genus, '' Lobocla'', endemic to East Asia. (2009):

Tree of Life Web Project The Tree of Life Web Project is an Internet project providing information about the diversity and phylogeny of life on Earth. This collaborative peer reviewed project began in 1995, and is written by biologists from around the world. The site ...

&ndash
Eudaminae
Version of 2009-JUN-11. Retrieved 2009-DEC-24.

Taxonomy

The Eudaminae have been recent subject to significant taxonomic revisions based on genome analysis, including by Brower & Warren 2009, and Li et al. 2019. Historically, the subfamily has been included as tribe Eudamini in subfamily Pyrginae, based on perceived similarities with two of the tribes in that subfamily, the

Celaenorrhinini The Celaenorrhinini are a tribe of spread-winged skippers in the skipper butterfly subfamily Pyrginae. When the Eudaminae were still included in the Pyrginae as a tribe, their delimitation versus the Celaenorrhinini was disputed, and there were ...

and

Pyrgini The Pyrgini are a tribe in the skipper butterfly subfamily Pyrginae. Formerly, when only four tribes of Pyrginae were recognized, the Pyrgini contained the largest number of genera among these. But this overly wide delimitation has since turned ...

. As of Li et al. 2019, the Eudaminae are divided into four tribes: Entheini,

Phocidini The Eudaminae are a subfamily of skipper butterflies ( family Hesperiidae). Their original type genus ''Eudamus'' is today a junior synonym of '' Urbanus''. They are largely found in the Neotropics, with some extending into temperate North Amer ...

, Eudamini and Oileidini.

Current status and subdivisions

Except where otherwise noted, the classification below follows Li et al., 2019:

Tribe Entheini

*''

Drephalys ''Drephalys'' is a Neotropical butterfly genus in the family Hesperiidae, in which it is placed in tribe Entheini. The genus was established in 1893 by Edward Yerbury Watson, who assigned ''Eudamus helixus'' (now '' Drephalys helixus'') as typ ...

'' Watson, 1893 *''

Udranomia ''Udranomia'' is a genus of Neotropical butterflies in the family Hesperiidae, in which it is placed in tribe Entheini. It was established in 1870 by Arthur Gardiner Butler. Species Per Li et al. 2019, the genus contains the following species ...

'' A. Butler, 1870 *''

Phanus ''Phanus'' is a genus of butterflies in the family Hesperiidae, in which it is placed in tribe Entheini. Species *'' Phanus albiapicalis'' Austin, 1993 Mexico *'' Phanus australis'' Miller, 1965 Brazil *'' Phanus confusis'' Austin, 1993 Mexico ...

'' Hübner,

819 __NOTOC__ Year 819 ( DCCCXIX) was a common year starting on Saturday (link will display the full calendar) of the Julian calendar. Events By place Europe * Spring – Emperor Louis I marries Judith of Bavaria in Aachen.Rogers, B ...

/small> *'' Hyalothyrus'' Mabille, 1878 *''

Entheus ''Entheus'' is a Neotropical genus of skippers in the family Hesperiidae, in which it is placed in the tribe Entheini The Eudaminae are a subfamily of skipper butterflies (family Hesperiidae). Their original type genus ''Eudamus'' is today ...

'' Hübner,

/small> *''

Augiades ''Augiades'' is a genus of butterflies in the family Hesperiidae, in which it is placed in tribe Entheini The Eudaminae are a subfamily of skipper butterflies (family Hesperiidae). Their original type genus ''Eudamus'' is today a junior synon ...

'' Hübner,

/small> *'' Tarsoctenus'' E. Watson, 1893

Tribe Phocidini

*''

Phocides ''Phocides'' is a genus of butterflies in the skipper family, Hesperiidae, in which it is placed in tribe Phocidini, of which it is the namesake genus. The distribution of the genus is primarily Neotropical, but a few species occur in the Near ...

'' Hübner,

/small> *'' Pseudonascus'' Austin, 2008 *'' Nascus'' E. Watson, 1893 *'' Aurina'' Evans, 1937 *'' Emmelus'' O. Mielke & Casagrande , 2016 *'' Porphyrogenes'' E. Watson, 1893 *'' Nicephellus'' Austin, 2008 *''

Salatis ''Salatis'' is a Neotropical genus of spread-winged skippers in the family Hesperiidae, in which they are placed to tribe Phocidini. Species of the genus ''Salatis'' are native to Central America and South America, including in the Amazon River ...

'' Evans, 1952 *'' Salantoia'' Grishin, 2019 *''

Sarmientoia ''Sarmientoia'' is a genus of skippers in the family Hesperiidae, in which they are placed to tribe Phocidini The Eudaminae are a subfamily of skipper butterflies (family Hesperiidae). Their original type genus ''Eudamus'' is today a junio ...

'' Berg, 1897 *'' Bungalotis'' E. Watson, 1893 *''

Euriphellus ''Euriphellus'' is a genus of Neotropical butterflies in the family Hesperiidae, in which it is placed in the tribe Phocidini. The genus was separated from ''Dyscophellus'' by George Traut Austin in 2008, and originally contained a single species ...

'' Austin, 2008 *''

Dyscophellus ''Dyscophellus'' is a genus of largely Neotropical butterflies in the family Hesperiidae. The genus has been the subject of recent revisions based on genome analysis. It is currently placed in tribe (biology), tribe Phocidini within the subfamily ...

'' Godman & Salvin, 1893 *''

Phareas ''Phareas'' is a genus of skippers in the family Hesperiidae, within which it is placed in tribe Phocidini The Eudaminae are a subfamily of skipper butterflies (family Hesperiidae). Their original type genus ''Eudamus'' is today a junior s ...

'' Westwood, 1852 *'' Ornilius'' Grishin, 2022 *'' Fulvatis'' Grishin, 2022 *''

Adina ADINA is a commercial engineering simulation software program that is developed and distributed worldwide by ADINA R & D, Inc. The company was founded in 1986 by Dr. Klaus-Jürgen Bathe, and is headquartered in Watertown, Massachusetts, United St ...

'' Grishin, 2022

Tribe Eudamini

Subtribe Eudamina

*''

Cecropterus ''Cecropterus'' is a genus of skipper (butterfly), skippers in the family Hesperiidae, within which it is placed in subtribe Eudamina. It has three subgenera: ''Cecropterus'', ''Thorybes'' and ''Murgaria''. Species As of 2019, the genus holds th ...

'' Herrich-Schäffer, 1869 *'' Spicauda'' Grishin, 2019 *''

'' Hübner,

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/small> *''

Telegonus Telegonus (; Ancient Greek: Τηλέγονος means "born afar") is the name shared by three different characters in Greek mythology. * Telegonus, a king of Egypt who was sometimes said to have married the nymph Io. * Telegonus, a Thracian son ...

'' Hübner,

/small> *'' Autochton'' Hübner, 1823 *'' Spathilepia'' A. Butler, 1870 *''

Astraptes ''Astraptes'', commonly known as the flasher butterflies, is a genus of skipper butterflies in the subfamily Eudaminae. They are found in the Nearctic and Neotropical ecozones. Species The following species are recognised in the genus ''Astrapte ...

'' Hübner,

/small> *''

Narcosius ''Narcosius'' is a genus of butterflies in the family Hesperiidae (Eudaminae The Eudaminae are a subfamily of skipper butterflies (family Hesperiidae). Their original type genus ''Eudamus'' is today a junior synonym of ''Urbanus''. They are ...

'' Steinhauser, 1986 *''

Proteides ''Proteides'' is a Neotropical genus of spread-winged skipper butterflies in the family Hesperiidae. Species The following two species are recognised in the genus ''Proteides'' *'' Proteides maysi'' (Lucas, 1857) Cuba *'' Proteides mercurius'' ...

'' Hübner,

/small> *''

Epargyreus ''Epargyreus'' is the genus of flasher butterflies, in the subfamily Eudaminae of skipper (butterfly), skipper butterflies. The species are found in the Nearctic realm, Nearctic and Neotropical realms. Taxonomy Etymology The genus name is derive ...

'' Hübner,

/small> *''

Chioides ''Chioides'' is a mainly Neotropical genus of skipper butterflies in the family Hesperiidae (Eudaminae) They range from the south-western United States to Argentina and there are several Antillean endemic species. They are robust and have hindw ...

'' Lindsey, 1921

Subtribe Loboclina

*'' Aguna'' R. Williams, 1927 *'' Zeutus'' Grishin, 2019 *'' Lobocla'' Moore, 1884 *'' Lobotractus'' Grishin, 2019 *''

Codatractus ''Codatractus'' is a Neotropical and Nearctic genus in the family Hesperiidae (Eudaminae). Species The following species are recognised in the genus ''Codatractus'': * ''Codatractus cyledis'' (Dyar, 1912) - Mexico * ''Codatractus bryaxis'' (Hew ...

'' Lindsey, 1921 *''

Zestusa ''Zestusa'' is a genus of skippers in the family Hesperiidae subfamily Eudaminae. Species of the genus occur in Mexico and the western United States. Species The following species are recognised in the genus ''Zestusa'': * ''Zestusa dorus '' ...

'' Lindsey, 1925 *''

Ridens ''Ridens'' is a genus of Neotropical butterflies in the family Hesperiidae (Eudaminae). Species *''Ridens allyni'' Freeman, 1979 - Mexico, Guatemala *''Ridens bidens'' Austin, 1998 - Brazil (Rondônia). *''Ridens biolleyi'' (Mabille, 1900) - Cos ...

'' Evans, 1952 *''

Venada ''Venada'' is a genus of skipper (butterfly), skippers in the family Hesperiidae. Species ''Venada'' consists of the following species: *''Venada advena'' *''Venada cacao'' *''Venada daneva'' *''Venada lamella'' Burns ''in'' Burns, Janzen, Hal ...

'' Evans, 1952

Subtribe Cephisina

*''

Cephise ''Cephise'' is a Neotropical genus of skipper butterflies of the subfamily Eudaminae, within which it is placed in subtribe Cephisina. Species Per Li et al. 2019, the genus contains the following species: *''Cephise aelius'' (Plötz, 1880) – ...

'' Evans, 1952

Subtribe Telemiadina

*''

Ectomis ''Ectomis'' is a genus of Neotropical and Nearctic butterflies in the family Hesperiidae (Eudaminae). Species The following species are recognised in the genus ''Ectomis'': * ''Ectomis'' (''Asina'') ** ''Ectomis gyges'' Evans, 1952 - Venezuela ...

'' Mabille, 1878 *''

Telemiades ''Telemiades'' is a genus of Neotropical butterflies in the family Hesperiidae (subfamily Eudaminae). Species The following species are recognised in the genus ''Telemiades'': *''Telemiades amphion'' (Carl Geyer, Geyer, 1832) **''T. amphion amph ...

'' Hübner,

/small> *''

Polygonus Polygonus may refer to: * Polygonus (genus), a genus of Nearctic and Neotropical spread-winged skippers * Polygonus (mythology) In Greek mythology, Polygonus (Ancient Greek: Πολύγονον means 'prolific') was the Thracian son of the sea go ...

'' Hübner, 825/small>

Tribe Oileidini

Subtribe Oileidina

*'' Flattoides'' Grishin, 2019 *'' Oileides'' Hübner, 825/small>

Subtribe Typhedanina

*''

Typhedanus ''Typhedanus'' is a genus of Neotropical butterflies in the family Hesperiidae (Eudaminae). Species The following species are recognosed in the genus ''Typhedanus'': * ''Typhedanus ampyx'' (Godman & Salvin,

893 __FORCETOC__ Year 893 (Roman numerals, DCCCXCIII) was a common year starting on Monday (link will display the full calendar) of the Julian calendar. Events By place Europe * Vladimir of Bulgaria, Vladimir, ruler (''Khan (title), khan' ...

- Venezuela to Mexico, Amazon ...

'' A. Butler, 1870 *''

Oechydrus ''Oechydrus'' is a genus of Neotropical butterflies in the family Hesperiidae (Eudaminae). Species There are two species recognised in the genus ''Oechydrus'': * '' Oechydrus chersis'' (Herrich-Schäffer, 1869) **''O. c. chersis'' (Herrich-Sc ...

'' E. Watson, 1893 *''

Cogia ''Cogia'' is a genus of Neotropical butterflies in the family Hesperiidae (Eudaminae). Species The following species are recognised in the genus ''Cogia'': *''Cogia abdul'' Hayward, 1947 *''Cogia azila'' Evans, 1953 *''Cogia aziris'' (Hewitson, ...

'' A. Butler, 1870 *''

Nerula ''Nerula'' is a genus of dicot skippers in the butterfly family Hesperiidae (Eudaminae The Eudaminae are a subfamily of skipper butterflies (family Hesperiidae). Their original type genus ''Eudamus'' is today a junior synonym of ''Urbanus'' ...

'' Mabille, 1888 *''

Marela Marela is a town and sub-prefecture in the Faranah Prefecture in the Faranah Region Faranah Region is located in east-central Guinea. It is bordered by the countries of Sierra Leone and Mali and the Guinean regions of Kankan, Mamou, Nzé ...

'' Mabille, 1903

Footnotes and references

Butterfly subfamilies Taxa named by Paul Mabille {{Hesperiidae-stub